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<document id="2C7648D7A9E15F8914C88976A2456CA1" ID-CLB-Dataset="88683" ID-DOI="10.5281/zenodo.6448815" ID-GBIF-Dataset="bbbf94d9-a910-4cda-97df-7eca124163ed" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6448815" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="admin" IM.metadata_approvedBy="admin" IM.taxonomicNames_approvedBy="admin" checkinTime="1635825784914" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="03AD87FAFFF8F61689AD3E81F971FA41" docLanguage="en" docName="hbmw_9_Pteropodidae_16.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Nyctimene wrightae Irwin 2017" docType="treatment" docVersion="14" lastPageNumber="118" masterDocId="FF94FF82FFC4F62A891E341CFFA5FF9B" masterDocTitle="Pteropodidae" masterLastPageNumber="162" masterPageNumber="16" pageNumber="118" updateTime="1719592658732" updateUser="admin">
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<mods:title id="73094168FCD816DE148837DBBA91C683">Pteropodidae</mods:title>
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<mods:namePart id="AA5B3A47FE8095A9A0F3699E01A3CE56">Don E. Wilson</mods:namePart>
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<mods:namePart id="54FAFAB00BCBF3520248EEA82CCBF099">Russell A. Mittermeier</mods:namePart>
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<mods:title id="A883DCE86D6E3A9C843138B60AD3BCE2">Handbook of the Mammals of the World Volume 9 Bats</mods:title>
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<treatment id="03AD87FAFFF8F61689AD3E81F971FA41" ID-DOI="http://doi.org/10.5281/zenodo.6788894" ID-GBIF-Taxon="196379470" ID-Zenodo-Dep="6788894" LSID="urn:lsid:plazi:treatment:03AD87FAFFF8F61689AD3E81F971FA41" httpUri="http://treatment.plazi.org/id/03AD87FAFFF8F61689AD3E81F971FA41" lastPageNumber="118" pageId="60" pageNumber="118">
<subSubSection id="C31E6567FFF8F61689AD3E81FEA6F550" box="[179,259,2717,2763]" pageId="60" pageNumber="118" type="multiple">
<paragraph id="8BBB36ECFFF8F61689AD3E81FEA6F550" blockId="60.[174,1283,2717,2844]" box="[179,259,2717,2763]" pageId="60" pageNumber="118">
<heading id="D0F38180FFF8F61689AD3E81FEA6F550" box="[179,259,2717,2763]" pageId="60" pageNumber="118">
<figureCitation id="133F2A69FFF8F61689AD3E81FEA6F550" box="[179,259,2717,2763]" captionStart="Plate 6: Pteropodidae" captionStartId="57.[107,137,3293,3318]" captionTargetBox="[11,2749,17,3661]" captionTargetPageId="56" captionText="95. African Straw-colored Fruit Bat (Eidolon helvum), 96. Madagascan Straw-colored Fruit Bat (Eidolon dupreanum), 98. Steadfast Tube-nosed Fruit Bat (Paranyctimene tenax), 99. Broad-striped Tube-nosed Fruit Bat (Nyctimene aello), 100. Round-eared Tube-nosed Fruit Bat (Nyctimene cyclotis), 101. Mountain Tube-nosed Fruit Bat (Nyctimene certans), 102. Happy Tube-nosed Fruit Bat (Nyctimene wrighiae), 103. Lesser Tube-nosed Fruit Bat (Nyctimene varius), 104. Common Tube-nosed Fruit Bat (Nyctimene albiventer), 105. Dragon Tube-nosed Fruit Bat (Nyctimene draconilla), 106. Demonic Tube-nosed Fruit Bat (Nyctimene masalai), 107. Malaita Tube-nosed Fruit Bat (Nyctimene malaitensis), 108. Umboi Tube-nosed Fruit Bat (Nyctimene vizcaccia), 109. Island Tube-nosed Fruit Bat (Nyctimene major), 110. Philippine Tube-nosed Fruit Bat (Nyctimene rabori), 111. Pallass Tube-nosed Fruit Bat (Nyctimene cephalotes), 112. Keasts Tube-nosed Fruit Bat (Nyctimene keasti), 113. Queensland Tube-nosed Fruit Bat (Nyctimene robinsont)" figureDoi="http://doi.org/10.5281/zenodo.6448867" httpUri="https://zenodo.org/record/6448867/files/figure.png" pageId="60" pageNumber="118">102.</figureCitation>
</heading>
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<subSubSection id="C31E6567FFF8F616880A3E81FCC1F550" box="[276,868,2717,2763]" pageId="60" pageNumber="118" type="vernacular_names">
<paragraph id="8BBB36ECFFF8F616880A3E81FCC1F550" blockId="60.[174,1283,2717,2844]" box="[276,868,2717,2763]" pageId="60" pageNumber="118">
<heading id="D0F38180FFF8F616880A3E81FCC1F550" box="[276,868,2717,2763]" pageId="60" pageNumber="118">
<vernacularName id="050746C2FFF8F616880A3E81FCC1F550" ID-CoL="7W3ZS" authorityName="Irwin" authorityYear="2017" box="[276,868,2717,2763]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" language="eng" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="wrightae">Happy Tube-nosed Fruit Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168A883E81FAA7F550" box="[918,1282,2717,2763]" pageId="60" pageNumber="118" type="nomenclature">
<paragraph id="8BBB36ECFFF8F6168A883E81FAA7F550" blockId="60.[174,1283,2717,2844]" box="[918,1282,2717,2763]" pageId="60" pageNumber="118">
<heading id="D0F38180FFF8F6168A883E81FAA7F550" box="[918,1282,2717,2763]" pageId="60" pageNumber="118">
<taxonomicName id="4C044D6FFFF8F6168A883E81FAA7F550" ID-CoL="7W3ZS" authorityName="Irwin" authorityYear="2017" box="[918,1282,2717,2763]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="wrightae">
<emphasis id="B970EAFEFFF8F6168A883E81FAA7F550" box="[918,1282,2717,2763]" italics="true" pageId="60" pageNumber="118">Nyctimene wrightae</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F61689AE3EC1FCBCF481" pageId="60" pageNumber="118" type="vernacular_names">
<paragraph id="8BBB36ECFFF8F61689AE3EC1FB61F569" blockId="60.[174,1283,2717,2844]" box="[176,1220,2781,2802]" pageId="60" pageNumber="118">
<heading id="D0F38180FFF8F61689AE3EC1FB61F569" box="[176,1220,2781,2802]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F61689AE3EC1FF59F569" bold="true" box="[176,252,2781,2802]" pageId="60" pageNumber="118">French:</emphasis>
<vernacularName id="050746C2FFF8F616881B3EC1FE7CF569" ID-CoL="7W3ZS" authorityName="Irwin" authorityYear="2017" box="[261,473,2781,2802]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" language="fra" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="wrightae">Nyctimeéne de Wright</vernacularName>
/
<emphasis id="B970EAFEFFF8F61688F33EC1FDEDF569" bold="true" box="[493,584,2781,2802]" pageId="60" pageNumber="118">German:</emphasis>
<vernacularName id="050746C2FFF8F6168B503EC1FCD0F569" ID-CoL="7W3ZS" authorityName="Irwin" authorityYear="2017" box="[590,885,2781,2802]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" language="deu" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="wrightae">Wright-Réhrennasenflughund</vernacularName>
/
<emphasis id="B970EAFEFFF8F6168A953EC1FC43F569" bold="true" box="[907,998,2781,2802]" pageId="60" pageNumber="118">Spanish:</emphasis>
<vernacularName id="050746C2FFF8F6168AEE3EC1FB61F569" ID-CoL="7W3ZS" authorityName="Irwin" authorityYear="2017" box="[1008,1220,2781,2802]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" language="esp" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="wrightae">Nyctimeno de Wright</vernacularName>
</heading>
</paragraph>
<paragraph id="8BBB36ECFFF8F61689B13F19FCBCF481" blockId="60.[174,1283,2717,2844]" box="[175,793,2821,2842]" pageId="60" pageNumber="118">
<heading id="D0F38180FFF8F61689B13F19FCBCF481" box="[175,793,2821,2842]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F61689B13F19FE02F481" bold="true" box="[175,423,2821,2842]" pageId="60" pageNumber="118">Other common names:</emphasis>
<vernacularName id="050746C2FFF8F61688B03F19FCBCF481" ID-CoL="7W3ZS" authorityName="Irwin" authorityYear="2017" box="[430,793,2821,2842]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" language="eng" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="wrightae">Wright's Happy Tube-nosed Fruit Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168A113F5BFAC6F4F3" box="[783,1379,2887,2920]" pageId="60" pageNumber="118" type="reference_group">
<paragraph id="8BBB36ECFFF8F6168A113F5BFAC6F4F3" blockId="60.[783,1383,2887,3315]" box="[783,1379,2887,2920]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F6168A113F5BFC0EF4F3" bold="true" box="[783,939,2887,2920]" pageId="60" pageNumber="118">Taxonomy.</emphasis>
<taxonomicName id="4C044D6FFFF8F6168AA43F5BFAFAF4F3" ID-CoL="7W3ZS" authority="Irwin, 2017" authorityName="Irwin" authorityYear="2017" box="[954,1375,2887,2920]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="wrightae">Nyctimene wrightae Irwin, 2017</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168A0F3F6FFBC2F444" pageId="60" pageNumber="118" type="materials_examined">
<paragraph id="8BBB36ECFFF8F6168A0F3F6FFBC2F444" blockId="60.[783,1383,2887,3315]" pageId="60" pageNumber="118">
<materialsCitation id="3B6C3CB1FFF8F6168A0F3F6FFBC2F444" ID-GBIF-Occurrence="3828391328" latitude="-6.25" longitude="142.78334" pageId="60" pageNumber="118">
“Namosado (
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<geoCoordinate id="EE30502BFFF8F6168D7C3F6FFB7BF40B" box="[1122,1246,2931,2960]" degrees="142" direction="east" minutes="47" orientation="longitude" pageId="60" pageNumber="118" precision="925" value="142.78334">142°47E</geoCoordinate>
,
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asl),
<collectingRegion id="49C0F80EFFF8F6168A443F86FAA9F42C" box="[858,1292,2970,2999]" country="Papua New Guinea" name="Southern Highlands" pageId="60" pageNumber="118">Southern Highlands Province</collectingRegion>
,
<collectingCountry id="F313767CFFF8F6168C3C3F86FAC0F42C" box="[1314,1381,2970,2999]" name="Papua New Guinea" pageId="60" pageNumber="118">PNG</collectingCountry>
[=
<collectingCountry id="F313767CFFF8F6168A263FDEFBEAF444" box="[824,1103,3010,3039]" name="Papua New Guinea" pageId="60" pageNumber="118">Papua New Guinea</collectingCountry>
].”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168A0F3FF9FB2CF357" pageId="60" pageNumber="118" type="discussion">
<paragraph id="8BBB36ECFFF8F6168A0F3FF9FB2CF357" blockId="60.[783,1383,2887,3315]" pageId="60" pageNumber="118">
<taxonomicName id="4C044D6FFFF8F6168A0F3FF9FBB4F39D" authorityName="Irwin" authorityYear="2017" box="[785,1041,3045,3078]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="wrightae">Nyctimene wrightae</taxonomicName>
is tentatively included in the
<taxonomicName id="4C044D6FFFF8F6168A633810FC77F3B6" authorityName="K. Andersen" authorityYear="1910" box="[893,978,3084,3117]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="cyclotis">cyclotis</taxonomicName>
species group, although it shares many traits with N. albwenter papuanus. Populations attributed to N. wnrightae have been treated as N.
<taxonomicName id="4C044D6FFFF8F6168DD4389FFABAF33F" authorityName="K. Andersen" authorityYear="1910" box="[1226,1311,3203,3236]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="60" pageNumber="118" phylum="Chordata" rank="species" species="cyclotis">cyclotis</taxonomicName>
and N. a. papuanus. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168A0E38CAFE9BF280" pageId="60" pageNumber="118" type="multiple">
<caption id="DF7B6664FFF8F6168A0E38CAFE9BF280" ID-DOI="http://doi.org/10.5281/zenodo.6803784" ID-Zenodo-Dep="6803784" httpUri="https://zenodo.org/record/6803784/files/figure.png" inLine="true" pageId="60" pageNumber="118" startId="60.[784,957,3286,3315]" targetBox="[169,760,2899,3312]" targetPageId="60">
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<emphasis id="B970EAFEFFF8F6168A0E38CAFC64F368" bold="true" box="[784,961,3286,3315]" pageId="60" pageNumber="118">Distribution.</emphasis>
New
<collectingCountry id="F313767CFFF8F6168D3E38CAFB23F368" box="[1056,1158,3286,3315]" name="Guinea-Bissau" pageId="60" pageNumber="118">Guinea</collectingCountry>
(W, N, C &amp; E lowlands).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C31E6567FFF8F61689AE3939F73DFBD4" pageId="60" pageNumber="118" type="description">
<paragraph id="8BBB36ECFFF8F61689AE3939F73DFBD4" blockId="60.[176,1382,3326,3473]" lastBlockId="60.[1449,2657,281,1500]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F61689AE3939FE0AF2D9" bold="true" box="[176,431,3365,3394]" pageId="60" pageNumber="118">Descriptive notes.</emphasis>
Head-body 84-86-
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, tail 19-
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, ear 9-
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, hindfoot 8:2—-15-
<quantity id="4CFC9B09FFF8F616887F394CFE10F2F2" box="[353,437,3408,3433]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="60" pageNumber="118" unit="mm" value="6.0">6 mm</quantity>
, forearm 52-1-63-
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; weight 34-
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. There is some sexual dimorphism relative to color during reproductive season; hairs on central thorax seem to be whiter and brighter on males, and males also develop striking yellowish orange patch on ventral flanks; females generally have duller fawn brown pelage. Rostrum of the Happy Tube-nosed Fruit Bat is short, with long tubular divergent nostrils. Ears are somewhat elongated, with bluntly pointed tips and thickened along top edges; eyes are large, with dark reddish-brown irises. Dorsal pelage is relatively variable, generally being medium or fawn brown and grayish brown on face, with dark thin dorsal stripe that generally runs from mid-back to tail and thickens toward rump. Ventral pelage is paler brown than dorsum. There also is yellow morph that has lighter yellow brown dorsal and ventral fur, with brown wing spotting and white spotting on forearm and wing digits. Wings and ears are dark brown; ears, tube nostrils, and wing digits have variable amounts of white or yellow spotting; wing membranes have variable amounts of white (occasionally yellow) and darker brown spotting. Wing attaches at second digit of foot. Second digit of wing has a claw, and wing attaches at second digit of foot. Tail is short, black, and wrinkled, and narrow uropatagium connects at base and stretches to calcar at ankles. Claws are black. Skull and mandible are robust; rostrum has shortest length relative to skull length of any species of
<taxonomicName id="4C044D6FFFF8F6168F583793F8B5FC2B" authorityName="Miller" authorityYear="1907" box="[1606,1808,911,944]" class="Mammalia" family="Pteropodidae" kingdom="Animalia" order="Chiroptera" pageId="60" pageNumber="116" phylum="Chordata" rank="subFamily" subFamily="Nyctimeninae">Nyctimeninae</taxonomicName>
. Single lower incisor is completely deciduous, falling out before adulthood; lower molars are broad and rounded in dorsal view; C replaces incisors and is long and powerful; P, is elongated and longer than P, and P; and they have broad dental arcade and palate with very short rostrum, giving them a “smiling” appearance that is alluded to by the common name.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168CB53049F6F9FBED" box="[1451,2396,1109,1142]" pageId="60" pageNumber="118" type="biology_ecology">
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<emphasis id="B970EAFEFFF8F6168CB53049F9BFFBED" bold="true" box="[1451,1562,1109,1142]" pageId="60" pageNumber="118">Habitat.</emphasis>
Lowland to montane rainforests at elevations of
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.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168CB43061F58FFB05" box="[1450,2602,1149,1182]" pageId="60" pageNumber="118" type="food_feeding">
<paragraph id="8BBB36ECFFF8F6168CB43061F58FFB05" blockId="60.[1449,2657,281,1500]" box="[1450,2602,1149,1182]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F6168CB43061F914FB05" bold="true" box="[1450,1713,1149,1182]" pageId="60" pageNumber="118">Food and Feeding.</emphasis>
The Happy Tube-nosed Fruit Bat is assumed to be frugivorous.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168CB530B8F7AAFB76" pageId="60" pageNumber="118" type="breeding">
<paragraph id="8BBB36ECFFF8F6168CB530B8F7AAFB76" blockId="60.[1449,2657,281,1500]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F6168CB530B8F997FB5E" bold="true" box="[1451,1586,1188,1221]" pageId="60" pageNumber="118">Breeding.</emphasis>
Pregnant Happy Tube-nosed Fruit Bats have been captured in January-February and females with volant young in July.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168CB730EFF6ACFA8F" box="[1449,2313,1267,1300]" pageId="60" pageNumber="118" type="activity">
<paragraph id="8BBB36ECFFF8F6168CB730EFF6ACFA8F" blockId="60.[1449,2657,281,1500]" box="[1449,2313,1267,1300]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F6168CB730EFF930FA8F" bold="true" box="[1449,1685,1267,1300]" pageId="60" pageNumber="118">Activity patterns.</emphasis>
Happy Tube-nosed Fruit Bats are nocturnal.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168CB53107F6F2FAA7" box="[1451,2391,1307,1340]" pageId="60" pageNumber="118" type="biology_ecology">
<paragraph id="8BBB36ECFFF8F6168CB53107F6F2FAA7" blockId="60.[1449,2657,281,1500]" box="[1451,2391,1307,1340]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F6168CB53107F7CDFAA7" bold="true" box="[1451,2152,1307,1340]" pageId="60" pageNumber="118">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168CB5315EF639FA29" pageId="60" pageNumber="118" type="conservation">
<paragraph id="8BBB36ECFFF8F6168CB5315EF639FA29" blockId="60.[1449,2657,281,1500]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F6168CB5315EF8ADFAF8" bold="true" box="[1451,1800,1346,1379]" pageId="60" pageNumber="118">Status and Conservation.</emphasis>
<collectingRegion id="49C0F80EFFF8F6168E0D315EF8EDFAF8" box="[1811,1864,1346,1379]" country="Maldives" name="Uthuru" pageId="60" pageNumber="118">Not</collectingRegion>
assessed on The IUCN Red List. Because the Happy Tubenosed Fruit Bat was recently recognized, very little is understood regarding its ecology and threats, although it seems to be widespread and relatively common.
</paragraph>
</subSubSection>
<subSubSection id="C31E6567FFF8F6168CB231DDF971FA41" box="[1452,1748,1473,1498]" pageId="60" pageNumber="118" type="bibRefCitation_list">
<paragraph id="8BBB36ECFFF8F6168CB231DDF971FA41" blockId="60.[1449,2657,281,1500]" box="[1452,1748,1473,1498]" pageId="60" pageNumber="118">
<emphasis id="B970EAFEFFF8F6168CB231DDF9E0FA41" bold="true" box="[1452,1605,1473,1498]" pageId="60" pageNumber="118">Bibliography.</emphasis>
Irwin (2017).
</paragraph>
</subSubSection>
</treatment>
</document>