143 lines
17 KiB
XML
143 lines
17 KiB
XML
<document id="8C511C953D50FBFCAA2FEEE555BE8D37" ID-DOI="10.11646/zootaxa.4576.3.6" ID-GBIF-Dataset="7abdc62c-c350-49dd-8060-4ed64da07362" ID-ISSN="1175-5326" ID-Zenodo-Dep="3715772" ID-ZooBank="56BC8573-D4A1-4B18-9BF6-7AB5F7984BFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584603665960" checkinUser="plazi" docAuthor="Muona, Jyrki" docDate="2019" docId="03D387E9FFA17779FF6FE88CFADDFD48" docLanguage="en" docName="zootaxa.4576.3.6.pdf" docOrigin="Zootaxa 4576 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudothroscus Muona 2019, new genus" docType="treatment" docVersion="5" lastPageNumber="528" masterDocId="FFEAFF91FFA7777EFFF8EE40FF81FFA7" masterDocTitle="Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species" masterLastPageNumber="543" masterPageNumber="521" pageNumber="527" updateTime="1698730058212" updateUser="plazi">
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<mods:title id="474C38F961949A3E06E63243978EDCC6">Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species</mods:title>
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<mods:namePart id="F74AFEE4189E0313565AE0031D85410E">Muona, Jyrki</mods:namePart>
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<mods:title id="94B17D4B2A24697920E3C693B66C3DCA">Zootaxa</mods:title>
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<mods:part id="A7A45D9C5B44E005097D542AEF4F3268">
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<mods:date id="E67D8179087D939AD33E7607D9BFA84B">2019</mods:date>
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<mods:number id="16DEF694D7190D83C6153E02763CCA21">2019-04-03</mods:number>
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<mods:number id="5D8DE1D138FE54DD9AF3F49AF31C2416">4576</mods:number>
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<mods:identifier id="9B0CCFCE2A25C85892ADF5BEE41A3C11" type="DOI">10.11646/zootaxa.4576.3.6</mods:identifier>
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<treatment id="03D387E9FFA17779FF6FE88CFADDFD48" ID-DOI="http://doi.org/10.5281/zenodo.3716780" ID-GBIF-Taxon="162639252" ID-Zenodo-Dep="3716780" LSID="urn:lsid:plazi:treatment:03D387E9FFA17779FF6FE88CFADDFD48" httpUri="http://treatment.plazi.org/id/03D387E9FFA17779FF6FE88CFADDFD48" lastPageId="7" lastPageNumber="528" pageId="6" pageNumber="527">
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<subSubSection id="C3606574FFA17778FF6FE88CFE5CF941" box="[151,477,1740,1766]" pageId="6" pageNumber="527" type="nomenclature">
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<paragraph id="8BC536FFFFA17778FF6FE88CFE5CF941" blockId="6.[151,477,1740,1766]" box="[151,477,1740,1766]" pageId="6" pageNumber="527">
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<heading id="D08D8193FFA17778FF6FE88CFE5CF941" bold="true" box="[151,477,1740,1766]" fontSize="11" level="1" pageId="6" pageNumber="527" reason="1">
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<emphasis id="B90EEAEDFFA17778FF6FE88CFE5CF941" bold="true" box="[151,477,1740,1766]" pageId="6" pageNumber="527">
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<taxonomicName id="4C7A4D7CFFA17778FF6FE88CFED4F941" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[151,341,1740,1766]" class="Insecta" family="Throscidae" genus="Pseudothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90EEAEDFFA17778FF6FE88CFED4F941" bold="true" box="[151,341,1740,1766]" italics="true" pageId="6" pageNumber="527">Pseudothroscus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D5796FFA17778FEA4E88CFE5CF941" box="[348,477,1740,1766]" pageId="6" pageNumber="527" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFA17778FF6FE94CFD22F885" box="[151,675,1804,1826]" pageId="6" pageNumber="527" type="materials_examined">
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<paragraph id="8BC536FFFFA17778FF6FE94CFD22F885" blockId="6.[151,675,1804,1826]" box="[151,675,1804,1826]" pageId="6" pageNumber="527">
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<emphasis id="B90EEAEDFFA17778FF6FE94CFE9DF885" bold="true" box="[151,284,1804,1826]" pageId="6" pageNumber="527">
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<typeStatus id="54C1885DFFA17778FF6FE94CFF4DF885" box="[151,204,1804,1826]" pageId="6" pageNumber="527">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C7A4D7CFFA17778FED2E94CFDA1F885" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[298,544,1804,1826]" class="Insecta" family="Throscidae" genus="Pseudothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B90EEAEDFFA17778FED2E94CFDA1F885" box="[298,544,1804,1826]" italics="true" pageId="6" pageNumber="527">Pseudothroscus balticus</emphasis>
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</taxonomicName>
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<emphasis id="B90EEAEDFFA17778FDDEE94CFD22F885" bold="true" box="[550,675,1804,1826]" pageId="6" pageNumber="527">
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<taxonomicNameLabel id="A23D5796FFA17778FDDEE94CFD22F885" box="[550,675,1804,1826]" pageId="6" pageNumber="527" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFA17779FF6FE907FA03FE0B" lastPageId="7" lastPageNumber="528" pageId="6" pageNumber="527" type="diagnosis">
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<paragraph id="8BC536FFFFA17778FF6FE907FBFBF8C7" blockId="6.[151,1436,1863,2033]" box="[151,1146,1863,1888]" pageId="6" pageNumber="527">
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<emphasis id="B90EEAEDFFA17778FF6FE907FE8BF8C7" bold="true" box="[151,266,1863,1888]" pageId="6" pageNumber="527">Diagnosis</emphasis>
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. Elateroidea synapomorphy: pro-mesothoracic joint with clicking mechanism.
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</paragraph>
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<paragraph id="8BC536FFFFA17778FF3FE92CFDB8F80F" blockId="6.[151,1436,1863,2033]" pageId="6" pageNumber="527">
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<taxonomicName id="4C7A4D7CFFA17778FF3FE92CFEC5F822" authorityName="Laporte" authorityYear="1840" box="[199,324,1900,1925]" class="Insecta" family="Throscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="family">Throscidae</taxonomicName>
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synapomorphies: a2 as wide as a3 and a4, antennal grooves running along notosternal suture and bending laterally along proleg cavity.
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</paragraph>
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<paragraph id="8BC536FFFFA17778FF3FE9F4FE24F857" blockId="6.[151,1436,1863,2033]" pageId="6" pageNumber="527">
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Synapomorphies of clade
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<taxonomicName id="4C7A4D7CFFA17778FE15E9F4FDE8F86A" authorityName="Laporte" authorityYear="1840" box="[493,617,1972,1997]" class="Insecta" family="Throscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="family">Throscidae</taxonomicName>
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without
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<taxonomicName id="4C7A4D7CFFA17778FD37E9F4FC89F86A" authorityName="Muona" authorityYear="1993" box="[719,776,1972,1997]" class="Insecta" family="Throscidae" genus="Jaira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">Jaira</taxonomicName>
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: metasternal discrimen absent, metacoxae with posteriorly projecting median apex.
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</paragraph>
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<paragraph id="8BC536FFFFA07779FF3FEED7FBBEFF5F" blockId="7.[151,1436,151,752]" pageId="7" pageNumber="528">Diagnostic other characters: pronotal lateral carinae complete, head without median carina, eyes large, antennal insertions located in fossae, antennal grooves posteriorly partly separated from proleg impressions by septum, pro- and mesoleg impressions only faintly indicated, tibiae simple, slender.</paragraph>
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<paragraph id="8BC536FFFFA07779FF3FEF45FB5BFEBB" blockId="7.[151,1436,151,752]" box="[199,1242,260,285]" pageId="7" pageNumber="528">
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None of the characters used in the analysis turned out to be apomorphies for
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<taxonomicName id="4C7A4D7CFFA07779FBDBEF44FB52FEBA" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[1059,1235,260,285]" class="Insecta" family="Throscidae" genus="Pseudothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="527" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90EEAEDFFA07779FBDBEF44FB52FEBA" box="[1059,1235,260,285]" italics="true" pageId="7" pageNumber="528">Pseudothroscus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BC536FFFFA07779FF3FEF67FA03FE0B" blockId="7.[151,1436,151,752]" pageId="7" pageNumber="528">
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<taxonomicName id="4C7A4D7CFFA07779FF3FEF67FEF6FEE7" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[199,375,295,320]" class="Insecta" family="Throscidae" genus="Pseudothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="527" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90EEAEDFFA07779FF3FEF67FEF6FEE7" box="[199,375,295,320]" italics="true" pageId="7" pageNumber="528">Pseudothroscus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C7A4D7CFFA07779FE4CEF69FDC8FEE7" authorityName="Kovalev & Kirejtshuk, 2013 in Kovalev, Kirejtshuk and Azar" authorityYear="2013" box="[436,585,297,320]" class="Insecta" family="Throscidae" genus="Potergosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEAEDFFA07779FE4CEF69FDC8FEE7" box="[436,585,297,320]" italics="true" pageId="7" pageNumber="528">Potergosoma</emphasis>
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</taxonomicName>
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share several plesiomorphies: complete lateral pronotal carinae, no tarsal grooves, simple t4, simple tibiae, poorly developed pro-mesoleg impressions, antennal fossae and strong serial elytral punctures.
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<taxonomicName id="4C7A4D7CFFA07779FE9AEF2FFD93FE2F" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[354,530,367,392]" class="Insecta" family="Throscidae" genus="Pseudothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="527" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90EEAEDFFA07779FE9AEF2FFD93FE2F" box="[354,530,367,392]" italics="true" pageId="7" pageNumber="528">Pseudothroscus</emphasis>
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</taxonomicName>
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lacks the
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<taxonomicName id="4C7A4D7CFFA07779FD7EEF31FC9AFE2F" authorityName="Kovalev & Kirejtshuk, 2013 in Kovalev, Kirejtshuk and Azar" authorityYear="2013" box="[646,795,369,392]" class="Insecta" family="Throscidae" genus="Potergosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEAEDFFA07779FD7EEF31FC9AFE2F" box="[646,795,369,392]" italics="true" pageId="7" pageNumber="528">Potergosoma</emphasis>
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</taxonomicName>
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apomorphies (reduced eyes, strong frontal keel) and has unique delicate ridges around the antennal fossae. This combination of characters places it in a genus of its own.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFA07779FF3FEFF7FB5CFD6A" pageId="7" pageNumber="528" type="description">
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<paragraph id="8BC536FFFFA07779FF3FEFF7FB5CFD6A" blockId="7.[151,1436,151,752]" pageId="7" pageNumber="528">
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<emphasis id="B90EEAEDFFA07779FF3FEFF7FED0FE77" bold="true" box="[199,337,439,464]" pageId="7" pageNumber="528">Description</emphasis>
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. Body fairly elongate, flattened, venter convex, dorsum somewhat less so (
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<figureCitation id="13412A7AFFA07779FB3CEFF7FA8CFE77" box="[1220,1293,439,464]" captionStart="FIGURE 4" captionStartId="7.[151,250,1482,1504]" captionTargetBox="[150,1436,802,1467]" captionTargetId="figure@7.[150,1436,802,1467]" captionTargetPageId="7" captionText="FIGURE 4. Pseudothroscus balticus n. sp., dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3715780" httpUri="https://zenodo.org/record/3715780/files/figure.png" pageId="7" pageNumber="528">Fig. 4</figureCitation>
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). Head with small antennal fossae, these with delicate ridges, frontoclypeus small, wide and short, front margin slightly arched, eyes large, labrum well sclerotized, fairly large (
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<figureCitation id="13412A7AFFA07779FD48EFBFFD75FDBF" box="[688,756,511,536]" captionStart="FIGURE 5" captionStartId="8.[151,250,1071,1093]" captionTargetBox="[150,1436,193,1050]" captionTargetId="figure@8.[150,1436,193,1050]" captionTargetPageId="8" captionText="FIGURE 5. Pseudothroscus balticus n. sp., ventral view. AC = antennal club; AF = antennal pit boarded by delicate ridge; L = labrum; LC = complete lateral pronotal ridge; MsT = simple mesotibia." figureDoi="http://doi.org/10.5281/zenodo.3715782" httpUri="https://zenodo.org/record/3715782/files/figure.png" pageId="7" pageNumber="528">Fig. 5</figureCitation>
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). Antennae slender, all antennomeres longer than wide, a2 as wide as a3 and a4, a9–a11 forming a club (
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<figureCitation id="13412A7AFFA07779FD8EEC64FD3BFD9A" box="[630,698,548,573]" captionStart="FIGURE 5" captionStartId="8.[151,250,1071,1093]" captionTargetBox="[150,1436,193,1050]" captionTargetId="figure@8.[150,1436,193,1050]" captionTargetPageId="8" captionText="FIGURE 5. Pseudothroscus balticus n. sp., ventral view. AC = antennal club; AF = antennal pit boarded by delicate ridge; L = labrum; LC = complete lateral pronotal ridge; MsT = simple mesotibia." figureDoi="http://doi.org/10.5281/zenodo.3715782" httpUri="https://zenodo.org/record/3715782/files/figure.png" pageId="7" pageNumber="528">Fig. 5</figureCitation>
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). Prosternum apically parallel-sided, sides diverging rectilinearily posteriorly, lateral carniae complete, apex quite narrow, rounded, anterior edge produced ventrally, curved (
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<figureCitation id="13412A7AFFA07779FAB2EC07FA0FFDC7" box="[1354,1422,583,608]" captionStart="FIGURE 5" captionStartId="8.[151,250,1071,1093]" captionTargetBox="[150,1436,193,1050]" captionTargetId="figure@8.[150,1436,193,1050]" captionTargetPageId="8" captionText="FIGURE 5. Pseudothroscus balticus n. sp., ventral view. AC = antennal club; AF = antennal pit boarded by delicate ridge; L = labrum; LC = complete lateral pronotal ridge; MsT = simple mesotibia." figureDoi="http://doi.org/10.5281/zenodo.3715782" httpUri="https://zenodo.org/record/3715782/files/figure.png" pageId="7" pageNumber="528">Fig. 5</figureCitation>
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). Antennal grooves partly separated from proleg impressions by septum, pro- and mesoleg impressions only faintly indicated, metathorax and abdomen without tarsal grooves. Elytral striae wide, deep, with large punctures, interstices slightly convex (
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<figureCitation id="13412A7AFFA07779FE33ECF4FD91FD6B" box="[459,528,692,717]" captionStart="FIGURE 4" captionStartId="7.[151,250,1482,1504]" captionTargetBox="[150,1436,802,1467]" captionTargetId="figure@7.[150,1436,802,1467]" captionTargetPageId="7" captionText="FIGURE 4. Pseudothroscus balticus n. sp., dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3715780" httpUri="https://zenodo.org/record/3715780/files/figure.png" pageId="7" pageNumber="528">Fig. 4</figureCitation>
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). Tibiae simple, without tarsal grooves, tarsi delicate, t4 simple.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFA07779FF3FEC97FADDFD48" box="[199,1372,727,752]" pageId="7" pageNumber="528" type="etymology">
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<paragraph id="8BC536FFFFA07779FF3FEC97FADDFD48" blockId="7.[151,1436,151,752]" box="[199,1372,727,752]" pageId="7" pageNumber="528">
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<emphasis id="B90EEAEDFFA07779FF3FEC97FEC7FD57" bold="true" box="[199,326,727,752]" pageId="7" pageNumber="528">Etymology</emphasis>
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. Superficially similar to
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<taxonomicName id="4C7A4D7CFFA07779FDA5EC99FD3EFD57" authorityName="Kugelann" authorityYear="1794" box="[605,703,729,752]" class="Insecta" family="Throscidae" genus="Trixagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEAEDFFA07779FDA5EC99FD3EFD57" box="[605,703,729,752]" italics="true" pageId="7" pageNumber="528">Trixagus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C7A4D7CFFA07779FD35EC97FCB2FD57" authorityName="Latreille" authorityYear="1796" box="[717,819,727,752]" class="Insecta" family="Throscidae" genus="Throscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEAEDFFA07779FD35EC97FCB2FD57" box="[717,819,727,752]" italics="true" pageId="7" pageNumber="528">Throscus</emphasis>
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</taxonomicName>
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being its junior synonym previously in wide use.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |