143 lines
17 KiB
XML
143 lines
17 KiB
XML
<document id="C64BFE1716ED5D513466930F37108E4A" ID="10.11646/zootaxa.4271.1.1" ID-CLB-Dataset="32974" ID-DOI="10.11646/zootaxa.4271.1.1" ID-GBIF-Dataset="f3ad9b4c-35d8-4a01-a14b-3dd968623358" ID-ISSN="1175-5326" ID-Zenodo-Dep="583192" ID-ZooBank="50496AC4-D639-49A7-9249-386B037DAE72" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495697318696" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2017" docId="0393E60DFFA6FF8AD2B08EDCECA5770B" docLanguage="en" docName="zootaxa.4271.1.1SLASH11141.pdf" docOrigin="Zootaxa 4271 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ferdina Mah, 2017, n. sp." docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="FFAA9E75FFB7FF9BD2278D50ED097120" masterDocTitle="Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="18" updateTime="1698439914132" updateUser="ExternalLinkService">
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<mods:title id="C0D0F0161CA9465C5F4B00B9E820C790">Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species</mods:title>
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<mods:name id="DF291FC95D0E0ADE267360C57EF0224A" type="personal">
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<mods:namePart id="C962A541080481CF74D0B655AA54643C">Mah, Christopher L.</mods:namePart>
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<mods:title id="FBA952E04DFB0B4CF8FE5C8DB12EAE19">Zootaxa</mods:title>
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<mods:date id="F52BFFF12FB4923048EE57A1F820EA1D">2017</mods:date>
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<mods:number id="29F636226BC35EE47E0D946CF85C8A64">4271</mods:number>
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<mods:classification id="1D967AEFDDB12BD96DC024EA15CD9ABA">journal article</mods:classification>
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<mods:identifier id="9641D0BCF54B7021716142D5FDCC302A" type="CLB-Dataset">32974</mods:identifier>
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<mods:identifier id="72B81C9EAE6D8DCC0A05991B61BE0BB4" type="DOI">10.11646/zootaxa.4271.1.1</mods:identifier>
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<treatment id="0393E60DFFA6FF8AD2B08EDCECA5770B" ID-DOI="http://doi.org/10.5281/zenodo.6016987" ID-GBIF-Taxon="130844950" ID-Zenodo-Dep="6016987" LSID="urn:lsid:plazi:treatment:0393E60DFFA6FF8AD2B08EDCECA5770B" httpUri="http://treatment.plazi.org/id/0393E60DFFA6FF8AD2B08EDCECA5770B" lastPageNumber="18" pageId="17" pageNumber="18">
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<subSubSection id="C3200490FFA6FF8AD2B08EDCE91D742A" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="8B85571BFFA6FF8AD2B08EDCECFE7286" blockId="17.[151,503,908,934]" box="[151,503,908,934]" pageId="17" pageNumber="18">
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<heading id="D0CDE077FFA6FF8AD2B08EDCECFE7286" bold="true" box="[151,503,908,934]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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<emphasis id="B94E8B09FFA6FF8AD2B08EDCECFE7286" bold="true" box="[151,503,908,934]" pageId="17" pageNumber="18">
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<taxonomicName id="4C3A2C98FFA6FF8AD2B08EDCEDF27286" box="[151,251,908,934]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">Ferdina</taxonomicName>
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<emphasis id="B94E8B09FFA6FF8AD3268EDCECFE7286" bold="true" box="[257,503,908,934]" italics="true" pageId="17" pageNumber="18">
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in the
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<taxonomicName id="4C3A2C98FFA6FF8AD3698EDCECFE7286" box="[334,503,908,934]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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</emphasis>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B85571BFFA6FF8AD2B08E82E91D742A" blockId="17.[151,1437,978,1579]" pageId="17" pageNumber="18">
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Observations of
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<taxonomicName id="4C3A2C98FFA6FF8AD3718E83ECA672CB" box="[342,431,979,1003]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFA6FF8AD3718E83ECA672CB" box="[342,431,979,1003]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
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</taxonomicName>
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specimens identify several characters that support placement within the
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<taxonomicName id="4C3A2C98FFA6FF8AD6DD8E82E89072CB" box="[1274,1433,978,1003]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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. These include the presence of goniasterid-like abactinal plates arranged into a flattened surface framed by a distinct marginal plate arrangement in juveniles of
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<taxonomicName id="4C3A2C98FFA6FF8AD0B3894BEE3F7512" box="[660,822,1051,1075]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="mena" status="sp. nov.">
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<emphasis id="B94E8B09FFA6FF8AD0B3894BEE3F7512" box="[660,822,1051,1075]" italics="true" pageId="17" pageNumber="18">Ferdina mena</emphasis>
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</taxonomicName>
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<emphasis id="B94E8B09FFA6FF8AD165894AEE8C7513" bold="true" box="[834,901,1050,1075]" pageId="17" pageNumber="18">
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<taxonomicNameLabel id="A27D3672FFA6FF8AD165894AEE8C7513" box="[834,901,1050,1075]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="13014B9EFFA6FF8AD1BE894AEEEA7513" box="[921,995,1050,1075]" captionStart="FIGURE 6" captionStartId="21.[151,250,1395,1417]" captionTargetBox="[200,1384,199,1337]" captionTargetId="figure@21.[151,1436,163,1374]" captionTargetPageId="21" captionText="FIGURE 6. Ferdina mena n. sp. Small goniasterid-like specimens. MNHN-IE- 2007 - 4941. Scale Bar = 1.5 mm A. Abactinal surface. B. A bactinal surface detail. C. Lateral view showing paired bare regions in each interradius. D. Actinal surface." httpUri="https://zenodo.org/record/583198/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
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). As outlined in the introduction, the placement of
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<taxonomicName id="4C3A2C98FFA6FF8AD315896EEC827576" box="[306,395,1086,1110]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFA6FF8AD315896EEC827576" box="[306,395,1086,1110]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
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</taxonomicName>
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in the
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<taxonomicName id="4C3A2C98FFA6FF8AD3FB896EEF717577" box="[476,632,1086,1111]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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was first considered by
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<bibRefCitation id="EFAB2AEAFFA6FF8AD1A8896EE9267577" author="Viguier" box="[911,1071,1086,1111]" pageId="17" pageNumber="18" refString="Viguier, C. (1878) Classification des Stellerides. Comptes Rendus de l'Academie des Sciences de Paris, 84, 681 - 583. [also in Annals of the Magazine of Natural History, 2 (5), 103 - 105]" type="journal article" year="1878">Viguier (1878</bibRefCitation>
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,
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<bibRefCitation id="EFAB2AEAFFA6FF8AD61B896FE97C7577" author="Viguier" box="[1084,1141,1087,1111]" pageId="17" pageNumber="18" refString="Viguier, C. (1879) Anatomie comparee du squelette des Stellerides. Archives de Zoologie Experimentale et Generale, 7, 33 - 250." type="journal article" year="1879">1879</bibRefCitation>
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) with some agreement by
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<bibRefCitation id="EFAB2AEAFFA6FF8AD2B08932EC3F755B" author="Fisher" box="[151,310,1122,1147]" pageId="17" pageNumber="18" refString="Fisher, W. K. (1919) Starfishes of the Philippine seas and adjacent waters. Bulletin of the United States National Museum, 3 (100), 1 - 547, 156 pls." type="journal article" year="1919">Fisher (1919)</bibRefCitation>
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. The placement of
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<taxonomicName id="4C3A2C98FFA6FF8AD0388933EF95755B" box="[543,668,1123,1147]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFA6FF8AD0388933EF95755B" box="[543,668,1123,1147]" italics="true" pageId="17" pageNumber="18">Neoferdina</emphasis>
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</taxonomicName>
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within the
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<taxonomicName id="4C3A2C98FFA6FF8AD10E8932EECC755B" box="[809,965,1122,1147]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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by
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<bibRefCitation id="EFAB2AEAFFA6FF8AD1DF8932E9F3755B" author="Mah" box="[1016,1274,1122,1147]" pageId="17" pageNumber="18" refString="Mah, C. L. & Foltz, D. W. (2011) Molecular Phylogeny of the Valvatacea (Asteroidea, Echinodermata). Zoological Journal of the Linnean Society, 161, 769 - 788." type="journal article" year="2011">Mah and Foltz (2011)</bibRefCitation>
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supported by molecular data, suggests that a closely related genus with shared characters (e.g., furrow spination, granulate tegument covering adambulacrals) such as
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<taxonomicName id="4C3A2C98FFA6FF8AD05E89FBEFDB75E3" box="[633,722,1195,1219]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFA6FF8AD05E89FBEFDB75E3" box="[633,722,1195,1219]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
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</taxonomicName>
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can also be plausibly classified within the
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<taxonomicName id="4C3A2C98FFA6FF8AD69689FAE85975E3" box="[1201,1360,1194,1219]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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.
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<bibRefCitation id="EFAB2AEAFFA6FF8AD77C89FAEC5C75C7" author="Blake" pageId="17" pageNumber="18" refString="Blake, D. B. & Reid, R. (1998) Some Albian (Cretaceous) Asteroids (Echinodermata) from Texas and their paleobiological implications. Journal of Paleontology, 72 (3), 512 - 532." type="journal article" year="1998">Blake and Reid (1998)</bibRefCitation>
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also noted goniasterid affinities when comparing members of the “
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<taxonomicName id="4C3A2C98FFA6FF8AD649899EE9C175C6" box="[1134,1224,1230,1254]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFA6FF8AD649899EE9C175C6" box="[1134,1224,1230,1254]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
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</taxonomicName>
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group”, including
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<emphasis id="B94E8B09FFA6FF8AD2B089A3EF03742B" box="[151,522,1267,1291]" italics="true" pageId="17" pageNumber="18">
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<taxonomicName id="4C3A2C98FFA6FF8AD2B089A3EDFA742B" box="[151,243,1267,1291]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">Ferdina</taxonomicName>
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,
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<taxonomicName id="4C3A2C98FFA6FF8AD2D989A3EC8F742B" box="[254,390,1267,1291]" class="Asteroidea" family="Ophidiasteridae" genus="Paraferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">Paraferdina</taxonomicName>
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<taxonomicName id="4C3A2C98FFA6FF8AD3AA89A3EF03742B" box="[397,522,1267,1291]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">Neoferdina</taxonomicName>
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</emphasis>
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and
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<taxonomicName id="4C3A2C98FFA6FF8AD06789A3EFCB742B" box="[576,706,1267,1291]" class="Asteroidea" family="Ophidiasteridae" genus="Sinoferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFA6FF8AD06789A3EFCB742B" box="[576,706,1267,1291]" italics="true" pageId="17" pageNumber="18">Sinoferdina</emphasis>
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</taxonomicName>
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with the Cretaceous
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<taxonomicName id="4C3A2C98FFA6FF8AD18989A3E904742B" box="[942,1037,1267,1291]" class="Asteroidea" family="Ophidiasteridae" genus="Denebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFA6FF8AD18989A3E904742B" box="[942,1037,1267,1291]" italics="true" pageId="17" pageNumber="18">Denebia</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3200490FFA6FF8AD2E08846ECA5770B" pageId="17" pageNumber="18" type="description">
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<paragraph id="8B85571BFFA6FF8AD2E08846ECA5770B" blockId="17.[151,1437,978,1579]" pageId="17" pageNumber="18">
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Relatively small individuals of
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<taxonomicName id="4C3A2C98FFA6FF8AD0158846EFD8740E" box="[562,721,1302,1326]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="mena" status="sp. nov.">
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<emphasis id="B94E8B09FFA6FF8AD0158846EFD8740E" box="[562,721,1302,1326]" italics="true" pageId="17" pageNumber="18">Ferdina mena</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27D3672FFA6FF8AD0FC8847EE19740E" box="[731,784,1303,1326]" pageId="17" pageNumber="18" rank="species">n.sp.</taxonomicNameLabel>
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(e.g. MNHN-IE-2007-4941 at R=
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<quantity id="4CC2FAFEFFA6FF8AD6828847E9FF740F" box="[1189,1270,1303,1327]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="cm" value="1.2">1.2 cm</quantity>
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) show a welldeveloped marginal series with a more quadrate and blocky morphology forming a distinct frame around the abactinal plates (
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<figureCitation id="13014B9EFFA6FF8AD370880EECAE7457" box="[343,423,1374,1399]" captionStart="FIGURE 6" captionStartId="21.[151,250,1395,1417]" captionTargetBox="[200,1384,199,1337]" captionTargetId="figure@21.[151,1436,163,1374]" captionTargetPageId="21" captionText="FIGURE 6. Ferdina mena n. sp. Small goniasterid-like specimens. MNHN-IE- 2007 - 4941. Scale Bar = 1.5 mm A. Abactinal surface. B. A bactinal surface detail. C. Lateral view showing paired bare regions in each interradius. D. Actinal surface." httpUri="https://zenodo.org/record/583198/files/figure.png" pageId="17" pageNumber="18">Figs. 6</figureCitation>
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A–C). Although the marginal plate morphology and arrangement appear to vary more in larger individuals, these smaller specimens suggest synapomorphies for placement in the
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<taxonomicName id="4C3A2C98FFA6FF8AD68B88D2E84274BB" box="[1196,1355,1410,1435]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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. These specimens also show the arm as quadrate in cross-section as opposed to larger individuals with a more rounded or convex abactinal surface. Although juveniles of the other two
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<taxonomicName id="4C3A2C98FFA6FF8AD14B889BEECC74C3" box="[876,965,1483,1507]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FFA6FF8AD14B889BEECC74C3" box="[876,965,1483,1507]" italics="true" pageId="17" pageNumber="18">Ferdina</emphasis>
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</taxonomicName>
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spp. were unavailable, those of
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<taxonomicName id="4C3A2C98FFA6FF8AD766889BE89174C2" box="[1345,1432,1483,1506]" class="Asteroidea" family="Ophidiasteridae" genus="Ferdina" kingdom="Animalia" order="Valvatida" pageId="17" pageNumber="18" phylum="Echinodermata" rank="species" species="mena">
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<emphasis id="B94E8B09FFA6FF8AD766889BE89174C2" box="[1345,1432,1483,1506]" italics="true" pageId="17" pageNumber="18">F. mena</emphasis>
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</taxonomicName>
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would suggest that there is a transformation in the shape/convexity of the abactinal surface and arm shape as this species increases in size.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |