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<mods:title id="1C6C929E9DF4272AF28CBEA076D77B6F">Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species</mods:title>
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<treatment id="0393E60DFFA7FF8AD2B0880FE98D7218" ID-DOI="http://doi.org/10.5281/zenodo.6016985" ID-GBIF-Taxon="130844949" ID-Zenodo-Dep="6016985" LSID="urn:lsid:plazi:treatment:0393E60DFFA7FF8AD2B0880FE98D7218" httpUri="http://treatment.plazi.org/id/0393E60DFFA7FF8AD2B0880FE98D7218" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<subSubSection id="C3200490FFA7FF8BD2B0880FEC9574FB" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B85571BFFA7FF8BD2B0880FEC8A745A" blockId="16.[151,387,1375,1402]" box="[151,387,1375,1402]" pageId="16" pageNumber="17">
<heading id="D0CDE077FFA7FF8BD2B0880FEC8A745A" bold="true" box="[151,387,1375,1402]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C3A2C98FFA7FF8BD2B0880FEC8A745A" authority="Gray 1840" authorityName="Gray" authorityYear="1840" box="[151,387,1375,1402]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA7FF8BD2B0880FEC8A745A" bold="true" box="[151,387,1375,1402]" pageId="16" pageNumber="17">
<emphasis id="B94E8B09FFA7FF8BD2B0880FEDF07459" bold="true" box="[151,249,1375,1401]" italics="true" pageId="16" pageNumber="17">Ferdina</emphasis>
<bibRefCitation id="EFAB2AEAFFA7FF8BD3278830EC8A745A" author="Gray" box="[256,387,1376,1402]" pageId="16" pageNumber="17" refString="Gray, J. E. (1840) A synopsis of the genera and species of the class Hypostoma (Asterias Linnaeus). Annals of the Magazine of Natural History, 6, 175 - 184, 275 - 290." type="journal article" year="1840">Gray 1840</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B85571BFFA7FF8BD2B088F6EC9574FB" blockId="16.[151,1435,1446,1499]" pageId="16" pageNumber="17">
<bibRefCitation id="EFAB2AEAFFA7FF8BD2B088F6EC03749C" author="Gray" box="[151,266,1446,1468]" pageId="16" pageNumber="17" refString="Gray, J. E. (1840) A synopsis of the genera and species of the class Hypostoma (Asterias Linnaeus). Annals of the Magazine of Natural History, 6, 175 - 184, 275 - 290." type="journal article" year="1840">Gray, 1840</bibRefCitation>
: 282; 1866: 1213;
<bibRefCitation id="EFAB2AEAFFA7FF8BD3F488F6EF47749C" author="Fisher" box="[467,590,1446,1468]" pageId="16" pageNumber="17" refString="Fisher, W. K. (1919) Starfishes of the Philippine seas and adjacent waters. Bulletin of the United States National Museum, 3 (100), 1 - 547, 156 pls." type="journal article" year="1919">Fisher 1919</bibRefCitation>
: 370; H.L. Clark 1921: 58;
<bibRefCitation id="EFAB2AEAFFA7FF8BD14488F6E911749C" author="Livingstone" box="[867,1048,1446,1468]" pageId="16" pageNumber="17" refString="Livingstone, A. (1931) On the restriction of the genus Ferdina Gray (Asteroidea). Australian Zoologist, 6 (4), 305 - 309." type="journal article" year="1931">Livingstone 1931</bibRefCitation>
: 305;
<bibRefCitation id="EFAB2AEAFFA7FF8BD67288F7E80C749C" author="Clark" box="[1109,1285,1446,1468]" pageId="16" pageNumber="17" refString="Clark, A. M. (1967) Notes on asteroids in the British Museum (Natural History) V. Nardoa and some other ophidiasterids. Bulletin of the British Museum (Natural History) Zoology, 15 (4), 169 - 198." type="journal article" year="1967">A.M. Clark 1967</bibRefCitation>
: 191;
<bibRefCitation id="EFAB2AEAFFA7FF8BD76688F6EC6F74FB" author="Marsh" pageId="16" pageNumber="17" refString="Marsh, L. M. &amp; Campbell, A. C. (1991) A new species of Ferdina (Echinodermata: Asteroidea) from the Sultanate of Oman with discussion of the relationships of the genus within the family Ophidiasteridae. Bulletin of the British Museum of Natural History, Zoology, 57 (2), 213 - 219." type="journal article" year="1991">Marsh &amp; Campbell 1991</bibRefCitation>
: 213.
</paragraph>
</subSubSection>
<subSubSection id="C3200490FFA7FF8BD2B08B57E9F47700" box="[151,1277,1543,1568]" pageId="16" pageNumber="17" type="description">
<paragraph id="8B85571BFFA7FF8BD2B08B57E9F47700" blockId="16.[151,1436,1543,2037]" box="[151,1277,1543,1568]" pageId="16" pageNumber="17">
<emphasis id="B94E8B09FFA7FF8BD2B08B57EC687700" bold="true" box="[151,353,1543,1568]" pageId="16" pageNumber="17">Included species.</emphasis>
<taxonomicName id="4C3A2C98FFA7FF8BD34F8B59EF7E7700" authority="Gray 1840" authorityName="Gray" authorityYear="1840" box="[360,631,1544,1568]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="flavescens">
<emphasis id="B94E8B09FFA7FF8BD34F8B59ECFE7700" box="[360,503,1544,1568]" italics="true" pageId="16" pageNumber="17">F. flavescens</emphasis>
<bibRefCitation id="EFAB2AEAFFA7FF8BD3DA8B58EF7E7700" author="Gray" box="[509,631,1544,1568]" pageId="16" pageNumber="17" refString="Gray, J. E. (1840) A synopsis of the genera and species of the class Hypostoma (Asterias Linnaeus). Annals of the Magazine of Natural History, 6, 175 - 184, 275 - 290." type="journal article" year="1840">Gray 1840</bibRefCitation>
</taxonomicName>
;
<emphasis id="B94E8B09FFA7FF8BD0A48B58EE417700" bold="true" box="[643,840,1543,1568]" pageId="16" pageNumber="17">
<taxonomicName id="4C3A2C98FFA7FF8BD0A48B58EFD5773F" box="[643,732,1544,1567]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="mena">
<emphasis id="B94E8B09FFA7FF8BD0A48B58EFD5773F" bold="true" box="[643,732,1544,1567]" italics="true" pageId="16" pageNumber="17">F. mena</emphasis>
</taxonomicName>
nov. sp.,
</emphasis>
<taxonomicName id="4C3A2C98FFA7FF8BD1698B59E9F17700" authority="Marsh &amp; Campbell 1991" authorityName="Marsh &amp; Campbell" authorityYear="1991" box="[846,1272,1543,1568]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="sadhensis">
<emphasis id="B94E8B09FFA7FF8BD1698B59EEDE7700" box="[846,983,1544,1568]" italics="true" pageId="16" pageNumber="17">F. sadhensis</emphasis>
<bibRefCitation id="EFAB2AEAFFA7FF8BD1F98B57E9F17700" author="Marsh" box="[990,1272,1543,1568]" pageId="16" pageNumber="17" refString="Marsh, L. M. &amp; Campbell, A. C. (1991) A new species of Ferdina (Echinodermata: Asteroidea) from the Sultanate of Oman with discussion of the relationships of the genus within the family Ophidiasteridae. Bulletin of the British Museum of Natural History, Zoology, 57 (2), 213 - 219." type="journal article" year="1991">Marsh &amp; Campbell 1991</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3200490FFA7FF8BD2E08B7CEF7C7764" box="[199,629,1580,1605]" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8B85571BFFA7FF8BD2E08B7CEF7C7764" blockId="16.[151,1436,1543,2037]" box="[199,629,1580,1605]" pageId="16" pageNumber="17">
<emphasis id="B94E8B09FFA7FF8BD2E08B7CEC6B7765" bold="true" box="[199,354,1580,1605]" pageId="16" pageNumber="17">Type species.</emphasis>
<taxonomicName id="4C3A2C98FFA7FF8BD34E8B7DEF7C7764" authority="Gray 1840" authorityName="Gray" authorityYear="1840" box="[361,629,1580,1604]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="flavescens">
<emphasis id="B94E8B09FFA7FF8BD34E8B7DECFE7764" box="[361,503,1580,1604]" italics="true" pageId="16" pageNumber="17">F. flavescens</emphasis>
<bibRefCitation id="EFAB2AEAFFA7FF8BD3D98B7CEF7C7764" author="Gray" box="[510,629,1580,1604]" pageId="16" pageNumber="17" refString="Gray, J. E. (1840) A synopsis of the genera and species of the class Hypostoma (Asterias Linnaeus). Annals of the Magazine of Natural History, 6, 175 - 184, 275 - 290." type="journal article" year="1840">Gray 1840</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3200490FFA7FF8AD2E08B1FE98D7218" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8B85571BFFA7FF8BD2E08B1FEE97763D" blockId="16.[151,1436,1543,2037]" pageId="16" pageNumber="17">
<emphasis id="B94E8B09FFA7FF8BD2E08B1FEC487748" bold="true" box="[199,321,1615,1640]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
Body stellate, elongate arms, small disk (R/r=3.95.6). Arms cylindrical with flattened actinal surface. Broad at base, but more tapering at tips Body surface covered by strongly fine to coarse granules extending from base of furrow spines. Bald surfaces present on first two paired superomarginals in each interradius
<taxonomicName id="4C3A2C98FFA7FF8BD7648BC9E8917790" box="[1347,1432,1689,1712]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="mena" status="sp. nov.">
<emphasis id="B94E8B09FFA7FF8BD7648BC9E8917790" box="[1347,1432,1689,1712]" italics="true" pageId="16" pageNumber="17">F. mena</emphasis>
</taxonomicName>
<emphasis id="B94E8B09FFA7FF8BD2B08BECEDDF77F5" bold="true" box="[151,214,1724,1749]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A27D3672FFA7FF8BD2B08BECEDDF77F5" box="[151,214,1724,1749]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, otherwise absent in other
<taxonomicName id="4C3A2C98FFA7FF8BD02D8BECEF6A77F4" box="[522,611,1724,1748]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA7FF8BD02D8BECEF6A77F4" box="[522,611,1724,1748]" italics="true" pageId="16" pageNumber="17">Ferdina</emphasis>
</taxonomicName>
spp. Large convex abactinal plates round to dome like in shape overlying smaller, irregularly round plates. These larger plates vary in size, distribution and abundance relative to the smaller plates. Single papular pores present between smaller abactinal plates.
</paragraph>
<paragraph id="8B85571BFFA7FF8BD2E08A77EEF276D4" blockId="16.[151,1436,1543,2037]" pageId="16" pageNumber="17">
Marginal plates variably regular or irregular in series, shape and size. Superomarginals variably strongly convex to more weakly so. Inferomarginals variably convex. Smaller secondary abactinal plates as well as papular pores present between larger marginal plates in some species. Superomarginals and inferomarginals in distinct, but recognizable series in
<taxonomicName id="4C3A2C98FFA7FF8BD3868AC5ECF2768C" box="[417,507,1941,1964]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="16" pageNumber="17" phylum="Echinodermata" rank="species" species="mena" status="sp. nov.">
<emphasis id="B94E8B09FFA7FF8BD3868AC5ECF2768C" box="[417,507,1941,1964]" italics="true" pageId="16" pageNumber="17">F. mena</emphasis>
</taxonomicName>
,
<emphasis id="B94E8B09FFA7FF8BD0358AC4EF5F768D" bold="true" box="[530,598,1940,1965]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A27D3672FFA7FF8BD0358AC4EF5F768D" box="[530,598,1940,1965]" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
whereas others show marginal plates in irregular, jumbled, sometimes completely discontinuous series. Inferomarginals tend to be in more recognizable series. Both superomarginal and inferomarginals tend to be in more regular and continuous distally along arm.
</paragraph>
<paragraph id="8B85571BFFA6FF8AD2E08DC7E860708D" blockId="17.[151,1437,151,824]" pageId="17" pageNumber="18">Actinal surface present in three to five distinct chevron like series extending only partially along the arm. Actinal plates vary in shape from more rounded to irregularly polygonal. Granulated tegument also variably fine to coarse grained depending on species. Papulae absent from actinal surface. Granulated tegument extends to lower base of furrow spines which occur in a single series along the tube foot groove. Furrow spines, thick with blunt tips, round to quadrate in cross-section, one or two per adambulacral plate. Adambulacral plates with no accessories other than the granulated tegument covering the body surface. Granulated tegument similarly covers the oral plate which has no other surficial accessories. One spine present on each oral plate (i.e., two directed into mouth) but as with the furrow spines, they are short, blunt with basal portion covered by granulated tegument.</paragraph>
<paragraph id="8B85571BFFA6FF8AD2E08CE7ECE17338" blockId="17.[151,1437,151,824]" pageId="17" pageNumber="18">
<materialsCitation id="3B525D46FFA6FF8AD2E08CE7ECE17338" ID-GBIF-Occurrence="1503739495" country="Oman" location="western Indian Ocean" municipality="Europa Island" pageId="17" pageNumber="18" specimenCount="1">
<emphasis id="B94E8B09FFA6FF8AD2E08CE7EF4570F0" bold="true" box="[199,588,439,464]" pageId="17" pageNumber="18">Taxonomic comments &amp; history.</emphasis>
<taxonomicName id="4C3A2C98FFA6FF8AD0748CE8EFA570F0" box="[595,684,440,464]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA6FF8AD0748CE8EFA570F0" box="[595,684,440,464]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
</taxonomicName>
is known primarily from the
<location id="8EE501C0FFA6FF8AD1DC8CE8E9F970F0" LSID="urn:lsid:plazi:treatment:0393E60DFFA7FF8AD2B0880FE98D7218:8EE501C0FFA6FF8AD1DC8CE8E9F970F0" box="[1019,1264,439,464]" country="Oman" municipality="Europa Island" name="western Indian Ocean" pageId="17" pageNumber="18">western Indian Ocean</location>
from intertidal to subtidal depths with known species occurring from
<collectingCountry id="F32D178BFFA6FF8AD0D98C8CEE4A70D4" box="[766,835,476,500]" name="Oman" pageId="17" pageNumber="18">Oman</collectingCountry>
to
<collectingMunicipality id="6BE1CD61FFA6FF8AD14F8C8DE90C70D5" box="[872,1029,476,501]" pageId="17" pageNumber="18">Europa Island</collectingMunicipality>
and
<collectingCountry id="F32D178BFFA6FF8AD61A8C8CE9CE70D5" box="[1085,1223,476,501]" name="Madagascar" pageId="17" pageNumber="18">Madagascar</collectingCountry>
. Nothing is known about its biology and ecology.
</materialsCitation>
</paragraph>
<paragraph id="8B85571BFFA6FF8AD2E08F74E98D7218" blockId="17.[151,1437,151,824]" pageId="17" pageNumber="18">
Historically,
<taxonomicName id="4C3A2C98FFA6FF8AD37D8F74ECBA731C" box="[346,435,548,572]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA6FF8AD37D8F74ECBA731C" box="[346,435,548,572]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EFAB2AEAFFA6FF8AD0A98F74EE12731C" author="Gray" box="[654,795,548,572]" pageId="17" pageNumber="18" refString="Gray, J. E. (1840) A synopsis of the genera and species of the class Hypostoma (Asterias Linnaeus). Annals of the Magazine of Natural History, 6, 175 - 184, 275 - 290." type="journal article" year="1840">Gray (1840)</bibRefCitation>
to accommodate
<taxonomicName id="4C3A2C98FFA6FF8AD1CC8F75E975731C" box="[1003,1148,548,572]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="flavescens">
<emphasis id="B94E8B09FFA6FF8AD1CC8F75E975731C" box="[1003,1148,548,572]" italics="true" pageId="17" pageNumber="18">F. flavescens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A2C98FFA6FF8AD69E8F75E838731C" box="[1209,1329,549,572]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="cumingi">
<emphasis id="B94E8B09FFA6FF8AD69E8F75E838731C" box="[1209,1329,549,572]" italics="true" pageId="17" pageNumber="18">F. cumingi</emphasis>
</taxonomicName>
.
<taxonomicName id="4C3A2C98FFA6FF8AD7648F74EDF8737F" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="cumingi">
<emphasis id="B94E8B09FFA6FF8AD7648F74EDF8737F" italics="true" pageId="17" pageNumber="18">Ferdina cumingi</emphasis>
</taxonomicName>
has since been reclassified into
<taxonomicName id="4C3A2C98FFA6FF8AD04A8F18EFE37340" box="[621,746,584,608]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA6FF8AD04A8F18EFE37340" box="[621,746,584,608]" italics="true" pageId="17" pageNumber="18">Neoferdina</emphasis>
</taxonomicName>
and many other
<taxonomicName id="4C3A2C98FFA6FF8AD1948F18E9057340" box="[947,1036,584,608]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA6FF8AD1948F18E9057340" box="[947,1036,584,608]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
</taxonomicName>
-like taxa have been recognized in subsequent work (see Marsh and Campbell 19991 for full review). In spite of this, relatively little work has been published on
<taxonomicName id="4C3A2C98FFA6FF8AD3138FC0EC877388" box="[308,398,656,680]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA6FF8AD3138FC0EC877388" box="[308,398,656,680]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
</taxonomicName>
itself.
<bibRefCitation id="EFAB2AEAFFA6FF8AD3C68FDFEE2E7388" author="Marsh" box="[481,807,655,680]" pageId="17" pageNumber="18" refString="Marsh, L. M. &amp; Campbell, A. C. (1991) A new species of Ferdina (Echinodermata: Asteroidea) from the Sultanate of Oman with discussion of the relationships of the genus within the family Ophidiasteridae. Bulletin of the British Museum of Natural History, Zoology, 57 (2), 213 - 219." type="journal article" year="1991">Marsh and Campbell (1991)</bibRefCitation>
provided a key to the “
<taxonomicName id="4C3A2C98FFA6FF8AD6658FC0E9927388" box="[1090,1179,656,680]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA6FF8AD6658FC0E9927388" box="[1090,1179,656,680]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
</taxonomicName>
group of genera” and distinguished
<taxonomicName id="4C3A2C98FFA6FF8AD3148FE4EC8473EC" box="[307,397,692,716]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA6FF8AD3148FE4EC8473EC" box="[307,397,692,716]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
</taxonomicName>
from other similar genera based on the presence of a bare marginal plates as well as size and morphology of abactinal and marginal plates. However, based on the description of the new
<taxonomicName id="4C3A2C98FFA6FF8AD68F8F88E80B73D0" box="[1192,1282,728,752]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA6FF8AD68F8F88E80B73D0" box="[1192,1282,728,752]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
</taxonomicName>
sp. described below, it is apparent that the presence of bare spots within the
<taxonomicName id="4C3A2C98FFA6FF8AD1568FACEEC37234" box="[881,970,764,788]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
<emphasis id="B94E8B09FFA6FF8AD1568FACEEC37234" box="[881,970,764,788]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
</taxonomicName>
“group” is more variable than had been previously documented. Its use as a key character in the key should be regarded carefully.
</paragraph>
</subSubSection>
</treatment>
</document>