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<document id="A5A294B8C9E709B2B2C8B465E96CEF6B" ID-DOI="10.1111/j.1096-3642.2009.00559.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5438272" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630441115732" checkinUser="valdenar" docAuthor="Huber, Bernhard A., Fischer, Nadine &amp; Astrin, Jonas J." docDate="2010" docId="03D3130F7A12741DDC25BF22FD2947E6" docLanguage="en" docName="j.1096-3642.2009.00559.x.pdf" docOrigin="Zoological Journal of the Linnean Society 158 (2)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00559.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Modisimus cienaga Huber &amp; Fischer &amp; Astrin 2010, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="287" masterDocId="FFEA6B777A387436DF62B903FFF14159" masterDocTitle="High level of endemism in Haiti’s last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules" masterLastPageNumber="299" masterPageNumber="244" pageNumber="286" updateTime="1699271919417" updateUser="plazi">
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<mods:title id="02CCEAF30493D3E9E36F62411CAF157A">High level of endemism in Haiti’s last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules</mods:title>
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<mods:namePart id="8857052DA7EEDA40D0962261691893BB">Huber, Bernhard A.</mods:namePart>
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<mods:namePart id="06D9B904FA1D208CD77AF2A0BC875000">Fischer, Nadine</mods:namePart>
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<mods:namePart id="AF65D5B146B7E363121FE54E91365B12">Astrin, Jonas J.</mods:namePart>
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<mods:title id="D5402FBC57E6098EC2FE41C0079C15BF">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="A8C6A6FCB321609B3C9B734496198E8C">2010</mods:date>
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<mods:identifier id="5471653EE5E3C2F98997457D27485422" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03D3130F7A12741DDC25BF22FD2947E6" LSID="urn:lsid:plazi:treatment:03D3130F7A12741DDC25BF22FD2947E6" httpUri="http://treatment.plazi.org/id/03D3130F7A12741DDC25BF22FD2947E6" lastPageId="43" lastPageNumber="287" pageId="42" pageNumber="286">
<subSubSection id="C360F1927A12741CDC25BF22FA65476F" box="[839,1428,1569,1590]" pageId="42" pageNumber="286" type="nomenclature">
<paragraph id="8BC5A2197A12741CDC25BF22FA65476F" blockId="42.[839,1428,1569,1590]" box="[839,1428,1569,1590]" pageId="42" pageNumber="286">
<heading id="D08D15757A12741CDC25BF22FA65476F" box="[839,1428,1569,1590]" centered="true" fontSize="9" level="2" pageId="42" pageNumber="286" reason="4">
<taxonomicName id="4C7AD99A7A12741CDC25BF22FAD7476C" ID-CoL="73N87" authority="HUBER &amp; FISCHER" authorityName="Huber &amp; Fischer &amp; Astrin" authorityYear="2010" box="[839,1318,1569,1590]" class="Arachnida" family="Pholcidae" genus="Modisimus" kingdom="Animalia" order="Araneae" pageId="42" pageNumber="286" phylum="Arthropoda" rank="species" species="cienaga" status="sp. nov.">
<emphasis id="B90E7E0B7A12741CDC25BF22FBB0476C" bold="true" box="[839,1089,1569,1590]" italics="true" pageId="42" pageNumber="286">MODISIMUS CIENAGA</emphasis>
HUBER &amp; FISCHER
</taxonomicName>
<emphasis id="B90E7E0B7A12741CDA4DBF27FA65476F" bold="true" box="[1327,1428,1569,1590]" pageId="42" pageNumber="286">
<taxonomicNameLabel id="A23DC3707A12741CDA4DBF27FA65476F" box="[1327,1428,1569,1590]" pageId="42" pageNumber="286" rank="species">SP. NOV.</taxonomicNameLabel>
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<subSubSection id="C360F1927A12741DDC32BF44FD2947E6" lastPageId="43" lastPageNumber="287" pageId="42" pageNumber="286" type="description">
<paragraph id="8BC5A2197A12741CDC32BF44FA7B4704" blockId="42.[848,1418,1607,1629]" box="[848,1418,1607,1629]" pageId="42" pageNumber="286">
(
<figureCitation id="1341BE9C7A12741CDC3ABF44FC5F4704" box="[856,942,1607,1629]" captionStart="Figures 17–37" captionStartId="13.[144,234,1662,1681]" captionTargetBox="[144,1426,194,1632]" captionTargetId="figure-0@13.[144,1426,194,1632]" captionTargetPageId="13" captionText="Figures 17–37. Habitus photos of males (except 33), showing typical and distinctive characters. 17, 18. M. femoratus. 19. M. jima sp. nov. 20. M. kiskeya sp. nov. 21, 22. M. angulatus sp. nov. 23, 24. M. epepye sp. nov. 25. M. mango sp. nov. 26. M. cienaga sp. nov. 27–29. M. seguin sp. nov. (arrows point to callus-like area). 30–34. M. vittatus. 35. M. palvet sp. nov. 36. M. berac sp. nov. 37. M. paraiso sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5438284" httpUri="https://zenodo.org/record/5438284/files/figure.png" pageId="42" pageNumber="286">Figs 26</figureCitation>
,
<figureCitation id="1341BE9C7A12741CDCDEBF44FBC34705" box="[956,1074,1607,1629]" captionStart="Figures 83–93" captionStartId="16.[164,254,1736,1755]" captionTargetBox="[258,1348,194,1704]" captionTargetId="figure-0@16.[258,1348,194,1704]" captionTargetPageId="16" captionText="Figures 83–93. SEM images of selected characters. 83–85. Male prosomata in frontal view: M. femoratus (83; arrow points to tiny anterior median eye, AME), M. toma sp. nov. (84), and M. cienaga sp. nov. (85). 86. Male prosoma, dorsal view (M. kiskeya sp. nov.). 87. Ocular area and carapace, frontodorsal view (M. cienaga sp. nov.). 88–91. Male cheliceral hairs: M. seguin sp. nov. (88), M. palvet sp. nov. (89), and M. toma sp. nov. (90, 91). 92, 93. Male gonopore, M. cienaga sp. nov. (92), M. palvet sp. nov. (93). Scale bars: 20 Mm (89); 40 Mm (90, 92, 93); 60 Mm (88, 91); 100 Mm (85, 87); 300 Mm (83, 84, 86)." figureDoi="http://doi.org/10.5281/zenodo.5438292" httpUri="https://zenodo.org/record/5438292/files/figure.png" pageId="42" pageNumber="286">85, 87, 92</figureCitation>
,
<figureCitation id="1341BE9C7A12741CDB22BF44FB524705" box="[1088,1187,1607,1628]" captionStart="Figures 94–103" captionStartId="17.[144,234,1678,1697]" captionTargetBox="[249,1319,194,1645]" captionTargetId="figure-0@17.[249,1319,194,1645]" captionTargetPageId="17" captionText="Figures 94–103. SEM images of selected characters. 94. Left procursus, retrolateral view (M. femoratus). 95. Right procursus and distal bulbal elements, retrolaterodorsal view (M. kiskeya sp. nov.). 96. Left palp, retrolateral view (M. toma sp. nov.). 97 Right procursus and distal bulbal elements, distal view (M. toma sp. nov.). 98. Left procursus, retrolateral view (M. seguin sp. nov.). 99. Left procursus, retrolateral view (M. cuadro sp. nov.). 100. Right procursus and distal bulbal elements, distal view (M. cienaga sp. nov.). 101. Palpal tarsal organ (M. cienaga sp. nov.). 102. Bulb with putative sperm duct opening (M. cienaga sp. nov.). 103. Left procursus tip, retrolateral view (M. cienaga sp. nov.). Abbreviations: b, genital bulb; ba, bulbal apophysis; p, procursus. Scale bars: 8 Mm (101); 30 Mm (100, 102, 103); 60 Mm (94, 95, 97); 100 Mm (96, 98, 99)." figureDoi="http://doi.org/10.5281/zenodo.5438294" httpUri="https://zenodo.org/record/5438294/files/figure.png" pageId="42" pageNumber="286">100–103</figureCitation>
,
<figureCitation id="1341BE9C7A12741CDBD3BF44FB2A4705" box="[1201,1243,1607,1628]" captionStart="Figures 104–114" captionStartId="18.[164,254,1648,1667]" captionTargetBox="[164,1446,195,1619]" captionTargetId="figure-0@18.[164,1446,195,1619]" captionTargetPageId="18" captionText="Figures 104–114. SEM images of selected characters. 104–107. Epigynum of M. femoratus, ventral view, without plug (104), ventroposterior view, with plug (105), and ventral views, with plug (106, 107). 108–110. Epigynum of M. toma sp. nov., ventral view without plug (108), ventral view with plug (109), and ventroposterior view with plug (110). 111, 112. Epigynum of M. cuadro sp. nov., ventral (111) and ventroposterior (112) views. 113, 114. Epigyna of M. cienaga sp. nov. (113) and M. palvet sp. nov. (114), ventral views. Abbreviation: gp, ‘genital plug’. Scale bars: 100 Mm (104, 105, 107–114); 200 Mm (106)." figureDoi="http://doi.org/10.5281/zenodo.5438296" httpUri="https://zenodo.org/record/5438296/files/figure.png" pageId="42" pageNumber="286">113</figureCitation>
,
<figureCitation id="1341BE9C7A12741CDB8BBF44FABD4705" box="[1257,1356,1607,1628]" captionStart="Figures 179–184" captionStartId="42.[164,254,1332,1351]" captionTargetBox="[325,1284,196,1300]" captionTargetId="figure-185@42.[323,1284,196,1302]" captionTargetPageId="42" captionText="Figures 179–184. Diagnostic characters. 179–181. Modisimus mango sp. nov. 179. Left palp, retrolateral view. 180. Male chelicerae, frontal view. 181. Cleared epigynum, dorsal view. 182–184. Modisimus cienaga sp. nov. 182. Left palp, retrolateral view. 183. Male chelicerae, frontal view. 184. Cleared epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5438318" httpUri="https://zenodo.org/record/5438318/files/figure.png" pageId="42" pageNumber="286">182–184</figureCitation>
,
<figureCitation id="1341BE9C7A12741CDA38BF44FA754704" box="[1370,1412,1607,1629]" captionStart="Figure 200" captionStartId="48.[164,243,1038,1057]" captionTargetBox="[387,1219,650,1008]" captionTargetId="figure-362@48.[387,1219,650,1008]" captionTargetPageId="48" captionText="Figure 200. Known distributions of Modisimus cienaga sp. nov. and species that are not yet assigned to any species group." figureDoi="http://doi.org/10.5281/zenodo.5438332" httpUri="https://zenodo.org/record/5438332/files/figure.png" pageId="42" pageNumber="286">200</figureCitation>
)
</paragraph>
<paragraph id="8BC5A2197A12741CDC58BF87FB33464D" blockId="42.[826,1441,1666,1812]" pageId="42" pageNumber="286">
<emphasis id="B90E7E0B7A12741CDC58BF87FC8447C0" box="[826,885,1668,1689]" italics="true" pageId="42" pageNumber="286">Type:</emphasis>
<materialsCitation id="3B12A8447A12741CDC1DBF80FB4F464D" collectingDate="2005-11-09" collectionCode="ZFMK" collectorName="Male &amp; La Cienaga &amp; B. A. Huber" country="Dominican Republic" elevation="1000" latitude="19.05" location="La Vega Province" longLatPrecision="1273" longitude="-70.88333" pageId="42" pageNumber="286" specimenCode="DR 17" specimenCount="1" stateProvince="La Vega" typeStatus="holotype">
<collectorName id="268FC7CF7A12741CDC1DBF80FC4947C0" box="[895,952,1667,1689]" pageId="42" pageNumber="286">Male</collectorName>
<typeStatus id="54C11CBB7A12741CDCA3BF80FBD347C0" box="[961,1058,1667,1689]" pageId="42" pageNumber="286" type="holotype">holotype</typeStatus>
from near
<collectorName id="268FC7CF7A12741CDBCABF87FADC47C0" box="[1192,1325,1667,1689]" pageId="42" pageNumber="286">La Ciénaga</collectorName>
(
<geoCoordinate id="EE4EC4DE7A12741CDA5DBF87FA6D47C0" box="[1343,1436,1666,1689]" degrees="19" direction="north" minutes="03" orientation="latitude" pageId="42" pageNumber="286" precision="925" value="19.05">
19°03
<emphasis id="B90E7E0B7A12741CDAE2BF81FA7747C0" box="[1408,1414,1666,1689]" italics="true" pageId="42" pageNumber="286">′</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE4EC4DE7A12741CDC58BFA1FC6A47E1" box="[826,923,1697,1720]" degrees="70" direction="west" minutes="53" orientation="longitude" pageId="42" pageNumber="286" precision="925" value="-70.88333">
70°53
<emphasis id="B90E7E0B7A12741CDC19BFA2FC7047E1" box="[891,897,1697,1720]" italics="true" pageId="42" pageNumber="286">′</emphasis>
W
</geoCoordinate>
; ~
<elevation id="0057452A7A12741CDCD5BFA1FBBA47E1" box="[951,1099,1698,1720]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="286" unit="m" value="1000.0">
<quantity id="4C820FFC7A12741CDCD5BFA1FBFA47E1" box="[951,1035,1698,1720]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="42" pageNumber="286" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
),
<collectingRegion id="49BE6CFB7A12741CDB07BFA0FAC747E1" box="[1125,1334,1698,1720]" country="Dominican Republic" name="La Vega" pageId="42" pageNumber="286">La Vega Province</collectingRegion>
,
<collectingCountry id="F36DE2897A12741CDA27BFA1FC3E478E" name="Dominican Republic" pageId="42" pageNumber="286">Dominican Republic</collectingCountry>
; forest along river, among the dry leaves of a dead epiphyte on the ground,
<date id="FFC484D97A12741CDBB6BFDCFA5047AC" box="[1236,1441,1759,1781]" pageId="42" pageNumber="286" value="2005-11-09">
<collectingDate id="EF807D317A12741CDBB6BFDCFA5047AC" box="[1236,1441,1759,1781]" pageId="42" pageNumber="286" value="2005-11-09">9 November 2005</collectingDate>
</date>
(
<collectorName id="268FC7CF7A12741CDC21BFFDFC39464D" box="[835,968,1790,1812]" pageId="42" pageNumber="286">B.A. Huber</collectorName>
), in
<collectionCode id="ED6B3ADC7A12741CDC9DBFFDFBB8464A" box="[1023,1097,1790,1811]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="286" type="Museum">ZFMK</collectionCode>
(
<specimenCode id="DBDC0A627A12741CDB38BFFDFB56464D" box="[1114,1191,1790,1812]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="42" pageNumber="286" type="Museum">DR 17</specimenCode>
a)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC5A2197A12741CDC58BE39FB0D4636" blockId="42.[826,1441,1850,1903]" pageId="42" pageNumber="286">
<emphasis id="B90E7E0B7A12741CDC58BE39FC494616" box="[826,952,1850,1871]" italics="true" pageId="42" pageNumber="286">Etymology:</emphasis>
The species name refers to the
<typeStatus id="54C11CBB7A12741CDA49BE38FAAD4609" box="[1323,1372,1851,1872]" pageId="42" pageNumber="286">type</typeStatus>
locality; it is used as a noun in apposition.
</paragraph>
<paragraph id="8BC5A2197A13741DDFF2B9C5FEB3400F" blockId="43.[144,759,198,342]" pageId="43" pageNumber="287">
<emphasis id="B90E7E0B7A13741DDFF2B9C5FEF74182" box="[144,262,198,219]" italics="true" pageId="43" pageNumber="287">Diagnosis:</emphasis>
Small, short-legged species, similar to
<taxonomicName id="4C7AD99A7A13741DDDBAB9C4FF0741A0" authorityName="Simon" authorityYear="1893" class="Arachnida" family="Pholcidae" genus="Hedypsilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="43" pageNumber="287" phylum="Arthropoda" rank="species" species="culicinus">
<emphasis id="B90E7E0B7A13741DDDBAB9C4FF0741A0" italics="true" pageId="43" pageNumber="287">M. culicinus</emphasis>
</taxonomicName>
(carapace pattern,
<figureCitation id="1341BE9C7A13741DDE92B9E7FDB041A3" box="[496,577,228,250]" captionStart="Figures 17–37" captionStartId="13.[144,234,1662,1681]" captionTargetBox="[144,1426,194,1632]" captionTargetId="figure-0@13.[144,1426,194,1632]" captionTargetPageId="13" captionText="Figures 17–37. Habitus photos of males (except 33), showing typical and distinctive characters. 17, 18. M. femoratus. 19. M. jima sp. nov. 20. M. kiskeya sp. nov. 21, 22. M. angulatus sp. nov. 23, 24. M. epepye sp. nov. 25. M. mango sp. nov. 26. M. cienaga sp. nov. 27–29. M. seguin sp. nov. (arrows point to callus-like area). 30–34. M. vittatus. 35. M. palvet sp. nov. 36. M. berac sp. nov. 37. M. paraiso sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5438284" httpUri="https://zenodo.org/record/5438284/files/figure.png" pageId="43" pageNumber="287">Fig. 26</figureCitation>
; palpal femur,
<figureCitation id="1341BE9C7A13741DDFF2B800FF1E4040" box="[144,239,259,281]" captionStart="Figures 179–184" captionStartId="42.[164,254,1332,1351]" captionTargetBox="[325,1284,196,1300]" captionTargetId="figure-185@42.[323,1284,196,1302]" captionTargetPageId="42" captionText="Figures 179–184. Diagnostic characters. 179–181. Modisimus mango sp. nov. 179. Left palp, retrolateral view. 180. Male chelicerae, frontal view. 181. Cleared epigynum, dorsal view. 182–184. Modisimus cienaga sp. nov. 182. Left palp, retrolateral view. 183. Male chelicerae, frontal view. 184. Cleared epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5438318" httpUri="https://zenodo.org/record/5438318/files/figure.png" pageId="43" pageNumber="287">Fig. 182</figureCitation>
), but male chelicerae (
<figureCitation id="1341BE9C7A13741DDD73B800FD9C4040" box="[529,621,259,281]" captionStart="Figures 179–184" captionStartId="42.[164,254,1332,1351]" captionTargetBox="[325,1284,196,1300]" captionTargetId="figure-185@42.[323,1284,196,1302]" captionTargetPageId="42" captionText="Figures 179–184. Diagnostic characters. 179–181. Modisimus mango sp. nov. 179. Left palp, retrolateral view. 180. Male chelicerae, frontal view. 181. Cleared epigynum, dorsal view. 182–184. Modisimus cienaga sp. nov. 182. Left palp, retrolateral view. 183. Male chelicerae, frontal view. 184. Cleared epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5438318" httpUri="https://zenodo.org/record/5438318/files/figure.png" pageId="43" pageNumber="287">Fig. 183</figureCitation>
) and male clypeus unmodified, distinctive procursus tip (
<figureCitation id="1341BE9C7A13741DDFF5B843FEC4400F" box="[151,309,320,342]" captionStart="Figures 94–103" captionStartId="17.[144,234,1678,1697]" captionTargetBox="[249,1319,194,1645]" captionTargetId="figure-0@17.[249,1319,194,1645]" captionTargetPageId="17" captionText="Figures 94–103. SEM images of selected characters. 94. Left procursus, retrolateral view (M. femoratus). 95. Right procursus and distal bulbal elements, retrolaterodorsal view (M. kiskeya sp. nov.). 96. Left palp, retrolateral view (M. toma sp. nov.). 97 Right procursus and distal bulbal elements, distal view (M. toma sp. nov.). 98. Left procursus, retrolateral view (M. seguin sp. nov.). 99. Left procursus, retrolateral view (M. cuadro sp. nov.). 100. Right procursus and distal bulbal elements, distal view (M. cienaga sp. nov.). 101. Palpal tarsal organ (M. cienaga sp. nov.). 102. Bulb with putative sperm duct opening (M. cienaga sp. nov.). 103. Left procursus tip, retrolateral view (M. cienaga sp. nov.). Abbreviations: b, genital bulb; ba, bulbal apophysis; p, procursus. Scale bars: 8 Mm (101); 30 Mm (100, 102, 103); 60 Mm (94, 95, 97); 100 Mm (96, 98, 99)." figureDoi="http://doi.org/10.5281/zenodo.5438294" httpUri="https://zenodo.org/record/5438294/files/figure.png" pageId="43" pageNumber="287">Figs 100, 103</figureCitation>
).
</paragraph>
<paragraph id="8BC5A2197A13741DDFF2B87CFD3345CB" blockId="43.[144,759,383,1170]" pageId="43" pageNumber="287">
<emphasis id="B90E7E0B7A13741DDFF2B87CFEBD40CD" box="[144,332,383,404]" italics="true" pageId="43" pageNumber="287">
Male (
<typeStatus id="54C11CBB7A13741DDFBDB87CFECF40CD" box="[223,318,383,404]" pageId="43" pageNumber="287" type="holotype">holotype</typeStatus>
):
</emphasis>
Total length, 1.2; carapace width, 0.55. Leg 1: 5.2 (1.4 + 0.2 + 1.4 + 1.6 + 0.5); tibia 2, 1.4; tibia 3, 0.7; tibia 4, 1.0. Tibia 1 L/d: 23. Habitus as in
<figureCitation id="1341BE9C7A13741DDFF2B8D8FEF140A8" box="[144,256,475,497]" captionStart="Figures 17–37" captionStartId="13.[144,234,1662,1681]" captionTargetBox="[144,1426,194,1632]" captionTargetId="figure-0@13.[144,1426,194,1632]" captionTargetPageId="13" captionText="Figures 17–37. Habitus photos of males (except 33), showing typical and distinctive characters. 17, 18. M. femoratus. 19. M. jima sp. nov. 20. M. kiskeya sp. nov. 21, 22. M. angulatus sp. nov. 23, 24. M. epepye sp. nov. 25. M. mango sp. nov. 26. M. cienaga sp. nov. 27–29. M. seguin sp. nov. (arrows point to callus-like area). 30–34. M. vittatus. 35. M. palvet sp. nov. 36. M. berac sp. nov. 37. M. paraiso sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5438284" httpUri="https://zenodo.org/record/5438284/files/figure.png" pageId="43" pageNumber="287">Figure 26</figureCitation>
, carapace light ochre-yellow, with three pairs of distinct dark spots and dark median Y-shaped mark, clypeus with a pair of brown stripes, narrowing towards chelicerae, sternum whitish; legs light ochreyellow, very indistinct darker rings subdistally on tibiae and femora, tips of femora and tibiae whitish; abdomen bluish grey with large black spots in a row on each side of heart and laterally; genital area light brown. Ocular area moderately elevated (
<figureCitation id="1341BE9C7A13741DDDFBBBD3FD1843BC" box="[665,745,719,742]" captionStart="Figures 83–93" captionStartId="16.[164,254,1736,1755]" captionTargetBox="[258,1348,194,1704]" captionTargetId="figure-0@16.[258,1348,194,1704]" captionTargetPageId="16" captionText="Figures 83–93. SEM images of selected characters. 83–85. Male prosomata in frontal view: M. femoratus (83; arrow points to tiny anterior median eye, AME), M. toma sp. nov. (84), and M. cienaga sp. nov. (85). 86. Male prosoma, dorsal view (M. kiskeya sp. nov.). 87. Ocular area and carapace, frontodorsal view (M. cienaga sp. nov.). 88–91. Male cheliceral hairs: M. seguin sp. nov. (88), M. palvet sp. nov. (89), and M. toma sp. nov. (90, 91). 92, 93. Male gonopore, M. cienaga sp. nov. (92), M. palvet sp. nov. (93). Scale bars: 20 Mm (89); 40 Mm (90, 92, 93); 60 Mm (88, 91); 100 Mm (85, 87); 300 Mm (83, 84, 86)." figureDoi="http://doi.org/10.5281/zenodo.5438292" httpUri="https://zenodo.org/record/5438292/files/figure.png" pageId="43" pageNumber="287">Fig. 85</figureCitation>
); thoracic furrow distinct (
<figureCitation id="1341BE9C7A13741DDED0BBEDFDF2425D" box="[434,515,750,772]" captionStart="Figures 83–93" captionStartId="16.[164,254,1736,1755]" captionTargetBox="[258,1348,194,1704]" captionTargetId="figure-0@16.[258,1348,194,1704]" captionTargetPageId="16" captionText="Figures 83–93. SEM images of selected characters. 83–85. Male prosomata in frontal view: M. femoratus (83; arrow points to tiny anterior median eye, AME), M. toma sp. nov. (84), and M. cienaga sp. nov. (85). 86. Male prosoma, dorsal view (M. kiskeya sp. nov.). 87. Ocular area and carapace, frontodorsal view (M. cienaga sp. nov.). 88–91. Male cheliceral hairs: M. seguin sp. nov. (88), M. palvet sp. nov. (89), and M. toma sp. nov. (90, 91). 92, 93. Male gonopore, M. cienaga sp. nov. (92), M. palvet sp. nov. (93). Scale bars: 20 Mm (89); 40 Mm (90, 92, 93); 60 Mm (88, 91); 100 Mm (85, 87); 300 Mm (83, 84, 86)." figureDoi="http://doi.org/10.5281/zenodo.5438292" httpUri="https://zenodo.org/record/5438292/files/figure.png" pageId="43" pageNumber="287">Fig. 87</figureCitation>
). PME–PME, 45 Mm; PME diameter, 55 Mm; PME–ALE, 45 Mm; no AME. Sternum wider than long (0.4/0.3), unmodified. Chelicerae unmodified (
<figureCitation id="1341BE9C7A13741DDE17BA49FE254239" box="[373,468,842,864]" captionStart="Figures 179–184" captionStartId="42.[164,254,1332,1351]" captionTargetBox="[325,1284,196,1300]" captionTargetId="figure-185@42.[323,1284,196,1302]" captionTargetPageId="42" captionText="Figures 179–184. Diagnostic characters. 179–181. Modisimus mango sp. nov. 179. Left palp, retrolateral view. 180. Male chelicerae, frontal view. 181. Cleared epigynum, dorsal view. 182–184. Modisimus cienaga sp. nov. 182. Left palp, retrolateral view. 183. Male chelicerae, frontal view. 184. Cleared epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5438318" httpUri="https://zenodo.org/record/5438318/files/figure.png" pageId="43" pageNumber="287">Fig. 183</figureCitation>
). Palps as in
<figureCitation id="1341BE9C7A13741DDD16BA49FD034206" box="[628,754,842,864]" captionStart="Figures 179–184" captionStartId="42.[164,254,1332,1351]" captionTargetBox="[325,1284,196,1300]" captionTargetId="figure-185@42.[323,1284,196,1302]" captionTargetPageId="42" captionText="Figures 179–184. Diagnostic characters. 179–181. Modisimus mango sp. nov. 179. Left palp, retrolateral view. 180. Male chelicerae, frontal view. 181. Cleared epigynum, dorsal view. 182–184. Modisimus cienaga sp. nov. 182. Left palp, retrolateral view. 183. Male chelicerae, frontal view. 184. Cleared epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5438318" httpUri="https://zenodo.org/record/5438318/files/figure.png" pageId="43" pageNumber="287">Figure 182</figureCitation>
, coxa with light retrolateral apophysis, femur with rounded basal and more pointed distal ventral apophyses, procursus complex distally, with membranous and sclerotized structures (
<figureCitation id="1341BE9C7A13741DDEB3BAC7FD9F4283" box="[465,622,964,986]" captionStart="Figures 94–103" captionStartId="17.[144,234,1678,1697]" captionTargetBox="[249,1319,194,1645]" captionTargetId="figure-0@17.[249,1319,194,1645]" captionTargetPageId="17" captionText="Figures 94–103. SEM images of selected characters. 94. Left procursus, retrolateral view (M. femoratus). 95. Right procursus and distal bulbal elements, retrolaterodorsal view (M. kiskeya sp. nov.). 96. Left palp, retrolateral view (M. toma sp. nov.). 97 Right procursus and distal bulbal elements, distal view (M. toma sp. nov.). 98. Left procursus, retrolateral view (M. seguin sp. nov.). 99. Left procursus, retrolateral view (M. cuadro sp. nov.). 100. Right procursus and distal bulbal elements, distal view (M. cienaga sp. nov.). 101. Palpal tarsal organ (M. cienaga sp. nov.). 102. Bulb with putative sperm duct opening (M. cienaga sp. nov.). 103. Left procursus tip, retrolateral view (M. cienaga sp. nov.). Abbreviations: b, genital bulb; ba, bulbal apophysis; p, procursus. Scale bars: 8 Mm (101); 30 Mm (100, 102, 103); 60 Mm (94, 95, 97); 100 Mm (96, 98, 99)." figureDoi="http://doi.org/10.5281/zenodo.5438294" httpUri="https://zenodo.org/record/5438294/files/figure.png" pageId="43" pageNumber="287">Figs 100, 103</figureCitation>
), bulb with simple distal apophysis and membranous dorsal structure (
<figureCitation id="1341BE9C7A13741DDE6EBD01FE9A4541" box="[268,363,1026,1048]" captionStart="Figures 94–103" captionStartId="17.[144,234,1678,1697]" captionTargetBox="[249,1319,194,1645]" captionTargetId="figure-0@17.[249,1319,194,1645]" captionTargetPageId="17" captionText="Figures 94–103. SEM images of selected characters. 94. Left procursus, retrolateral view (M. femoratus). 95. Right procursus and distal bulbal elements, retrolaterodorsal view (M. kiskeya sp. nov.). 96. Left palp, retrolateral view (M. toma sp. nov.). 97 Right procursus and distal bulbal elements, distal view (M. toma sp. nov.). 98. Left procursus, retrolateral view (M. seguin sp. nov.). 99. Left procursus, retrolateral view (M. cuadro sp. nov.). 100. Right procursus and distal bulbal elements, distal view (M. cienaga sp. nov.). 101. Palpal tarsal organ (M. cienaga sp. nov.). 102. Bulb with putative sperm duct opening (M. cienaga sp. nov.). 103. Left procursus tip, retrolateral view (M. cienaga sp. nov.). Abbreviations: b, genital bulb; ba, bulbal apophysis; p, procursus. Scale bars: 8 Mm (101); 30 Mm (100, 102, 103); 60 Mm (94, 95, 97); 100 Mm (96, 98, 99)." figureDoi="http://doi.org/10.5281/zenodo.5438294" httpUri="https://zenodo.org/record/5438294/files/figure.png" pageId="43" pageNumber="287">Fig. 102</figureCitation>
). Legs without spines and curved hairs; no short vertical hairs on femora; retrolateral trichobothrium on tibia 1 at 31%; prolateral trichobothrium apparently absent on tibia 1, but present on other tibiae; tarsus 1 with ~15 pseudosegments.
</paragraph>
<paragraph id="8BC5A2197A13741DDFF2BDB8FD2C4589" blockId="43.[144,733,1211,1233]" box="[144,733,1211,1233]" pageId="43" pageNumber="287">
<emphasis id="B90E7E0B7A13741DDFF2BDB8FEF14589" box="[144,256,1211,1232]" italics="true" pageId="43" pageNumber="287">Variation:</emphasis>
Tibia
<quantity id="4C820FFC7A13741DDE2DBDB8FE8C4588" box="[335,381,1211,1233]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="43" pageNumber="287" unit="in" value="1.0">1 in</quantity>
two other males: 1.3 and 1.4.
</paragraph>
<paragraph id="8BC5A2197A13741DDFF2BDFAFD5844F1" blockId="43.[144,758,1273,1448]" pageId="43" pageNumber="287">
<emphasis id="B90E7E0B7A13741DDFF2BDFAFF184457" box="[144,233,1273,1294]" italics="true" pageId="43" pageNumber="287">Female:</emphasis>
In general similar to male, but dark rings on legs more distinct, and with additional rings on femora (proximally and medially) and tibiae (proximally). Tibia
<quantity id="4C820FFC7A13741DDE55BC56FE9B4432" box="[311,362,1365,1387]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="43" pageNumber="287" unit="in" value="1.0">1 in</quantity>
<specimenCount id="9D7C69907A13741DDE1ABC55FDD34432" box="[376,546,1365,1387]" pageId="43" pageNumber="287" type="female">seven females</specimenCount>
: 0.96–1.10 (mean 1.03). Epigynum, very simple externally; dorsal view as in
<figureCitation id="1341BE9C7A13741DDFB1BC91FEA044F1" box="[211,337,1426,1448]" captionStart="Figures 179–184" captionStartId="42.[164,254,1332,1351]" captionTargetBox="[325,1284,196,1300]" captionTargetId="figure-185@42.[323,1284,196,1302]" captionTargetPageId="42" captionText="Figures 179–184. Diagnostic characters. 179–181. Modisimus mango sp. nov. 179. Left palp, retrolateral view. 180. Male chelicerae, frontal view. 181. Cleared epigynum, dorsal view. 182–184. Modisimus cienaga sp. nov. 182. Left palp, retrolateral view. 183. Male chelicerae, frontal view. 184. Cleared epigynum, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5438318" httpUri="https://zenodo.org/record/5438318/files/figure.png" pageId="43" pageNumber="287">Figure 184</figureCitation>
, with contiguous pore plates.
</paragraph>
<paragraph id="8BC5A2197A13741DDFF2BCD2FEF5475F" blockId="43.[144,758,1489,1542]" pageId="43" pageNumber="287">
<emphasis id="B90E7E0B7A13741DDFF2BCD2FED044BF" box="[144,289,1489,1510]" italics="true" pageId="43" pageNumber="287">Distribution:</emphasis>
Known from
<typeStatus id="54C11CBB7A13741DDD66BCD1FDC444BE" box="[516,565,1490,1511]" pageId="43" pageNumber="287">type</typeStatus>
locality only (
<figureCitation id="1341BE9C7A13741DDFFABCF3FF09475F" box="[152,248,1520,1542]" captionStart="Figure 200" captionStartId="48.[164,243,1038,1057]" captionTargetBox="[387,1219,650,1008]" captionTargetId="figure-362@48.[387,1219,650,1008]" captionTargetPageId="48" captionText="Figure 200. Known distributions of Modisimus cienaga sp. nov. and species that are not yet assigned to any species group." figureDoi="http://doi.org/10.5281/zenodo.5438332" httpUri="https://zenodo.org/record/5438332/files/figure.png" pageId="43" pageNumber="287">Fig. 200</figureCitation>
).
</paragraph>
<paragraph id="8BC5A2197A13741DDFF2BF2DFD2947E6" blockId="43.[144,759,1582,1727]" pageId="43" pageNumber="287">
<emphasis id="B90E7E0B7A13741DDFF2BF2DFE86471A" box="[144,375,1582,1603]" italics="true" pageId="43" pageNumber="287">Material examined:</emphasis>
<materialsCitation id="3B12A8447A13741DDEEABF2DFD5E473A" collectorName="La Cienaga" country="Dominican Republic" location="La Vega Province" pageId="43" pageNumber="287" specimenCount="1" specimenCount-male="1" stateProvince="La Vega" typeStatus="holotype">
<collectingCountry id="F36DE2897A13741DDEEABF2DFD8D471D" box="[392,636,1582,1604]" name="Dominican Republic" pageId="43" pageNumber="287">Dominican Republic</collectingCountry>
:
<collectingRegion id="49BE6CFB7A13741DDDF2BF2CFF07473A" country="Dominican Republic" name="La Vega" pageId="43" pageNumber="287">La Vega Province</collectingRegion>
, near
<collectorName id="268FC7CF7A13741DDE5FBF4DFE33473A" box="[317,450,1613,1635]" pageId="43" pageNumber="287">La Ciénaga</collectorName>
,
<specimenCount id="9D7C69907A13741DDEACBF4EFE04473A" box="[462,501,1613,1635]" pageId="43" pageNumber="287" type="male">1♂</specimenCount>
,
<typeStatus id="54C11CBB7A13741DDD61BF4EFD95473A" box="[515,612,1613,1635]" pageId="43" pageNumber="287" type="holotype">holotype</typeStatus>
above
</materialsCitation>
;
<materialsCitation id="3B12A8447A13741DDDD9BF4DFDF847F9" collectionCode="ZFMK" collectorName="La Cienaga" country="Dominican Republic" location="La Vega Province" pageId="43" pageNumber="287" specimenCount="10" specimenCount-female="7" specimenCount-male="3" stateProvince="La Vega Province" typeStatus="holotype">
same data,
<specimenCount id="9D7C69907A13741DDFB7BF6FFF0D47D8" box="[213,252,1643,1665]" pageId="43" pageNumber="287" type="male">2♂</specimenCount>
and
<specimenCount id="9D7C69907A13741DDE5CBF6FFE9147D8" box="[318,352,1643,1665]" pageId="43" pageNumber="287" type="female">7♀</specimenCount>
(
<collectionCode id="ED6B3ADC7A13741DDE11BF6FFE3047D8" box="[371,449,1644,1665]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="43" pageNumber="287" type="Museum">ZFMK</collectionCode>
;
<specimenCount id="9D7C69907A13741DDEB2BF6FFE0647D8" box="[464,503,1643,1665]" pageId="43" pageNumber="287" type="male">1♂</specimenCount>
used for SEM; chelicerae of other male lost; DR 17)
</materialsCitation>
;
<materialsCitation id="3B12A8447A13741DDD75BF88FD2547E6" collectionCode="ZFMK" collectorName="La Cienaga" country="Dominican Republic" location="La Vega Province" pageId="43" pageNumber="287" specimenCode="DR 100-13" specimenCount="4" specimenCount-female="3" specimenCount-juvenile="1" stateProvince="La Vega Province" typeStatus="holotype">
same data,
<specimenCount id="9D7C69907A13741DDDC2BF89FD3347F9" box="[672,706,1674,1696]" pageId="43" pageNumber="287" type="female">3♀</specimenCount>
and
<specimenCount id="9D7C69907A13741DDFF2BFAAFEEF47E6" box="[144,286,1705,1727]" pageId="43" pageNumber="287" type="juvenile">one juvenile</specimenCount>
, in pure ethanol (
<collectionCode id="ED6B3ADC7A13741DDE90BFAAFDB147E7" box="[498,576,1705,1726]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="43" pageNumber="287" type="Museum">ZFMK</collectionCode>
,
<specimenCode id="DBDC0A627A13741DDD2CBFAAFD3D47E6" box="[590,716,1705,1727]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="43" pageNumber="287" type="Museum">DR 100-13</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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