295 lines
37 KiB
XML
295 lines
37 KiB
XML
<document id="E2964F8917056D4F60BAC63439F1061F" ID="10.11646/zootaxa.4271.1.1" ID-CLB-Dataset="32974" ID-DOI="10.11646/zootaxa.4271.1.1" ID-GBIF-Dataset="f3ad9b4c-35d8-4a01-a14b-3dd968623358" ID-ISSN="1175-5326" ID-Zenodo-Dep="583192" ID-ZooBank="50496AC4-D639-49A7-9249-386B037DAE72" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495697318696" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2017" docId="0393E60DFF9EFFB0D2B08DC7EC52727C" docLanguage="en" docName="zootaxa.4271.1.1SLASH11141.pdf" docOrigin="Zootaxa 4271 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Neoferdina annae Mah, 2017, n. sp." docType="treatment" docVersion="9" lastPageNumber="44" masterDocId="FFAA9E75FFB7FF9BD2278D50ED097120" masterDocTitle="Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species" masterLastPageNumber="72" masterPageNumber="1" pageNumber="42" updateTime="1698439914132" updateUser="ExternalLinkService">
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<mods:mods id="52E977AA598F915656E715FA90F4AA2B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0BA74ACD26AE4D74EC394335101AFF09">
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<mods:title id="87F3749E4DF0BA2527BD8493A07265DC">Overview of the Ferdina - like Goniasteridae (Echinodermata: Asteroidea) including a new subfamily, three new genera and fourteen new species</mods:title>
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<mods:name id="6F0E86F41A322C5AEB5161926C400F55" type="personal">
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<mods:roleTerm id="E13A9E7268E91EFDAC0E46585870E144">Author</mods:roleTerm>
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<mods:namePart id="35F29C0F2938591B0893B2B7EFB79FA5">Mah, Christopher L.</mods:namePart>
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<mods:typeOfResource id="C4FE6FE058A051A127EE0145A416A45C">text</mods:typeOfResource>
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<mods:title id="81D3AC9F08A33E669952C149989504F1">Zootaxa</mods:title>
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<mods:part id="B82E64BB027A338925599667FF899DE9">
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<mods:date id="EBE75AAB9C7FC0A85E0C04E80780F7F2">2017</mods:date>
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<mods:number id="5EE4D78D807BAE3A2D70F22ACD5EF850">4271</mods:number>
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<mods:detail id="150E1663813B86511305AA1E409C2BB9" type="issue">
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<mods:number id="CDA3C21724F5CE89EBBB94D9231729D2">1</mods:number>
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<mods:classification id="395A4A29958C487912EAE28C23F707F3">journal article</mods:classification>
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<mods:identifier id="EFBC59EC0A6DEF4600B24F0C6D8F9ED4" type="CLB-Dataset">32974</mods:identifier>
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<mods:identifier id="CCD3F4281300C1A18D753D0C8E10A174" type="DOI">10.11646/zootaxa.4271.1.1</mods:identifier>
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<mods:identifier id="07EE86CB4AC56201579D5E297AEA8082" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4867F42A79F79E06B2479760C19E15FD" type="Zenodo-Dep">583192</mods:identifier>
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<treatment id="0393E60DFF9EFFB0D2B08DC7EC52727C" ID-DOI="http://doi.org/10.5281/zenodo.6017011" ID-GBIF-Taxon="130844974" ID-Zenodo-Dep="6017011" LSID="urn:lsid:plazi:treatment:0393E60DFF9EFFB0D2B08DC7EC52727C" httpUri="http://treatment.plazi.org/id/0393E60DFF9EFFB0D2B08DC7EC52727C" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
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<subSubSection id="C3200490FF9EFFB2D2B08DC7EC3271F3" pageId="41" pageNumber="42" type="nomenclature">
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<paragraph id="8B85571BFF9EFFB2D2B08DC7ECB27192" blockId="41.[151,443,151,211]" box="[151,443,151,178]" pageId="41" pageNumber="42">
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<heading id="D0CDE077FF9EFFB2D2B08DC7ECB27192" bold="true" box="[151,443,151,178]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
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<emphasis id="B94E8B09FF9EFFB2D2B08DC7ECB27192" bold="true" box="[151,443,151,178]" pageId="41" pageNumber="42">
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<taxonomicName id="4C3A2C98FF9EFFB2D2B08DC7EC7B7191" ID-CoL="46H82" box="[151,370,151,177]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="annae" status="sp. nov.">
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<emphasis id="B94E8B09FF9EFFB2D2B08DC7EC7B7191" bold="true" box="[151,370,151,177]" italics="true" pageId="41" pageNumber="42">Neoferdina annae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27D3672FF9EFFB2D35E8DC8ECB27192" box="[377,443,152,178]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B85571BFF9EFFB2D2B08DEAEC3271F3" blockId="41.[151,443,151,211]" box="[151,315,186,211]" pageId="41" pageNumber="42">
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<figureCitation id="13014B9EFF9EFFB2D2B08DEAEC0A71F3" box="[151,259,186,211]" captionStart="FIGURE 13" captionStartId="42.[151,250,1846,1868]" captionTargetBox="[165,1408,225,1788]" captionTargetId="figure@42.[151,1436,193,1824]" captionTargetPageId="42" captionText="FIGURE 13. Neoferdina annae n. sp. Holotype, CASIZ 218854. Scale Bar = 3.5 mm. A. Abactinal surface. B. Abactinal disk plate detail. C. Abactinal-lateral view showing superomarginal / abactinal contact. D. Detail of arm tip showing superomarginal plates. E. Actinal surface. F. Arm tip showing paired bald patches on penultimate abutted superomarginal plates." httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="41" pageNumber="42">Figure 13</figureCitation>
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A–F
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</paragraph>
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</subSubSection>
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<subSubSection id="C3200490FF9EFFB2D2B08C52EEFA703B" box="[151,1011,258,283]" pageId="41" pageNumber="42" type="etymology">
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<paragraph id="8B85571BFF9EFFB2D2B08C52EEFA703B" blockId="41.[151,1437,258,1939]" box="[151,1011,258,283]" pageId="41" pageNumber="42">
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<emphasis id="B94E8B09FF9EFFB2D2B08C52EC17703B" bold="true" box="[151,286,258,283]" pageId="41" pageNumber="42">Etymology.</emphasis>
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This species is named for my colleague and friend Anna Sutton.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3200490FF9EFFB2D2E08C76ECAF731B" pageId="41" pageNumber="42" type="diagnosis">
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<paragraph id="8B85571BFF9EFFB2D2E08C76EE03708B" blockId="41.[151,1437,258,1939]" pageId="41" pageNumber="42">
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<emphasis id="B94E8B09FF9EFFB2D2E08C76EC48701F" bold="true" box="[199,321,294,319]" pageId="41" pageNumber="42">Diagnosis.</emphasis>
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This species is distinctive for having a nearly continuous granular tegument which covers all plate surfaces save for the paired bald circles on the paired penultimate superomarginals. Abactinal plates are flat and mostly homogeneous in size and shape with no large, convex plates as seen in most other
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<taxonomicName id="4C3A2C98FF9EFFB2D69B8C3EE83070A6" box="[1212,1337,366,390]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FF9EFFB2D69B8C3EE83070A6" box="[1212,1337,366,390]" italics="true" pageId="41" pageNumber="42">Neoferdina</emphasis>
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</taxonomicName>
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species. Distalmost superomarginal plates are abutted at arm tip.
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</paragraph>
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<paragraph id="8B85571BFF9EFFB2D2E08CE6ECAF731B" blockId="41.[151,1437,258,1939]" pageId="41" pageNumber="42">
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<emphasis id="B94E8B09FF9EFFB2D2E08CE6ECDD70EF" bold="true" box="[199,468,438,463]" pageId="41" pageNumber="42">Taxonomic comments.</emphasis>
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This species is similar to
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<taxonomicName id="4C3A2C98FF9EFFB2D1208CE6EEE870EE" box="[775,993,438,462]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="insolita">
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<emphasis id="B94E8B09FF9EFFB2D1208CE6EEE870EE" box="[775,993,438,462]" italics="true" pageId="41" pageNumber="42">Neoferdina insolita</emphasis>
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</taxonomicName>
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, in that it shares the absence of large, convex bald plates on the abactinal surface. It differs from nearly all other
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<taxonomicName id="4C3A2C98FF9EFFB2D1CF8C8BE96C70D3" box="[1000,1125,475,499]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FF9EFFB2D1CF8C8BE96C70D3" box="[1000,1125,475,499]" italics="true" pageId="41" pageNumber="42">Neoferdina</emphasis>
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</taxonomicName>
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species in that there are no bald patches on the surfaces of the superomarginal or inferomarginal plates, save for the two superomarginals adjacent to the terminal.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3200490FF9EFFB0D2E08F16E8517388" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42" type="description">
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<paragraph id="8B85571BFF9EFFB2D2E08F16ECDA7283" blockId="41.[151,1437,258,1939]" pageId="41" pageNumber="42">
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<taxonomicName id="4C3A2C98FF9EFFB2D2E08F16EC9C737E" box="[199,405,582,606]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="annae" status="sp. nov.">
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<emphasis id="B94E8B09FF9EFFB2D2E08F16EC9C737E" box="[199,405,582,606]" italics="true" pageId="41" pageNumber="42">Neoferdina annae</emphasis>
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</taxonomicName>
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<emphasis id="B94E8B09FF9EFFB2D3868F16ECEC737F" bold="true" box="[417,485,582,607]" pageId="41" pageNumber="42">
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<taxonomicNameLabel id="A27D3672FF9EFFB2D3868F16ECEC737F" box="[417,485,582,607]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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invites comparison with
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<taxonomicName id="4C3A2C98FF9EFFB2D1308F16E94B737E" box="[791,1090,582,606]" class="Asteroidea" family="Ophidiasteridae" genus="Paraferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="laccadivensis">
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<emphasis id="B94E8B09FF9EFFB2D1308F16E94B737E" box="[791,1090,582,606]" italics="true" pageId="41" pageNumber="42">Paraferdina laccadivensis</emphasis>
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</taxonomicName>
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with which it shares several characters, including the flat, round shaped, and numerous abactinal plates and especially the granular tegument covering the surface, including all of the marginal plates, nearly all of which are lacking bare patches.
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<taxonomicName id="4C3A2C98FF9EFFB2D7388FDEEDD573EA" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="annae" status="sp. nov.">
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<emphasis id="B94E8B09FF9EFFB2D7388FDEEDD573EA" italics="true" pageId="41" pageNumber="42">Neoferdina annae</emphasis>
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</taxonomicName>
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<emphasis id="B94E8B09FF9EFFB2D2C38FE2EC2C73EB" bold="true" box="[228,293,690,715]" pageId="41" pageNumber="42">
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<taxonomicNameLabel id="A27D3672FF9EFFB2D2C38FE2EC2C73EB" box="[228,293,690,715]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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differs in that it possesses a single circular bare patch on each of two penultimate superomarginals, and elongate superomarginals in contrast to
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<taxonomicName id="4C3A2C98FF9EFFB2D08B8F87EE6C73CE" box="[684,869,726,750]" class="Asteroidea" family="Ophidiasteridae" genus="Paraferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="laccadivensis">
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<emphasis id="B94E8B09FF9EFFB2D08B8F87EE6C73CE" box="[684,869,726,750]" italics="true" pageId="41" pageNumber="42">P. laccadivensis</emphasis>
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</taxonomicName>
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which displays no known bare patches on the superomarginals and which has more quadrate to elongate superomarginals.
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<taxonomicName id="4C3A2C98FF9EFFB2D1D48FABE9507232" box="[1011,1113,763,786]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="annae">
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<emphasis id="B94E8B09FF9EFFB2D1D48FABE9507232" box="[1011,1113,763,786]" italics="true" pageId="41" pageNumber="42">N. annae</emphasis>
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</taxonomicName>
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also displays a more stellate body shape with more triangular shaped arms that are more similar to those in
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<taxonomicName id="4C3A2C98FF9EFFB2D60A8E4EE9A37216" box="[1069,1194,798,822]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FF9EFFB2D60A8E4EE9A37216" box="[1069,1194,798,822]" italics="true" pageId="41" pageNumber="42">Neoferdina</emphasis>
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</taxonomicName>
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than in
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<taxonomicName id="4C3A2C98FF9EFFB2D72A8E4EE89C7216" box="[1293,1429,798,822]" class="Asteroidea" family="Ophidiasteridae" genus="Paraferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="genus">
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<emphasis id="B94E8B09FF9EFFB2D72A8E4EE89C7216" box="[1293,1429,798,822]" italics="true" pageId="41" pageNumber="42">Paraferdina</emphasis>
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</taxonomicName>
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. Given these character differences between
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<taxonomicName id="4C3A2C98FF9EFFB2D0AC8E13EFFD727A" box="[651,756,835,858]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="annae">
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<emphasis id="B94E8B09FF9EFFB2D0AC8E13EFFD727A" box="[651,756,835,858]" italics="true" pageId="41" pageNumber="42">N. annae</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C3A2C98FF9EFFB2D1158E13EEEC727B" box="[818,997,835,859]" class="Asteroidea" family="Ophidiasteridae" genus="Paraferdina" kingdom="Animalia" order="Valvatida" pageId="41" pageNumber="42" phylum="Echinodermata" rank="species" species="laccadivensis">
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<emphasis id="B94E8B09FF9EFFB2D1158E13EEEC727B" box="[818,997,835,859]" italics="true" pageId="41" pageNumber="42">P laccadivensis</emphasis>
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</taxonomicName>
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, this species creates ambiguity at the taxonomic boundary between these two genera. Further phylogenetic testing is needed to properly establish the boundaries for these genera.
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</paragraph>
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<paragraph id="8B85571BFF9EFFB2D2E08EFEEEB872E6" blockId="41.[151,1437,258,1939]" box="[199,945,942,967]" pageId="41" pageNumber="42">
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<emphasis id="B94E8B09FF9EFFB2D2E08EFEEC5072E7" bold="true" box="[199,345,942,967]" pageId="41" pageNumber="42">Occurrence.</emphasis>
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Balut Island, Philippines. approximately
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<quantity id="4CC2FAFEFF9EFFB2D10C8EFEEEA472E6" box="[811,941,942,967]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="41" pageNumber="42" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">100–150 m</quantity>
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.
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</paragraph>
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<paragraph id="8B85571BFF9EFFB2D2E08E82ECA5752F" blockId="41.[151,1437,258,1939]" pageId="41" pageNumber="42">
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<emphasis id="B94E8B09FF9EFFB2D2E08E82EC5072CB" bold="true" box="[199,345,978,1003]" pageId="41" pageNumber="42">Description.</emphasis>
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Body stellate (R/r=3.36–4.25), arms triangular in shape with weakly curved interradial arcs (
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<figureCitation id="13014B9EFF9EFFB2D7448E82EDBA752F" captionStart="FIGURE 13" captionStartId="42.[151,250,1846,1868]" captionTargetBox="[165,1408,225,1788]" captionTargetId="figure@42.[151,1436,193,1824]" captionTargetPageId="42" captionText="FIGURE 13. Neoferdina annae n. sp. Holotype, CASIZ 218854. Scale Bar = 3.5 mm. A. Abactinal surface. B. Abactinal disk plate detail. C. Abactinal-lateral view showing superomarginal / abactinal contact. D. Detail of arm tip showing superomarginal plates. E. Actinal surface. F. Arm tip showing paired bald patches on penultimate abutted superomarginal plates." httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="41" pageNumber="42">Figs. 13</figureCitation>
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A, B). Body flattened.
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</paragraph>
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<paragraph id="8B85571BFF9EFFB2D2E0894AEEEF74BB" blockId="41.[151,1437,258,1939]" pageId="41" pageNumber="42">
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Abactinal surface flattened, plates oval to circular to irregularly polygonal, (
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<figureCitation id="13014B9EFF9EFFB2D63B894AE97F7513" box="[1052,1142,1050,1075]" captionStart="FIGURE 13" captionStartId="42.[151,250,1846,1868]" captionTargetBox="[165,1408,225,1788]" captionTargetId="figure@42.[151,1436,193,1824]" captionTargetPageId="42" captionText="FIGURE 13. Neoferdina annae n. sp. Holotype, CASIZ 218854. Scale Bar = 3.5 mm. A. Abactinal surface. B. Abactinal disk plate detail. C. Abactinal-lateral view showing superomarginal / abactinal contact. D. Detail of arm tip showing superomarginal plates. E. Actinal surface. F. Arm tip showing paired bald patches on penultimate abutted superomarginal plates." httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="41" pageNumber="42">Figs. 13</figureCitation>
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A, B). Plate surface flat to weakly convex, mostly homogeneous in size but with smaller more homogeneously sized plates adjacent to contact with superomarginal plates. Abactinal plates covered with continuous granular tegument extending onto marginal and actinal plate surfaces. Individual granules fine, 10–30, usually 20 along a 1.0 mm line. If granular tegument is removed, plate surfaces are smooth and reddish in color with no convexities or other structures. glassy nodules absent. Granules obscuring plate boundaries on abactinal surface, but darker in color between plates. Abactinal plates five or six along arm base, tapering out to a single series where superomarginals abut over midline. Carinal series with irregular plates variably smaller and oval shaped to three times as large and round in shape, alternating in series (
|
||
<figureCitation id="13014B9EFF9EFFB2D32A886AEC6D7473" box="[269,356,1338,1363]" captionStart="FIGURE 13" captionStartId="42.[151,250,1846,1868]" captionTargetBox="[165,1408,225,1788]" captionTargetId="figure@42.[151,1436,193,1824]" captionTargetPageId="42" captionText="FIGURE 13. Neoferdina annae n. sp. Holotype, CASIZ 218854. Scale Bar = 3.5 mm. A. Abactinal surface. B. Abactinal disk plate detail. C. Abactinal-lateral view showing superomarginal / abactinal contact. D. Detail of arm tip showing superomarginal plates. E. Actinal surface. F. Arm tip showing paired bald patches on penultimate abutted superomarginal plates." httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="41" pageNumber="42">Fig. 13</figureCitation>
|
||
B). Papulae distinct, ranging from four to eight surrounding each abactinal plate, primarily occurring along arm and basal arm regions on disk. Absent from interradial regions and central disk regions. Madreporite polygonal to round, sulci well developed. Pedicellariae absent.
|
||
</paragraph>
|
||
<paragraph id="8B85571BFF9EFFB2D2E088F6EC9376B3" blockId="41.[151,1437,258,1939]" pageId="41" pageNumber="42">
|
||
Superomarginal plates 21–32 per interradius (arm tip to arm tip), inferomarginals 22–26 per interradius in the
|
||
<typeStatus id="5481E9B9FF9EFFB2D2B0889AEDF074C3" box="[151,249,1482,1507]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13014B9EFF9EFFB2D32C889AEC5674C3" box="[267,351,1482,1507]" captionStart="FIGURE 13" captionStartId="42.[151,250,1846,1868]" captionTargetBox="[165,1408,225,1788]" captionTargetId="figure@42.[151,1436,193,1824]" captionTargetPageId="42" captionText="FIGURE 13. Neoferdina annae n. sp. Holotype, CASIZ 218854. Scale Bar = 3.5 mm. A. Abactinal surface. B. Abactinal disk plate detail. C. Abactinal-lateral view showing superomarginal / abactinal contact. D. Detail of arm tip showing superomarginal plates. E. Actinal surface. F. Arm tip showing paired bald patches on penultimate abutted superomarginal plates." httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="41" pageNumber="42">Fig. 13</figureCitation>
|
||
A),
|
||
<quantity id="4CC2FAFEFF9EFFB2D3A9889AECFD74C3" box="[398,500,1482,1507]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" metricValueMax="9.144" metricValueMin="8.636" pageId="41" pageNumber="42" unit="in" value="35.0" valueMax="36.0" valueMin="34.0">34–36 in</quantity>
|
||
the
|
||
<typeStatus id="5481E9B9FF9EFFB2D00D889BEF8774C3" box="[554,654,1483,1507]" pageId="41" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
. Proximal marginal plates with 1:1 correspondence becoming more offset distally along the arms. Proximalmost superomarginals wide and quadrate in shape with subsequent plates becoming more elongate along arm becoming more quadrate adjacent to terminal plate. Superomarginal surface weakly convex to flat (
|
||
<figureCitation id="13014B9EFF9EFFB2D38C8B66EF0F776F" box="[427,518,1590,1615]" captionStart="FIGURE 13" captionStartId="42.[151,250,1846,1868]" captionTargetBox="[165,1408,225,1788]" captionTargetId="figure@42.[151,1436,193,1824]" captionTargetPageId="42" captionText="FIGURE 13. Neoferdina annae n. sp. Holotype, CASIZ 218854. Scale Bar = 3.5 mm. A. Abactinal surface. B. Abactinal disk plate detail. C. Abactinal-lateral view showing superomarginal / abactinal contact. D. Detail of arm tip showing superomarginal plates. E. Actinal surface. F. Arm tip showing paired bald patches on penultimate abutted superomarginal plates." httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="41" pageNumber="42">Figs 13</figureCitation>
|
||
B,C). Some segments along arm smaller, irregular in shape (presumably from damage). Superomarginal contact with abactinal plates round and convex. Distalmost two to three superomarginals abutted over midline (
|
||
<figureCitation id="13014B9EFF9EFFB2D3B08B2EECE277B7" box="[407,491,1662,1687]" captionStart="FIGURE 13" captionStartId="42.[151,250,1846,1868]" captionTargetBox="[165,1408,225,1788]" captionTargetId="figure@42.[151,1436,193,1824]" captionTargetPageId="42" captionText="FIGURE 13. Neoferdina annae n. sp. Holotype, CASIZ 218854. Scale Bar = 3.5 mm. A. Abactinal surface. B. Abactinal disk plate detail. C. Abactinal-lateral view showing superomarginal / abactinal contact. D. Detail of arm tip showing superomarginal plates. E. Actinal surface. F. Arm tip showing paired bald patches on penultimate abutted superomarginal plates." httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="41" pageNumber="42">Fig. 13</figureCitation>
|
||
D). Granular tegument on marginal plates continuous from abactinal plate surface save for one distinct round bald, dark colored patch on the surface of distalmost superomarginals, on the
|
||
<typeStatus id="5481E9B9FF9EFFB2D71D8BF2E895779B" box="[1338,1436,1698,1723]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
patches are restricted to paired superomarginals adjacent to the terminal plate (
|
||
<figureCitation id="13014B9EFF9EFFB2D6288B96E96277FF" box="[1039,1131,1734,1759]" captionStart="FIGURE 13" captionStartId="42.[151,250,1846,1868]" captionTargetBox="[165,1408,225,1788]" captionTargetId="figure@42.[151,1436,193,1824]" captionTargetPageId="42" captionText="FIGURE 13. Neoferdina annae n. sp. Holotype, CASIZ 218854. Scale Bar = 3.5 mm. A. Abactinal surface. B. Abactinal disk plate detail. C. Abactinal-lateral view showing superomarginal / abactinal contact. D. Detail of arm tip showing superomarginal plates. E. Actinal surface. F. Arm tip showing paired bald patches on penultimate abutted superomarginal plates." httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="41" pageNumber="42">Figs. 13</figureCitation>
|
||
D, F). Thus, each arm with paired dark spots on each arm tip. Inferomarginal plates broadly similar in overall size and shape distribution. Distalmost inferomarginals covered with coarser granules, several bearing thorny to spine-like tips. Single short granular spinelet present on two to three inferomarginals adjacent to the terminal. Inferomarginal adjacent to terminal plate also strongly convex. Terminal plates triangular in shape, with no granular tegument.
|
||
<collectingRegion id="49FE99F9FF9EFFB2D75D8A07E895764E" box="[1402,1436,1879,1902]" country="Maldives" name="Uthuru" pageId="41" pageNumber="42">No</collectingRegion>
|
||
pedicellariae observed.
|
||
</paragraph>
|
||
<caption id="DF450793FF9DFFB1D2B08A66E80576A9" httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="42" pageNumber="43" targetBox="[165,1408,225,1788]" targetPageId="42">
|
||
<paragraph id="8B85571BFF9DFFB1D2B08A66E80576A9" blockId="42.[151,1436,1845,1929]" pageId="42" pageNumber="43">
|
||
<emphasis id="B94E8B09FF9DFFB1D2B08A66EF9A766B" bold="true" box="[151,659,1845,1868]" pageId="42" pageNumber="43">
|
||
FIGURE 13.
|
||
<taxonomicName id="4C3A2C98FF9DFFB1D3018A65ECED766B" box="[294,484,1845,1867]" class="Asteroidea" family="Goniasteridae" genus="Neoferdina" kingdom="Animalia" order="Valvatida" pageId="42" pageNumber="43" phylum="Echinodermata" rank="species" species="annae" status="sp. nov.">
|
||
<emphasis id="B94E8B09FF9DFFB1D3018A65ECED766B" bold="true" box="[294,484,1845,1867]" italics="true" pageId="42" pageNumber="43">Neoferdina annae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27D3672FF9DFFB1D3CD8A66EF2A766B" box="[490,547,1846,1867]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
|
||
Holotype,
|
||
</emphasis>
|
||
CASIZ 218854. Scale Bar=3.5 mm.
|
||
<emphasis id="B94E8B09FF9DFFB1D62B8A66E92C766B" bold="true" box="[1036,1061,1846,1867]" pageId="42" pageNumber="43">A.</emphasis>
|
||
Abactinal surface.
|
||
<emphasis id="B94E8B09FF9DFFB1D6CC8A66E80B766B" bold="true" box="[1259,1282,1846,1867]" pageId="42" pageNumber="43">B.</emphasis>
|
||
Abactinal disk plate detail.
|
||
<emphasis id="B94E8B09FF9DFFB1D3338A05EC25764B" bold="true" box="[276,300,1877,1899]" pageId="42" pageNumber="43">C.</emphasis>
|
||
Abactinal-lateral view showing superomarginal/abactinal contact.
|
||
<emphasis id="B94E8B09FF9DFFB1D1F38A05EEE4764A" bold="true" box="[980,1005,1877,1898]" pageId="42" pageNumber="43">D.</emphasis>
|
||
Detail of arm tip showing superomarginal plates.
|
||
<emphasis id="B94E8B09FF9DFFB1D2F98A23EDFF76A8" bold="true" box="[222,246,1907,1928]" pageId="42" pageNumber="43">E.</emphasis>
|
||
Actinal surface.
|
||
<emphasis id="B94E8B09FF9DFFB1D3848A23ECBF76A8" bold="true" box="[419,438,1907,1928]" pageId="42" pageNumber="43">F.</emphasis>
|
||
Arm tip showing paired bald patches on penultimate abutted superomarginal plates.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B85571BFF9CFFB0D2E08DC7E80D70D5" blockId="43.[151,1437,151,861]" pageId="43" pageNumber="44">
|
||
Actinal surface composed of five to six actinal intermediate series, four of which terminate along the arm at various points away from the disk (
|
||
<figureCitation id="13014B9EFF9CFFB0D0018DECEF7E71F4" box="[550,631,188,213]" captionStart="FIGURE 13" captionStartId="42.[151,250,1846,1868]" captionTargetBox="[165,1408,225,1788]" captionTargetId="figure@42.[151,1436,193,1824]" captionTargetPageId="42" captionText="FIGURE 13. Neoferdina annae n. sp. Holotype, CASIZ 218854. Scale Bar = 3.5 mm. A. Abactinal surface. B. Abactinal disk plate detail. C. Abactinal-lateral view showing superomarginal / abactinal contact. D. Detail of arm tip showing superomarginal plates. E. Actinal surface. F. Arm tip showing paired bald patches on penultimate abutted superomarginal plates." httpUri="https://zenodo.org/record/583205/files/figure.png" pageId="43" pageNumber="44">Fig. 13</figureCitation>
|
||
E). Two incomplete actinal intermediate plate series on disk. Individual actinal plates quadrate to polygonal in shape. Actinal surface covered by same continuous granular integument covering abactinal and marginal surface.
|
||
<collectingRegion id="49FE99F9FF9CFFB0D05A8C55EF96703C" box="[637,671,261,284]" country="Maldives" name="Uthuru" pageId="43" pageNumber="44">No</collectingRegion>
|
||
spines or other accessories present on actinal surface other than granular integument. Furrow spines, round and quadrate in cross-section, present in a single series along adambulacral furrow. Two to three (mostly three) short blunt spines per adambulacral plate. Granular integument extends up to base and half-way along spines. Oral plates with a large oral spine on each paired plate, each projecting into mouth. Thus, two oral spines project into the mouth per interradius. Each of these, approximately two to three times as thick as the furrow spines.
|
||
<collectingRegion id="49FE99F9FF9CFFB0D0FF8CE8EFF370EF" box="[728,762,440,463]" country="Maldives" name="Uthuru" pageId="43" pageNumber="44">No</collectingRegion>
|
||
other accessories present on furrow spines other than the granular integument. Oral plates strongly convex or raised above plane of actinal intermediate plates.
|
||
</paragraph>
|
||
<paragraph id="8B85571BFF9CFFB0D2E08CAFE8517388" blockId="43.[151,1437,151,861]" pageId="43" pageNumber="44">
|
||
Although color in life was unavailable, patterns remained evident on the
|
||
<typeStatus id="5481E9B9FF9CFFB0D62C8CAFE9787338" box="[1035,1137,511,536]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
. Plates on the central disk were dark brown/orange with pink in the center. The dark brown coloration from the disk was present interradially on the disk between the primary circlet to the interradial superomarginals. Abactinal coloration was dark pink to brown. Actinal intermediate region and color from lateral to actinolateral regions was lighter but similar to that of the abactinal surface. The areas around the mouth and along the adambulacral furrows were yellow to white.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3200490FF9CFFB0D2E08FE4EC52727C" pageId="43" pageNumber="44" type="materials_examined">
|
||
<paragraph id="8B85571BFF9CFFB0D2E08FE4EC52727C" blockId="43.[151,1437,151,861]" pageId="43" pageNumber="44">
|
||
<materialsCitation id="3B525D46FF9CFFB0D2E08FE4E9FC73ED" ID-GBIF-Occurrence="1503739453" box="[199,1269,692,717]" collectionCode="CASIZ" country="Philippines" location="Balut Island" pageId="43" pageNumber="44" specimenCode="CASIZ 218854" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B94E8B09FF9CFFB0D2E08FE4EF3873ED" bold="true" box="[199,561,692,717]" pageId="43" pageNumber="44">
|
||
Material examined.
|
||
<typeStatus id="5481E9B9FF9CFFB0D39C8FE4EF2573ED" box="[443,556,692,717]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="DB9CFF60FF9CFFB0D01D8FE4EFE573ED" box="[570,748,692,717]" collectionCode="CASIZ" pageId="43" pageNumber="44">
|
||
<collectionCode id="ED2BCFDEFF9CFFB0D01D8FE4EF8273EC" box="[570,651,692,716]" pageId="43" pageNumber="44">CASIZ</collectionCode>
|
||
218854
|
||
</specimenCode>
|
||
;
|
||
<location id="8EE501C0FF9CFFB0D0DE8FE4EE8F73ED" LSID="urn:lsid:plazi:treatment:0393E60DFF9EFFB0D2B08DC7EC52727C:8EE501C0FF9CFFB0D0DE8FE4EE8F73ED" box="[761,902,692,717]" country="Philippines" name="Balut Island" pageId="43" pageNumber="44">Balut Island</location>
|
||
,
|
||
<collectingCountry id="F32D178BFF9CFFB0D1B48FE4E91D73ED" box="[915,1044,692,717]" name="Philippines" pageId="43" pageNumber="44">Philippines</collectingCountry>
|
||
, approximately
|
||
<quantity id="4CC2FAFEFF9CFFB0D6EB8FE4E9FC73ED" box="[1228,1269,692,717]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="43" pageNumber="44" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">100</quantity>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="3B525D46FF9CFFB0D7228FE4ECC37234" ID-GBIF-Occurrence="1503739427" collectionCode="CASIZ" country="Philippines" elevation="125" location="Balut Island" pageId="43" pageNumber="44" specimenCode="CASIZ 218871" specimenCount="150" typeStatus="paratype">
|
||
<specimenCount id="9D3C9C92FF9CFFB0D7228FE4E85E73EC" box="[1285,1367,692,717]" pageId="43" pageNumber="44" type="generic">150 m.</specimenCount>
|
||
; Coll.
|
||
<collectingMethod id="527B2F0CFF9CFFB0D2B08F87ECAC73CF" box="[151,421,727,752]" pageId="43" pageNumber="44">taken with tangle net</collectingMethod>
|
||
. 1 dry spec. R=3.7 r=1.1.
|
||
<emphasis id="B94E8B09FF9CFFB0D12F8F87EE8073D0" bold="true" box="[776,905,727,752]" pageId="43" pageNumber="44">
|
||
<typeStatus id="5481E9B9FF9CFFB0D12F8F87EE8D73D0" box="[776,900,727,752]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="DB9CFF60FF9CFFB0D1BC8F88E95C73D0" box="[923,1109,728,752]" collectionCode="CASIZ" pageId="43" pageNumber="44">
|
||
<collectionCode id="ED2BCFDEFF9CFFB0D1BC8F88EEE573D0" box="[923,1004,728,752]" pageId="43" pageNumber="44">CASIZ</collectionCode>
|
||
218871
|
||
</specimenCode>
|
||
;
|
||
<location id="8EE501C0FF9CFFB0D64C8F87E80873D0" LSID="urn:lsid:plazi:treatment:0393E60DFF9EFFB0D2B08DC7EC52727C:8EE501C0FF9CFFB0D64C8F87E80873D0" box="[1131,1281,727,752]" country="Philippines" name="Balut Island" pageId="43" pageNumber="44">Balut Island</location>
|
||
,
|
||
<collectingCountry id="F32D178BFF9CFFB0D7308F87E89173D0" box="[1303,1432,727,752]" name="Philippines" pageId="43" pageNumber="44">Philippines</collectingCountry>
|
||
, approximately
|
||
<quantity id="4CC2FAFEFF9CFFB0D3648FACECC37234" box="[323,458,764,789]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="43" pageNumber="44" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">
|
||
<elevation id="0017B028FF9CFFB0D3648FACECC37234" box="[323,458,764,789]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="43" pageNumber="44" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">
|
||
100–
|
||
<specimenCount id="9D3C9C92FF9CFFB0D35C8FACECC37234" box="[379,458,764,789]" pageId="43" pageNumber="44" type="generic">150 m</specimenCount>
|
||
</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.;
|
||
<materialsCitation id="3B525D46FF9CFFB0D3F88FACEC627217" ID-GBIF-Occurrence="1503739441" collectionCode="CASIZ" country="Philippines" elevation="200" location="Balut Island" pageId="43" pageNumber="44" specimenCode="CASIZ 219697" specimenCount="1" typeStatus="paratype">
|
||
Coll.
|
||
<collectingMethod id="527B2F0CFF9CFFB0D0078FACEE117234" box="[544,792,764,789]" pageId="43" pageNumber="44">taken with tangle net</collectingMethod>
|
||
; 1 dry spec. R=5.1 r=1.2.
|
||
<specimenCode id="DB9CFF60FF9CFFB0D6778FACE9F67235" box="[1104,1279,764,789]" collectionCode="CASIZ" pageId="43" pageNumber="44">
|
||
<collectionCode id="ED2BCFDEFF9CFFB0D6778FACE9A87234" box="[1104,1185,764,788]" pageId="43" pageNumber="44">CASIZ</collectionCode>
|
||
219697
|
||
</specimenCode>
|
||
<location id="8EE501C0FF9CFFB0D72D8FACE8917235" LSID="urn:lsid:plazi:treatment:0393E60DFF9EFFB0D2B08DC7EC52727C:8EE501C0FF9CFFB0D72D8FACE8917235" box="[1290,1432,764,789]" country="Philippines" name="Balut Island" pageId="43" pageNumber="44">Balut Island</location>
|
||
,
|
||
<collectingCountry id="F32D178BFF9CFFB0D2B08E4FEC1C7218" box="[151,277,799,824]" name="Philippines" pageId="43" pageNumber="44">Philippines</collectingCountry>
|
||
<quantity id="4CC2FAFEFF9CFFB0D3398E70EC627217" box="[286,363,800,824]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="m" value="200.0">
|
||
<specimenCount id="9D3C9C92FF9CFFB0D3398E70EC627217" box="[286,363,800,824]" pageId="43" pageNumber="44" type="generic">
|
||
<elevation id="0017B028FF9CFFB0D3398E70EC627217" box="[286,363,800,824]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="m" value="200.0">200 m</elevation>
|
||
</specimenCount>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B525D46FF9CFFB0D35F8E4FE9CB7217" ID-GBIF-Occurrence="1503739469" box="[376,1218,799,824]" collectionCode="CASIZ" country="Philippines" elevation="200" location="Balut Island" pageId="43" pageNumber="44" specimenCode="CASIZ 219698" specimenCount="1" typeStatus="paratype">
|
||
<collectingMethod id="527B2F0CFF9CFFB0D35F8E4FECFA7217" box="[376,499,799,824]" pageId="43" pageNumber="44">Tangle net</collectingMethod>
|
||
. R=4.5 r=1.3;
|
||
<specimenCode id="DB9CFF60FF9CFFB0D0BB8E70EE437218" box="[668,842,799,824]" collectionCode="CASIZ" pageId="43" pageNumber="44">
|
||
<collectionCode id="ED2BCFDEFF9CFFB0D0BB8E70EFE47218" box="[668,749,800,824]" pageId="43" pageNumber="44">CASIZ</collectionCode>
|
||
219698
|
||
</specimenCode>
|
||
<location id="8EE501C0FF9CFFB0D1748E4FEEE97218" LSID="urn:lsid:plazi:treatment:0393E60DFF9EFFB0D2B08DC7EC52727C:8EE501C0FF9CFFB0D1748E4FEEE97218" box="[851,992,799,824]" country="Philippines" name="Balut Island" pageId="43" pageNumber="44">Balut Island</location>
|
||
,
|
||
<collectingCountry id="F32D178BFF9CFFB0D1C98E4FE9657218" box="[1006,1132,799,824]" name="Philippines" pageId="43" pageNumber="44">Philippines</collectingCountry>
|
||
<quantity id="4CC2FAFEFF9CFFB0D6528E70E9CB7217" box="[1141,1218,800,824]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="m" value="200.0">
|
||
<specimenCount id="9D3C9C92FF9CFFB0D6528E70E9CB7217" box="[1141,1218,800,824]" pageId="43" pageNumber="44" type="generic">
|
||
<elevation id="0017B028FF9CFFB0D6528E70E9CB7217" box="[1141,1218,800,824]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="m" value="200.0">200 m</elevation>
|
||
</specimenCount>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.
|
||
<collectingMethod id="527B2F0CFF9CFFB0D6E88E4FE8437217" box="[1231,1354,799,824]" pageId="43" pageNumber="44">Tangle net</collectingMethod>
|
||
. R=4.5 r=1.0. 1 dry spec.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |