115 lines
13 KiB
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115 lines
13 KiB
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<document id="35414693F96BC818E72EEF24BEDA5F03" ID="10.11646/zootaxa.4271.1.1" ID-CLB-Dataset="32974" ID-DOI="10.11646/zootaxa.4271.1.1" ID-GBIF-Dataset="f3ad9b4c-35d8-4a01-a14b-3dd968623358" ID-ISSN="1175-5326" ID-Zenodo-Dep="583192" ID-ZooBank="50496AC4-D639-49A7-9249-386B037DAE72" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495697318696" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2017" docId="0393E60DFFB3FF9FD2B08F3CEDE2740F" docLanguage="en" docName="zootaxa.4271.1.1SLASH11141.pdf" docOrigin="Zootaxa 4271 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Goniasteridae" docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="FFAA9E75FFB7FF9BD2278D50ED097120" masterDocTitle="Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="5" updateTime="1698439914132" updateUser="ExternalLinkService">
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<mods:title id="7026E6A80CAF06180B67DEB2EAF5C5F9">Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species</mods:title>
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<mods:name id="3B579B8F25234753096C1CF80FE6632D" type="personal">
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<mods:namePart id="839B177E67697FE9404A3A9E5E642E21">Mah, Christopher L.</mods:namePart>
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<mods:typeOfResource id="6161A055E739B9DBA28436E90F7005D5">text</mods:typeOfResource>
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<mods:title id="4D284213E99E9045B56E2DAD786AB7FD">Zootaxa</mods:title>
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<mods:part id="BC98C117895A3B65FD54060C21383ED4">
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<mods:date id="D1B6BB124351E54F2EA1B808FF903FCE">2017</mods:date>
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<mods:detail id="7625A07448D2554173382390589ADBE7" type="volume">
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<mods:number id="DBED911C9A433A1962092AC7F65DF569">4271</mods:number>
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<mods:number id="6092BBF244BF2D945A4FFF25F836D773">1</mods:number>
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<mods:classification id="06CA2E610E08CFA2CAB6D570DC6AA271">journal article</mods:classification>
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<mods:identifier id="B8C8AD43D6FF22F45A69E885A0D6AA65" type="CLB-Dataset">32974</mods:identifier>
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<mods:identifier id="00662023A66C302A0C675F93CF837DCB" type="DOI">10.11646/zootaxa.4271.1.1</mods:identifier>
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<treatment id="0393E60DFFB3FF9FD2B08F3CEDE2740F" ID-DOI="http://doi.org/10.5281/zenodo.6016973" ID-GBIF-Taxon="130844941" ID-Zenodo-Dep="6016973" LSID="urn:lsid:plazi:treatment:0393E60DFFB3FF9FD2B08F3CEDE2740F" httpUri="http://treatment.plazi.org/id/0393E60DFFB3FF9FD2B08F3CEDE2740F" lastPageNumber="5" pageId="4" pageNumber="5">
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<subSubSection id="C3200490FFB3FF9FD2B08F3CE96D7217" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="8B85571BFFB3FF9FD2B08F3CEC7673A6" blockId="4.[151,383,620,646]" box="[151,383,620,646]" pageId="4" pageNumber="5">
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<heading id="D0CDE077FFB3FF9FD2B08F3CEC7673A6" bold="true" box="[151,383,620,646]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
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<emphasis id="B94E8B09FFB3FF9FD2B08F3CEC7673A6" bold="true" box="[151,383,620,646]" pageId="4" pageNumber="5">
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The
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<taxonomicName id="4C3A2C98FFB3FF9FD2E88F3CEC7673A6" ID-CoL="8RYP7" box="[207,383,620,646]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B85571BFFB3FF9FD2B08FE2E96D7217" blockId="4.[151,1437,690,1327]" pageId="4" pageNumber="5">
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Goniasterids occur primarily at continental shelf depths with many taxa extending below
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<quantity id="4CC2FAFEFFB3FF9FD6B38FE3E9D573EA" box="[1172,1244,691,715]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="m" value="200.0">200 m</quantity>
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(
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<bibRefCitation id="EFAB2AEAFFB3FF9FD6CB8FE2EDDD73CF" author="Mah" pageId="4" pageNumber="5" refString="Mah, C. & Blake, D. B. (2012) Global Diversity and Phylogeny of the Asteroidea (Echinodermata). PLoS ONE, 7 (4), e 35644. http: // dx. doi. org / 10.1371 / journal. pone. 0035644" type="journal article" year="2012">Mah and Blake 2012</bibRefCitation>
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). Exceptionally, some taxa occur in shallow or intertidal settings and below
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<quantity id="4CC2FAFEFFB3FF9FD6098F87E98D73CE" box="[1070,1156,727,751]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="5" unit="m" value="1000.0">1000 m</quantity>
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in lower bathyal/abyssal settings (e.g., Mah 2015). Ecologically, goniasterids vary from detritivores, such as
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<emphasis id="B94E8B09FFB3FF9FD6B58FABE87E7233" box="[1170,1399,763,787]" italics="true" pageId="4" pageNumber="5">
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<taxonomicName id="4C3A2C98FFB3FF9FD6B58FABE87D7233" box="[1170,1396,763,787]" class="Asteroidea" family="Goniasteridae" genus="Mediaster" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="species" species="aequalis">Mediaster aequalis</taxonomicName>
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,
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</emphasis>
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to predators, such as the corallivorous
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<taxonomicName id="4C3A2C98FFB3FF9FD00D8E4FEFB97216" box="[554,688,799,822]" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFB3FF9FD00D8E4FEFB97216" box="[554,688,799,822]" italics="true" pageId="4" pageNumber="5">Hippasteria</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C3A2C98FFB3FF9FD0C18E4EEE787216" box="[742,881,798,822]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFB3FF9FD0C18E4EEE787216" box="[742,881,798,822]" italics="true" pageId="4" pageNumber="5">Evoplosoma</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EFAB2AEAFFB3FF9FD1588E4EE9087217" author="Mah" box="[895,1025,798,823]" pageId="4" pageNumber="5" refString="Mah, C. L. (2015 a) A new Atlantic species of Evoplosoma with taxonomic summary and in situ observations of Atlantic deep - sea corallivorous Goniasteridae (Valvatida; Asteroidea). Marine Biodiversity Records, 8 (e 5), 1 - 8. http: // dx. doi. org / 10.1017 / S 1755267214001407" type="journal article" year="2015" yearSuffix="a">Mah 2015a</bibRefCitation>
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,
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<bibRefCitation id="EFAB2AEAFFB3FF9FD62A8E4EE9517217" author="Mah" box="[1037,1112,798,823]" pageId="4" pageNumber="5" refString="Mah, C. L. (2015 b) New species, corallivory, in situ video observations and overview of the Goniasteridae (Valvatida, Asteroidea) in the Hawaiian Region. Zootaxa, 3926 (2), 211 - 228." type="journal article" year="2015" yearSuffix="b">2015b</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3200490FFB3FF9FD2E08E13EDE2740F" pageId="4" pageNumber="5" type="description">
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<paragraph id="8B85571BFFB3FF9FD2E08E13ECAF752F" blockId="4.[151,1437,690,1327]" pageId="4" pageNumber="5">
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Most
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<taxonomicName id="4C3A2C98FFB3FF9FD32A8E12ECA0727B" box="[269,425,834,859]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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are identified by a heavily calcified endoskeleton, a marginal plate series of blocky ossicles forming a distinct border around the periphery of the body, papulae limited to the abactinal surface, and a large disk with a relatively flattened body shape. This fundamental body form has been observed in other families in the
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<taxonomicName id="4C3A2C98FFB3FF9FD2D08EFEEC6D72E7" box="[247,356,942,967]" class="Asteroidea" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="order">Valvatida</taxonomicName>
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, including the
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<taxonomicName id="4C3A2C98FFB3FF9FD0168EFEEFEC72E7" box="[561,741,942,967]" class="Asteroidea" family="Ophidiasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Ophidiasteridae</taxonomicName>
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,
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<taxonomicName id="4C3A2C98FFB3FF9FD0D98EFEEE8672E7" box="[766,911,942,967]" class="Asteroidea" family="Oreasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Oreasteridae</taxonomicName>
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,
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<taxonomicName id="4C3A2C98FFB3FF9FD1818EFEE95972E7" box="[934,1104,942,967]" class="Asteroidea" family="Odontasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Odontasteridae</taxonomicName>
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among others, suggesting plesiomorphy and/or convergence (e.g.,
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<bibRefCitation id="EFAB2AEAFFB3FF9FD05B8E82EE0072CB" author="Blake" box="[636,777,978,1003]" pageId="4" pageNumber="5" refString="Blake, D. B. (1987) Classification and phylogeny of post - Paleozoic sea stars (Asteroidea: Echinodermata). Journal of Natural History, 21, 481 - 528." type="journal article" year="1987">Blake 1987</bibRefCitation>
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). Limits of the goniasterid body form have not been comprehensively tested.
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</paragraph>
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<paragraph id="8B85571BFFB3FF9FD2E0894AEDE2740F" blockId="4.[151,1437,690,1327]" pageId="4" pageNumber="5">
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Although goniasterid monophyly has been at least partially upheld by molecular data (
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<bibRefCitation id="EFAB2AEAFFB3FF9FD684894AE8997513" author="Mah" box="[1187,1424,1050,1075]" pageId="4" pageNumber="5" refString="Mah, C. L. & Foltz, D. W. (2011) Molecular Phylogeny of the Valvatacea (Asteroidea, Echinodermata). Zoological Journal of the Linnean Society, 161, 769 - 788." type="journal article" year="2011">Mah and Foltz 2011</bibRefCitation>
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), numerous goniasterid genera display morphology intermediate with other families (
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<taxonomicName id="4C3A2C98FFB3FF9FD661896EE9F57577" box="[1094,1276,1086,1111]" class="Asteroidea" family="Ophidiasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Ophidiasteridae</taxonomicName>
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,
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<taxonomicName id="4C3A2C98FFB3FF9FD72F896EE8917577" box="[1288,1432,1086,1111]" class="Asteroidea" family="Oreasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Oreasteridae</taxonomicName>
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, etc.) within the Valvatacea suggesting shared ancestries.
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<taxonomicName id="4C3A2C98FFB3FF9FD10C8933EE9C755B" box="[811,917,1123,1147]" class="Asteroidea" family="Oreasteridae" genus="Anthenea" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFB3FF9FD10C8933EE9C755B" box="[811,917,1123,1147]" italics="true" pageId="4" pageNumber="5">Anthenea</emphasis>
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</taxonomicName>
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and
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<emphasis id="B94E8B09FFB3FF9FD1F58933E97B755B" box="[978,1138,1123,1147]" italics="true" pageId="4" pageNumber="5">Goniodiscaste</emphasis>
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r, for example, have been assigned to both the
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<taxonomicName id="4C3A2C98FFB3FF9FD35A89D6EF1075BF" box="[381,537,1158,1183]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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and the
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<taxonomicName id="4C3A2C98FFB3FF9FD05F89D6EE0C75BF" box="[632,773,1158,1183]" class="Asteroidea" family="Oreasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Oreasteridae</taxonomicName>
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by different authors. As discussed below, several members of the
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<taxonomicName id="4C3A2C98FFB3FF9FD2C989FAEC9675E3" box="[238,415,1194,1219]" class="Asteroidea" family="Ophidiasteridae" kingdom="Animalia" order="Valvatida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="family">Ophidiasteridae</taxonomicName>
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have been historically considered as goniasterid-like, although these have not been considered as contentious as those taxa intermediate between goniasterids and oreasterids, the discovery of specimens displaying ophidiasterid/goniasterid characters challenges accepted taxonomic definitions for these groups.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |