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<mods:title id="6D470ADEAED2D3ED127BB766AB6BC8D7">First confirmed record of the sesarmid crab, Parasesarma bengalense (Davie, 2003) (Decapoda: Brachyura) in Indian waters</mods:title>
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<mods:namePart id="DEFC2244A93B76916B1802072F30AF24">Pati, Sameer Kumar</mods:namePart>
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<mods:namePart id="98DEB36C2BA1F14180F0ABA8F12BC9A1">Rani, Varghese</mods:namePart>
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<mods:namePart id="1CDCD8DDAAF34D47B5C0D7BA9E70B52E">Sujila, Pallikkara Subrahmanian</mods:namePart>
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<mods:namePart id="FE1F93E642E28C22E1C3E675C8BC25D3">Nadan, Sivasankaran Bijoy</mods:namePart>
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<heading id="D0CD4F613D5D3B35FEF16114FD0A0F15" bold="true" box="[271,753,1111,1140]" fontSize="12" level="1" pageId="2" pageNumber="3" reason="0">
<taxonomicName id="4C3A838E3D5D3B35FEF16114FD0A0F15" ID-CoL="4DBY6" authority="(Davie, 2003)" authorityName="Davie" authorityYear="2003" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[271,753,1111,1140]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5D3B35FEF16114FD0A0F15" bold="true" box="[271,753,1111,1140]" pageId="2" pageNumber="3">
<emphasis id="B94E241F3D5D3B35FEF16114FDC20F15" bold="true" box="[271,569,1111,1140]" italics="true" pageId="2" pageNumber="3">Parasesarma bengalense</emphasis>
(
<bibRefCitation id="EFAB85FC3D5D3B35FDB6611BFD130F15" author="Davie" box="[584,744,1111,1140]" firstAuthor="Davie" journalOrPublisher="Raffles Bulletin of Zoology" pageId="2" pageNumber="3" pagination="387 - 391" part="51" refId="ref2963" refString="Davie, P. J. F. 2003. A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal. Raffles Bulletin of Zoology, 51: 387 - 391." title="A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal" type="journal article" year="2003">Davie, 2003</bibRefCitation>
)
</emphasis>
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<heading id="D0CD4F613D5D3B35FE2D61D9FDD50FD0" box="[467,558,1172,1202]" fontSize="12" level="2" pageId="2" pageNumber="3" reason="1">
(
<figureCitation id="1301E4883D5D3B35FE2161D8FDDB0FD3" box="[479,544,1172,1202]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
)
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<taxonomicName id="4C3A838E3D5D3B35FF4B61AFFD6A0F9E" ID-CoL="4F44C" authority="Davie, 2003: 388" authorityName="Davie" authorityPageNumber="388" authorityYear="2003" box="[181,657,1250,1279]" class="Malacostraca" family="Sesarmidae" genus="Perisesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5D3B35FF4B61AFFE460F9E" box="[181,445,1250,1279]" italics="true" pageId="2" pageNumber="3">Perisesarma bengalense</emphasis>
<treatmentCitation id="0A9BDE1C3D5D3B35FE3D61AEFD6A0F9E" author="Davie, P. J. F." box="[451,657,1250,1279]" page="388" pageId="2" pageNumber="3" year="2003">
<bibRefCitation id="EFAB85FC3D5D3B35FE3D61AEFD6A0F9E" author="Davie" box="[451,657,1250,1279]" firstAuthor="Davie" journalOrPublisher="Raffles Bulletin of Zoology" pageId="2" pageNumber="3" pagination="387 - 391" part="51" refId="ref2963" refString="Davie, P. J. F. 2003. A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal. Raffles Bulletin of Zoology, 51: 387 - 391." title="A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal" type="journal article" year="2003">Davie, 2003: 388</bibRefCitation>
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.
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<taxonomicName id="4C3A838E3D5D3B35FF4B6045FCDE0E47" authority="- Bouillon et al., 2004: 83" authorityName="Bouillon" authorityPageNumber="83" authorityYear="2004" box="[181,805,1288,1318]" class="Malacostraca" family="Sesarmidae" genus="Perisesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="B94E241F3D5D3B35FF4B6045FE360E44" box="[181,461,1288,1317]" italics="true" pageId="2" pageNumber="3">Perisesarma bengalensis</emphasis>
<treatmentCitation id="0A9BDE1C3D5D3B35FE026044FCDE0E47" author="Bouillon, S &amp; Moens, T &amp; Overmeer, I. &amp; Koedam, N. &amp; Dehairs, F." box="[508,805,1288,1318]" page="83" pageId="2" pageNumber="3" year="2004">
<bibRefCitation id="EFAB85FC3D5D3B35FE026044FCDE0E47" author="Bouillon" box="[508,805,1288,1318]" etAl="et al." firstAuthor="Bouillon" journalOrPublisher="Marine Ecology Progress Series" pageId="2" pageNumber="3" pagination="77 - 88" part="278" refId="ref2843" refString="Bouillon, S.; Moens, T.; Overmeer, I.; Koedam, N. and Dehairs, F. 2004. Resource utilization patterns of epifauna from mangrove forests with contrasting inputs of local versus imported organic matter. Marine Ecology Progress Series, 278: 77 - 88." title="Resource utilization patterns of epifauna from mangrove forests with contrasting inputs of local versus imported organic matter" type="journal article" year="2004">
Bouillon
<emphasis id="B94E241F3D5D3B35FD916045FD590E44" box="[623,674,1288,1317]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
., 2004: 83
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(list).
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<taxonomicName id="4C3A838E3D5D3B35FF4B601BFD230E12" ID-CoL="4F44C" authority="- Ng et al., 2008: 222" authorityName="Ng" authorityPageNumber="222" authorityYear="2008" box="[181,728,1366,1395]" class="Malacostraca" family="Sesarmidae" genus="Perisesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5D3B35FF4B601BFE450E12" box="[181,446,1366,1395]" italics="true" pageId="2" pageNumber="3">Perisesarma bengalense</emphasis>
<treatmentCitation id="0A9BDE1C3D5D3B35FE13601AFD230E12" author="Ng, P. K. L &amp; Guinot, D. &amp; Davie, P. J. F." box="[493,728,1366,1395]" page="222" pageId="2" pageNumber="3" year="2008">
<bibRefCitation id="EFAB85FC3D5D3B35FE13601AFD230E12" author="Ng" box="[493,728,1366,1395]" etAl="et al." firstAuthor="Ng" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="2" pageNumber="3" pagination="1 - 286" part="17" refId="ref3481" refString="Ng, P. K. L.; Guinot, D. and Davie, P. J. F. 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world" type="journal article" year="2008">
Ng
<emphasis id="B94E241F3D5D3B35FDE7601BFDAB0E12" box="[537,592,1366,1395]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
, 2008: 222
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(list);
<treatmentCitation id="0A9BDE1C3D5D3B35FF256030FE5C0EFB" author="Davie, P. J. F." box="[219,423,1404,1434]" page="197" pageId="2" pageNumber="3" year="2010">
<bibRefCitation id="EFAB85FC3D5D3B35FF256030FE5C0EFB" author="Davie" box="[219,423,1404,1434]" editor="Fransen &amp; S. De Grave &amp; P. K. L. Ng" firstAuthor="Davie" journalOrPublisher="Brill" pageId="2" pageNumber="3" pagination="195 - 207" refId="ref3002" refString="Davie, P. J. F. 2010. A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia. p. 195 - 207. In: C. Fransen; S. De Grave and P. K. L. Ng (eds), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14. Leiden, Brill." title="A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia" type="book chapter" volumeTitle="Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14" year="2010">Davie, 2010: 197</bibRefCitation>
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(list).
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<taxonomicName id="4C3A838E3D5D3B35FF4B60E8FEAE0E86" ID-CoL="4DBY6" authority="- Shahdadi and Schubart, 2017: 534" authorityName="- Shahdadi and Schubart" authorityPageNumber="534" authorityYear="2017" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5D3B35FF4B60E8FE340EA1" box="[181,463,1443,1472]" italics="true" pageId="2" pageNumber="3">Parasesarma bengalense</emphasis>
<treatmentCitation id="0A9BDE1C3D5D3B35FE0160EFFEAE0E86" author="Shahdadi, A. &amp; Schubart, C. D." page="534" pageId="2" pageNumber="3" year="2017">
<bibRefCitation id="EFAB85FC3D5D3B35FE0160EFFEAE0E86" author="Shahdadi and Schubart" firstAuthor="Shahdadi" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="2" pageNumber="3" pagination="517 - 548" part="182" refId="ref3761" refString="Shahdadi, A. and Schubart, C. D. 2017. Taxonomic review of Perisesarma (Decapoda: Brachyura: Sesarmidae) and closely related genera based on morphology and molecular phylogenetics: new classification, two new genera and the questionable phylogenetic value of the epibranchial tooth. Zoological Journal of the Linnean Society, 182: 517 - 548." title="Taxonomic review of Perisesarma (Decapoda: Brachyura: Sesarmidae) and closely related genera based on morphology and molecular phylogenetics: new classification, two new genera and the questionable phylogenetic value of the epibranchial tooth" type="journal article" year="2017">Shahdadi and Schubart, 2017: 534</bibRefCitation>
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(list and new combination).
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<emphasis id="B94E241F3D5D3B35FF25635BFE450D55" box="[219,446,1559,1588]" italics="true" pageId="2" pageNumber="3">Material examined</emphasis>
.
<materialsCitation id="3B52F2503D5D3B35FE32635BFCDA0DAE" ID-GBIF-Occurrence="2456362991" collectingDate="1991-04-09" collectionCode="N" collectorName="S. V. Mulay" country="India" latitude="17.164" location="Ganapatipule" longLatPrecision="76" longitude="73.259" municipality="Malgund" pageId="2" pageNumber="3" specimenCode="ZSI-WRC C.1286" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="MAHARASHTRA">
<collectingCountry id="F32DB89D3D5D3B35FE32635BFDD60D55" box="[460,557,1559,1588]" name="India" pageId="2" pageNumber="3" value="India">INDIA</collectingCountry>
:
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<specimenCount id="9D3C33843D5D3B35FF216372FEE70D3A" box="[223,284,1598,1627]" pageId="2" pageNumber="3" type="male">male</specimenCount>
(12.20 ×
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<specimenCount id="9D3C33843D5D3B35FDA46372FD550D3A" box="[602,686,1598,1627]" pageId="2" pageNumber="3" type="female">female</specimenCount>
(13.67 ×
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),
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north of
<location id="8EE5AED63D5D3B35FF4B63C7FEA90DC9" LSID="urn:lsid:plazi:treatment:0393491B3D5D3B33FEF16114FE1B0D80:8EE5AED63D5D3B35FF4B63C7FEA90DC9" box="[181,338,1675,1704]" country="India" latitude="17.164" longLatPrecision="76" longitude="73.259" municipality="Malgund" name="Ganapatipule" pageId="2" pageNumber="3" stateProvince="MAHARASHTRA">Ganapatipule</location>
,
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district (
<geoCoordinate id="EE0E9ECA3D5D3B35FDD163C7FD5B0DC9" box="[559,672,1675,1704]" degrees="17.164" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="17.164">17.164°N</geoCoordinate>
,
<geoCoordinate id="EE0E9ECA3D5D3B35FD5763C7FCE90DC9" box="[681,786,1675,1704]" degrees="73.259" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="73.259">73.259°E</geoCoordinate>
),
<date id="FF84DECD3D5D3B35FF4B63FEFEAF0DAE" box="[181,340,1714,1743]" pageId="2" pageNumber="3" value="1991-04-09">
<collectingDate id="EFC027253D5D3B35FF4B63FEFEAF0DAE" box="[181,340,1714,1743]" pageId="2" pageNumber="3" value="1991-04-09">9 April 1991</collectingDate>
</date>
, coll.
<collectorName id="26CF9DDB3D5D3B35FE6363FEFDE60DAE" box="[413,541,1714,1743]" pageId="2" pageNumber="3">S.V. Mulay</collectorName>
(
<specimenCode id="DB9C50763D5D3B35FDD363FEFCEC0DAE" box="[557,791,1714,1743]" pageId="2" pageNumber="3">ZSI-WRC C.1286</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B52F2503D5D3B35FF4B6395FBC40A64" ID-GBIF-Occurrence="2456362798" collectingDate="2018-10-21" collectingDateMax="2019-03-09" collectingDateMin="2018-10-21" collectionCode="N" collectorName="V. Rani" country="India" county="Kasaragod" latitude="12.261" location="Estuary" longLatPrecision="77" longitude="75.165" municipality="Narakkal" pageId="2" pageNumber="3" specimenCode="ZSI-WRC C.1914" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="KERALA">
<collectingRegion id="49FE36EF3D5D3B35FF4B6395FEDC0D97" box="[181,295,1753,1782]" country="India" name="Kerala" pageId="2" pageNumber="3">KERALA</collectingRegion>
<specimenCount id="9D3C33843D5D3B35FEAE6395FE5D0D94" box="[336,422,1752,1782]" pageId="2" pageNumber="3" type="male">2 males</specimenCount>
(16.10 ×
<quantity id="4CC255E83D5D3B35FDED6395FD760D94" box="[531,653,1753,1782]" metricMagnitude="-2" metricUnit="m" metricValue="1.294" pageId="2" pageNumber="3" unit="mm" value="12.94">12.94 mm</quantity>
; 11.87 ×
<quantity id="4CC255E83D5D3B35FD0C6394FF1D0C7D" metricMagnitude="-3" metricUnit="m" metricValue="9.86" pageId="2" pageNumber="3" unit="mm" value="9.86">9.86 mm</quantity>
) and
<specimenCount id="9D3C33843D5D3B35FEDC63B3FE950C7D" box="[290,366,1791,1820]" pageId="2" pageNumber="3" type="female">female</specimenCount>
(12.52 ×
<quantity id="4CC255E83D5D3B35FE2763B3FDAB0C7D" box="[473,592,1791,1820]" metricMagnitude="-2" metricUnit="m" metricValue="1.052" pageId="2" pageNumber="3" unit="mm" value="10.52">10.52 mm</quantity>
), Thejaswini River, near Palayi Kadavu, Kasaragod district (
<geoCoordinate id="EE0E9ECA3D5D3B35FD5A626AFCDB0C22" box="[676,800,1830,1859]" degrees="12.261" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="12.261">12.261°N</geoCoordinate>
,
<geoCoordinate id="EE0E9ECA3D5D3B35FF4B6200FECB0C08" box="[181,304,1868,1897]" degrees="75.165" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="75.165">75.165°E</geoCoordinate>
),
<date id="FF84DECD3D5D3B35FEB86201FDDE0C0B" box="[326,549,1868,1898]" pageId="2" pageNumber="3" value="2018-10-21">
<collectingDate id="EFC027253D5D3B35FEB86201FDDE0C0B" box="[326,549,1868,1898]" pageId="2" pageNumber="3" value="2018-10-21">21 October 2018</collectingDate>
</date>
, coll. P.S. Sujila and P.S. Lakshmi (ZSI-WRC C.1910);
<specimenCount id="9D3C33843D5D3B35FDA7623FFD4E0CF1" box="[601,693,1907,1936]" pageId="2" pageNumber="3" type="male">2 males</specimenCount>
(17.13 ×
<quantity id="4CC255E83D5D3B35FF4B62D6FECC0CD7" box="[181,311,1946,1975]" metricMagnitude="-2" metricUnit="m" metricValue="1.399" pageId="2" pageNumber="3" unit="mm" value="13.99">13.99 mm</quantity>
; 11.59 ×
<quantity id="4CC255E83D5D3B35FE5662D6FDE10CD6" box="[424,538,1946,1975]" metricMagnitude="-3" metricUnit="m" metricValue="9.52" pageId="2" pageNumber="3" unit="mm" value="9.52">9.52 mm</quantity>
), Valiyavattom Island, near
<collectingMunicipality id="6BE162773D5D3B35FC7665D5FC010BD7" box="[904,1018,153,182]" pageId="2" pageNumber="3">Narakkal</collectingMunicipality>
, Cochin
<location id="8EE5AED63D5D3B35FB9765D6FB330BD7" LSID="urn:lsid:plazi:treatment:0393491B3D5D3B33FEF16114FE1B0D80:8EE5AED63D5D3B35FB9765D6FB330BD7" box="[1129,1224,154,182]" country="Iceland" county="Kasaragod" latitude="12.261" longLatPrecision="77" longitude="75.165" municipality="Narakkal" name="Estuary" pageId="2" pageNumber="3" stateProvince="KERALA">Estuary</location>
,
<location id="8EE5AED63D5D3B35FB2B65D5FAA70BD7" LSID="urn:lsid:plazi:treatment:0393491B3D5D3B33FEF16114FE1B0D80:8EE5AED63D5D3B35FB2B65D5FAA70BD7" box="[1237,1372,153,182]" country="Iceland" county="Kasaragod" latitude="12.261" longLatPrecision="77" longitude="75.165" municipality="Narakkal" name="Ernakulam" pageId="2" pageNumber="3" stateProvince="KERALA">Ernakulam</location>
district (
<geoCoordinate id="EE0E9ECA3D5D3B35FCAA658DFC2B0BBF" box="[852,976,193,222]" degrees="10.044" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="10.044">10.044°N</geoCoordinate>
,
<geoCoordinate id="EE0E9ECA3D5D3B35FC22658CFBA80BBC" box="[988,1107,192,221]" degrees="76.234" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="76.234">76.234°E</geoCoordinate>
),
<date id="FF84DECD3D5D3B35FB96658DFAE20BBF" box="[1128,1305,192,222]" pageId="2" pageNumber="3" value="2019-03-09">
<collectingDate id="EFC027253D5D3B35FB96658DFAE20BBF" box="[1128,1305,192,222]" pageId="2" pageNumber="3" value="2019-03-09">9 March 2019</collectingDate>
</date>
, coll.
<collectorName id="26CF9DDB3D5D3B35FA9E658DFA400BBC" box="[1376,1467,192,221]" pageId="2" pageNumber="3">V. Rani</collectorName>
(
<specimenCode id="DB9C50763D5D3B35FCAD65A4FBCD0A64" box="[851,1078,232,261]" pageId="2" pageNumber="3">ZSI-WRC C.1914</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C320AB863D5D3B35FC8F647AFA9D0D6A" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8B85F80D3D5D3B35FC8F647AFA9D0D6A" blockId="2.[843,1467,309,1547]" pageId="2" pageNumber="3">
<emphasis id="B94E241F3D5D3B35FC8F647AFC270A33" box="[881,988,310,338]" italics="true" pageId="2" pageNumber="3">Diagnosis</emphasis>
. Carapace subrectangular, slightly broader than long (cw/cl 1.21.5); regions well defined; postfrontal region with 4 prominent, rounded lobes; medial lobes slightly broader than lateral lobes; lateral lobes not separated from inner orbital rim; front deflexed downwards, margin sinuous; anterolateral margin with large, acutely triangular external orbital tooth and smaller, acutely triangular epibranchial tooth, separated from each other by distinct notch; lateral margins subparallel, slightly concave, with numerous short setae; cornea slightly extending beyond tip of external orbital tooth (
<figureCitation id="1301E4883D5D3B35FB9E67AEFB4D099E" box="[1120,1206,738,767]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="2" pageNumber="3">Fig. 2A</figureCitation>
). Male cheliped with 2 transverse pectinate crests on upper surface of palm; primary pectinate crest with 1418 tall, broad corneous teeth; secondary pectinate crest shorter than primary pectinate crest, with 1315 relatively shorter corneous teeth; fingers serrated along cutting margins; fixed finger with strong subdistal tooth along cutting margin; dactylus with strong subdistal tooth and prominent medial tooth along cutting margin, and 1618 almost symmetrical, distinct tubercles on upper surface; proximal 110 dactylar tubercles closely spaced, transversely broadened; distal dactylar tubercles relatively widely spaced, rounded; outer surfaces of palm and proximal portion of dactylus relatively more coarsely granular (
<figureCitation id="1301E4883D5D3B35FADE6048FA5E0E40" box="[1312,1445,1284,1313]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="2" pageNumber="3">Fig. 2BE</figureCitation>
). Ambulatory legs moderately long, stout (
<figureCitation id="1301E4883D5D3B35FAB36066FA5C0E29" box="[1357,1447,1322,1352]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="2" pageNumber="3">Fig. 2A</figureCitation>
). Male pleon moderately broad; sixth pleonal somite approximately 2.1 times broader than long (
<figureCitation id="1301E4883D5D3B35FAAB6034FA5C0EF7" box="[1365,1447,1400,1430]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="2" pageNumber="3">Fig. 2F</figureCitation>
). G1 moderately stout, gently curved; apical process corneous, relatively shorter and stouter, bent at angle of about 60° from longitudinal axis (
<figureCitation id="1301E4883D5D3B35FB0560A1FAA90D6A" box="[1275,1362,1517,1547]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="2" pageNumber="3">Fig. 2G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320AB863D5D3B33FC8F6377FE6D0EB2" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8B85F80D3D5D3B35FC8F6377FAFA0C23" blockId="2.[843,1467,1595,1975]" pageId="2" pageNumber="3">
<emphasis id="B94E241F3D5D3B35FC8F6377FC2F0D39" box="[881,980,1595,1624]" italics="true" pageId="2" pageNumber="3">Remarks</emphasis>
.
<taxonomicName id="4C3A838E3D5D3B35FC1E6370FB100D39" authorityName="- Shahdadi and Schubart" authorityYear="2017" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[992,1259,1595,1624]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5D3B35FC1E6370FB100D39" box="[992,1259,1595,1624]" italics="true" pageId="2" pageNumber="3">Parasesarma bengalense</emphasis>
</taxonomicName>
mostly resembles
<taxonomicName id="4C3A838E3D5D3B35FCB5632FFB3C0D1E" authority="(Campbell, 1967)" authorityName="Campbell" authorityYear="1967" baseAuthorityName="Campbell" baseAuthorityYear="1967" box="[843,1223,1634,1663]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="darwinense">
<emphasis id="B94E241F3D5D3B35FCB5632FFC1E0D1E" box="[843,997,1634,1663]" italics="true" pageId="2" pageNumber="3">P. darwinense</emphasis>
(Campbell, 1967)
</taxonomicName>
, an endemic species of northwestern Australia, in terms of morphological characters (see
<bibRefCitation id="EFAB85FC3D5D3B35FBFC63FCFB6C0DAC" author="Davie" box="[1026,1175,1712,1741]" firstAuthor="Davie" journalOrPublisher="Raffles Bulletin of Zoology" pageId="2" pageNumber="3" pagination="387 - 391" part="51" refId="ref2963" refString="Davie, P. J. F. 2003. A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal. Raffles Bulletin of Zoology, 51: 387 - 391." title="A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal" type="journal article" year="2003">Davie, 2003</bibRefCitation>
).
<taxonomicName id="4C3A838E3D5D3B35FB5463FDFA410DAC" authorityName="- Shahdadi and Schubart" authorityYear="2017" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[1194,1466,1712,1741]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5D3B35FB5463FDFA410DAC" box="[1194,1466,1712,1741]" italics="true" pageId="2" pageNumber="3">Parasesarma bengalense</emphasis>
</taxonomicName>
is nevertheless distinguished from
<taxonomicName id="4C3A838E3D5D3B35FB086394FA6D0D95" baseAuthorityName="Campbell" baseAuthorityYear="1967" box="[1270,1430,1751,1780]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="darwinense">
<emphasis id="B94E241F3D5D3B35FB086394FA6D0D95" box="[1270,1430,1751,1780]" italics="true" pageId="2" pageNumber="3">P. darwinense</emphasis>
</taxonomicName>
by the characters of the male cheliped, male sixth pleonal somite, and G1 (
<bibRefCitation id="EFAB85FC3D5D3B35FBF16269FB5D0C23" author="Davie" box="[1039,1190,1829,1858]" firstAuthor="Davie" journalOrPublisher="Raffles Bulletin of Zoology" pageId="2" pageNumber="3" pagination="387 - 391" part="51" refId="ref2963" refString="Davie, P. J. F. 2003. A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal. Raffles Bulletin of Zoology, 51: 387 - 391." title="A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal" type="journal article" year="2003">Davie, 2003</bibRefCitation>
;
<bibRefCitation id="EFAB85FC3D5D3B35FB4E6269FB0F0C23" author="Davie" box="[1200,1268,1829,1858]" editor="Fransen &amp; S. De Grave &amp; P. K. L. Ng" firstAuthor="Davie" journalOrPublisher="Brill" pageId="2" pageNumber="3" pagination="195 - 207" refId="ref3002" refString="Davie, P. J. F. 2010. A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia. p. 195 - 207. In: C. Fransen; S. De Grave and P. K. L. Ng (eds), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14. Leiden, Brill." title="A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia" type="book chapter" volumeTitle="Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14" year="2010">2010</bibRefCitation>
).
</paragraph>
<paragraph id="8B85F80D3D5D3B33FC8F6200FCDE0F32" blockId="2.[843,1467,1595,1975]" lastBlockId="4.[181,806,153,1761]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
On the other hand, the molecular data of
<bibRefCitation id="EFAB85FC3D5D3B35FAB26200FBC50CF1" author="Shahdadi and Schubart" firstAuthor="Shahdadi" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="2" pageNumber="3" pagination="517 - 548" part="182" refId="ref3761" refString="Shahdadi, A. and Schubart, C. D. 2017. Taxonomic review of Perisesarma (Decapoda: Brachyura: Sesarmidae) and closely related genera based on morphology and molecular phylogenetics: new classification, two new genera and the questionable phylogenetic value of the epibranchial tooth. Zoological Journal of the Linnean Society, 182: 517 - 548." title="Taxonomic review of Perisesarma (Decapoda: Brachyura: Sesarmidae) and closely related genera based on morphology and molecular phylogenetics: new classification, two new genera and the questionable phylogenetic value of the epibranchial tooth" type="journal article" year="2017">Shahdadi and Schubart (2017)</bibRefCitation>
showed that
<taxonomicName id="4C3A838E3D5D3B35FB2A6238FA9B0CF1" authorityName="- Shahdadi and Schubart" authorityYear="2017" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[1236,1376,1907,1936]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5D3B35FB2A6238FA9B0CF1" box="[1236,1376,1907,1936]" italics="true" pageId="2" pageNumber="3">P.bengalense</emphasis>
</taxonomicName>
forms a distinct clade along with
<taxonomicName id="4C3A838E3D5D3B35FB9562D7FB310CD6" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[1131,1226,1946,1975]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="B94E241F3D5D3B35FB9562D7FB310CD6" box="[1131,1226,1946,1975]" italics="true" pageId="2" pageNumber="3">P. bidens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A838E3D5D3B33FB2B62D7FE6F0BD6" authority="(Rahayu and Davie, 2002)" authorityName="Rahayu and Davie" authorityYear="2002" baseAuthorityName="Rahayu and Davie" baseAuthorityYear="2002" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" lastPageId="4" lastPageNumber="5" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cricotum">
<emphasis id="B94E241F3D5D3B35FB2B62D7FAB40CD6" box="[1237,1359,1947,1975]" italics="true" pageId="2" pageNumber="3">P. cricotum</emphasis>
<bibRefCitation id="EFAB85FC3D5D3B33FAAA62D6FE6F0BD6" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">(Rahayu and Davie, 2002)</bibRefCitation>
</taxonomicName>
, and the more closely related
<taxonomicName id="4C3A838E3D5B3B33FCEF65D6FD8B0BBC" authority="(A. Milne-Edwards, 1869)" authorityName="A. Milne-Edwards" authorityYear="1869" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1869" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="guttatum">
<emphasis id="B94E241F3D5B3B33FCEF65D6FEE40BBC" italics="true" pageId="4" pageNumber="5">P. guttatum</emphasis>
(A. Milne-Edwards, 1869)
</taxonomicName>
. Among these, only
<taxonomicName id="4C3A838E3D5B3B33FF0E65ABFEAB0A65" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[240,336,231,260]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="B94E241F3D5B3B33FF0E65ABFEAB0A65" box="[240,336,231,260]" italics="true" pageId="4" pageNumber="5">P. bidens</emphasis>
</taxonomicName>
is known to occur in Indian waters with a wider distribution (see
<bibRefCitation id="EFAB85FC3D5B3B33FE0F6441FD1A0A4B" author="Trivedi" box="[497,737,269,298]" etAl="et al." firstAuthor="Trivedi" journalOrPublisher="Zootaxa" pageId="4" pageNumber="5" pagination="1 - 83" part="4502" refId="ref4069" refString="Trivedi, J. N.; Trivedi, D. J.; Vachhrajani, K. D. and Ng, P. K. L. 2018. An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India. Zootaxa, 4502: 1 - 83." title="An annotated checklist of the marine brachyuran crabs (Crustacea: Decapoda: Brachyura) of India" type="journal article" year="2018">
Trivedi
<emphasis id="B94E241F3D5B3B33FDAC6442FD7E0A4B" box="[594,645,269,298]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
., 2018
</bibRefCitation>
). The juvenile crabs of
<taxonomicName id="4C3A838E3D5B3B33FE866478FDFC0A30" authorityName="- Shahdadi and Schubart" authorityYear="2017" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[376,519,308,337]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5B3B33FE866478FDFC0A30" box="[376,519,308,337]" italics="true" pageId="4" pageNumber="5">P. bengalense</emphasis>
</taxonomicName>
are likely to be confused with
<taxonomicName id="4C3A838E3D5B3B33FF086417FEA50A16" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[246,350,346,375]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="B94E241F3D5B3B33FF086417FEA50A16" box="[246,350,346,375]" italics="true" pageId="4" pageNumber="5">P. bidens</emphasis>
</taxonomicName>
due to similarities in some carapace morphology. For instance, both the species have a prominent epibranchial tooth (
<figureCitation id="1301E4883D5B3B33FDD964EBFD800AA5" box="[551,635,423,452]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
; see
<bibRefCitation id="EFAB85FC3D5B3B33FD4B64E4FF0F0A8A" author="De Haan" editor="P. F. von Siebold" firstAuthor="De Haan" pageId="4" pageNumber="5" pagination="65 - 72" part="pl. 16" refId="ref3178" refString="De Haan, W. 1837. Crustacea. p. 65 - 72, pl. 16. In: P. F. von Siebold (ed), Fauna Japonica sive Descriptio Animalium, Quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suscepto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit. Leiden, Lugduni-Batavorum." title="Crustacea" type="book chapter" volumeTitle="Fauna Japonica sive Descriptio Animalium, Quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suscepto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit" year="1837">De Haan, 1837</bibRefCitation>
: pl. 16, fig.4), and the male sixth pleonal somite of
<taxonomicName id="4C3A838E3D5B3B33FF4B64B9FEBF0970" authorityName="- Shahdadi and Schubart" authorityYear="2017" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[181,324,500,529]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5B3B33FF4B64B9FEBF0970" box="[181,324,500,529]" italics="true" pageId="4" pageNumber="5">P.bengalense</emphasis>
</taxonomicName>
is almost equally broad as that of
<taxonomicName id="4C3A838E3D5B3B33FD3864B9FCDD0970" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[710,806,500,529]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="B94E241F3D5B3B33FD3864B9FCDD0970" box="[710,806,500,529]" italics="true" pageId="4" pageNumber="5">P. bidens</emphasis>
</taxonomicName>
(
<figureCitation id="1301E4883D5B3B33FF3F6756FEEB0959" box="[193,272,538,568]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="4" pageNumber="5">Fig. 2F</figureCitation>
; see
<bibRefCitation id="EFAB85FC3D5B3B33FEBB6756FE290959" author="Davie" box="[325,466,538,568]" editor="Fransen &amp; S. De Grave &amp; P. K. L. Ng" firstAuthor="Davie" journalOrPublisher="Brill" pageId="4" pageNumber="5" pagination="195 - 207" refId="ref3002" refString="Davie, P. J. F. 2010. A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia. p. 195 - 207. In: C. Fransen; S. De Grave and P. K. L. Ng (eds), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14. Leiden, Brill." title="A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia" type="book chapter" volumeTitle="Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14" year="2010">Davie, 2010</bibRefCitation>
;
<bibRefCitation id="EFAB85FC3D5B3B33FE256757FD710959" author="De Haan" box="[475,650,539,568]" editor="P. F. von Siebold" firstAuthor="De Haan" pageId="4" pageNumber="5" pagination="65 - 72" part="pl. 16" refId="ref3178" refString="De Haan, W. 1837. Crustacea. p. 65 - 72, pl. 16. In: P. F. von Siebold (ed), Fauna Japonica sive Descriptio Animalium, Quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suscepto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit. Leiden, Lugduni-Batavorum." title="Crustacea" type="book chapter" volumeTitle="Fauna Japonica sive Descriptio Animalium, Quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suscepto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit" year="1837">De Haan, 1837</bibRefCitation>
: pl. 16, fig.4). The male cheliped of both the species, in particular, has long transverse pectinate crests on the upper surface of the palm and distinct tubercles on the upper surface of the dactylus (
<figureCitation id="1301E4883D5B3B33FE7E67F8FDE709B0" box="[384,540,692,722]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="4" pageNumber="5">Fig. 2A, B, E</figureCitation>
; see
<bibRefCitation id="EFAB85FC3D5B3B33FDA567F8FD0F09B3" author="Davie" box="[603,756,692,722]" editor="Fransen &amp; S. De Grave &amp; P. K. L. Ng" firstAuthor="Davie" journalOrPublisher="Brill" pageId="4" pageNumber="5" pagination="195 - 207" refId="ref3002" refString="Davie, P. J. F. 2010. A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia. p. 195 - 207. In: C. Fransen; S. De Grave and P. K. L. Ng (eds), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14. Leiden, Brill." title="A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia" type="book chapter" volumeTitle="Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14" year="2010">Davie, 2010</bibRefCitation>
;
<bibRefCitation id="EFAB85FC3D5B3B33FCFE67F9FEBD0999" author="De Haan" editor="P. F. von Siebold" firstAuthor="De Haan" pageId="4" pageNumber="5" pagination="65 - 72" part="pl. 16" refId="ref3178" refString="De Haan, W. 1837. Crustacea. p. 65 - 72, pl. 16. In: P. F. von Siebold (ed), Fauna Japonica sive Descriptio Animalium, Quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suscepto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit. Leiden, Lugduni-Batavorum." title="Crustacea" type="book chapter" volumeTitle="Fauna Japonica sive Descriptio Animalium, Quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suscepto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit" year="1837">De Haan, 1837</bibRefCitation>
: pl. 16, fig. 4). The dactylar tubercles on the male cheliped, however, are greater in number (1618), and transversely broadened proximally and rounded distally in
<taxonomicName id="4C3A838E3D5B3B33FE706603FDE3080A" authorityName="- Shahdadi and Schubart" authorityYear="2017" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[398,536,846,875]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5B3B33FE706603FDE3080A" box="[398,536,846,875]" italics="true" pageId="4" pageNumber="5">P.bengalense</emphasis>
</taxonomicName>
(
<figureCitation id="1301E4883D5B3B33FDD66602FD8E080D" box="[552,629,846,876]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="4" pageNumber="5">Fig. 2E</figureCitation>
) whereas these tubercles are lesser in number (12 or 13) and oval in
<taxonomicName id="4C3A838E3D5B3B33FCEC663AFF0708D9" baseAuthorityName="De Haan" baseAuthorityYear="1835" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="B94E241F3D5B3B33FCEC663AFF0708D9" italics="true" pageId="4" pageNumber="5">P. bidens</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EFAB85FC3D5B3B33FED366D7FE3B08D8" author="Davie" box="[301,448,923,953]" editor="Fransen &amp; S. De Grave &amp; P. K. L. Ng" firstAuthor="Davie" journalOrPublisher="Brill" pageId="4" pageNumber="5" pagination="195 - 207" refId="ref3002" refString="Davie, P. J. F. 2010. A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia. p. 195 - 207. In: C. Fransen; S. De Grave and P. K. L. Ng (eds), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14. Leiden, Brill." title="A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia" type="book chapter" volumeTitle="Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14" year="2010">Davie, 2010</bibRefCitation>
). In addition, both the primary and secondary transverse crests on the palm of the male cheliped are relatively shorter (1418 and 1315 teeth, respectively) in
<taxonomicName id="4C3A838E3D5B3B33FE90615CFE060F4D" authorityName="- Shahdadi and Schubart" authorityYear="2017" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[366,509,1039,1068]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5B3B33FE90615CFE060F4D" box="[366,509,1039,1068]" italics="true" pageId="4" pageNumber="5">P. bengalense</emphasis>
</taxonomicName>
(
<figureCitation id="1301E4883D5B3B33FDF06143FDA50F4D" box="[526,606,1039,1068]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="4" pageNumber="5">Fig. 2B</figureCitation>
) than in
<taxonomicName id="4C3A838E3D5B3B33FD38615CFCDE0F4D" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[710,805,1039,1068]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="B94E241F3D5B3B33FD38615CFCDE0F4D" box="[710,805,1039,1068]" italics="true" pageId="4" pageNumber="5">P. bidens</emphasis>
</taxonomicName>
(1922 and 1520 teeth, respectively) (
<bibRefCitation id="EFAB85FC3D5B3B33FD7C6179FCE20F32" author="Davie" box="[642,793,1077,1107]" editor="Fransen &amp; S. De Grave &amp; P. K. L. Ng" firstAuthor="Davie" journalOrPublisher="Brill" pageId="4" pageNumber="5" pagination="195 - 207" refId="ref3002" refString="Davie, P. J. F. 2010. A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia. p. 195 - 207. In: C. Fransen; S. De Grave and P. K. L. Ng (eds), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14. Leiden, Brill." title="A new species of Perisesarma (Brachyura, Sesarmidae) from Western Australia" type="book chapter" volumeTitle="Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustacean Monographs, Volume 14" year="2010">Davie, 2010</bibRefCitation>
).
</paragraph>
<caption id="DF45A8853D5C3B34FF4B6271FC1E0CD4" ID-DOI="http://doi.org/10.5281/zenodo.3550119" ID-Zenodo-Dep="3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="3" pageNumber="4" startId="3.[181,250,1853,1876]" targetBox="[233,1415,151,1823]" targetPageId="3">
<paragraph id="8B85F80D3D5C3B34FF4B6271FC1E0CD4" blockId="3.[181,1467,1853,1973]" pageId="3" pageNumber="4">
<emphasis id="B94E241F3D5C3B34FF4B6271FEEF0C35" bold="true" box="[181,276,1853,1876]" pageId="3" pageNumber="4">Figure 2.</emphasis>
<taxonomicName id="4C3A838E3D5C3B34FEE26272FD6E0C34" authority="(Davie, 2003)" authorityName="Davie" authorityYear="2003" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[284,661,1853,1877]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5C3B34FEE26272FDFB0C34" box="[284,512,1853,1877]" italics="true" pageId="3" pageNumber="4">Parasesarma bengalense</emphasis>
(
<bibRefCitation id="EFAB85FC3D5C3B34FDF06271FD760C34" author="Davie" box="[526,653,1853,1877]" firstAuthor="Davie" journalOrPublisher="Raffles Bulletin of Zoology" pageId="3" pageNumber="4" pagination="387 - 391" part="51" refId="ref2963" refString="Davie, P. J. F. 2003. A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal. Raffles Bulletin of Zoology, 51: 387 - 391." title="A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal" type="journal article" year="2003">Davie, 2003</bibRefCitation>
)
</taxonomicName>
, male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C).
</paragraph>
</caption>
<paragraph id="8B85F80D3D5B3B33FF256111FE6D0EB2" blockId="4.[181,806,153,1761]" pageId="4" pageNumber="5">
<taxonomicName id="4C3A838E3D5B3B33FF256111FE1D0F18" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[219,486,1116,1145]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5B3B33FF256111FE1D0F18" box="[219,486,1116,1145]" italics="true" pageId="4" pageNumber="5">Parasesarma bengalense</emphasis>
</taxonomicName>
is easily distinguished from the other Indian species of
<taxonomicName id="4C3A838E3D5B3B33FE0261CFFD750FFE" authorityName="De Man" authorityYear="1895" box="[508,654,1155,1183]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B94E241F3D5B3B33FE0261CFFD750FFE" box="[508,654,1155,1183]" italics="true" pageId="4" pageNumber="5">Parasesarma</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3A838E3D5B3B33FD5C61CFFCE50FFE" baseAuthorityName="Heller" baseAuthorityYear="1865" box="[674,798,1155,1183]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="asperum">
<emphasis id="B94E241F3D5B3B33FD5C61CFFCE50FFE" box="[674,798,1155,1183]" italics="true" pageId="4" pageNumber="5">P. asperum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A838E3D5B3B33FF4B61E6FECC0FA7" authorityName="Naderloo and Schubart" authorityYear="2010" box="[181,311,1194,1222]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="persicum">
<emphasis id="B94E241F3D5B3B33FF4B61E6FECC0FA7" box="[181,311,1194,1222]" italics="true" pageId="4" pageNumber="5">P. persicum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A838E3D5B3B33FEBB61E6FE4B0FA7" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[325,432,1194,1222]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pictum">
<emphasis id="B94E241F3D5B3B33FEBB61E6FE4B0FA7" box="[325,432,1194,1222]" italics="true" pageId="4" pageNumber="5">P. pictum</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C3A838E3D5B3B33FE0A61E6FD8F0FA7" baseAuthorityName="Latreille" baseAuthorityYear="1803" box="[500,628,1193,1222]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="plicatum">
<emphasis id="B94E241F3D5B3B33FE0A61E6FD8F0FA7" box="[500,628,1193,1222]" italics="true" pageId="4" pageNumber="5">P. plicatum</emphasis>
</taxonomicName>
) mainly by its prominent epibranchial tooth with a distinct notch (
<figureCitation id="1301E4883D5B3B33FF3C61BAFEE00E72" box="[194,283,1270,1299]" captionStart="Figure 2" captionStartId="3.[181,250,1853,1876]" captionTargetBox="[233,1415,151,1823]" captionTargetId="figure@3.[233,1415,151,1823]" captionTargetPageId="3" captionText="Figure 2. Parasesarma bengalense (Davie, 2003), male (16.10 × 12.94 mm) (ZSI-WRC C.1910). A, overall dorsal view; B, upper view of transverse crests on palm of right chela; C, frontal view of primary transverse crest on palm of right chela; D, outer view of right cheliped; E, dorsal view of dactylar tubercles on right cheliped; F, male pleonal somites 46 and telson; G, leff G1 denuded of setae. Scale bars = 10 mm (A), 4 mm (D), 2 mm (E, F), 1 mm (B, G), 0.5 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.3550119" httpUri="https://zenodo.org/record/3550119/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
) (
<emphasis id="B94E241F3D5B3B33FEC361BBFEAF0E72" box="[317,340,1271,1299]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
. indistinct epibranchial tooth with an indistinct notch or mostly with a small prominence; see
<bibRefCitation id="EFAB85FC3D5B3B33FF1C600FFE650E01" author="De Haan" box="[226,414,1347,1376]" editor="P. F. von Siebold" firstAuthor="De Haan" pageId="4" pageNumber="5" pagination="65 - 72" part="pl. 16" refId="ref3178" refString="De Haan, W. 1837. Crustacea. p. 65 - 72, pl. 16. In: P. F. von Siebold (ed), Fauna Japonica sive Descriptio Animalium, Quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suscepto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit. Leiden, Lugduni-Batavorum." title="Crustacea" type="book chapter" volumeTitle="Fauna Japonica sive Descriptio Animalium, Quae in Itinere per Japoniam, Jussu et Auspiciis Superiorum, qui Summum in India Batava Imperium Tenent, Suscepto, Annis 1823 - 1830 Collegit, Notis, Observationibus et Adumbrationibus Illustravit" year="1837">De Haan, 1837</bibRefCitation>
: pl. 16, fig. 6;
<bibRefCitation id="EFAB85FC3D5B3B33FDBB600FFD240E01" author="Heller" box="[581,735,1347,1376]" firstAuthor="Heller" pageId="4" pageNumber="5" pagination="1 - 280" refId="ref3300" refString="Heller, C. 1865. Crustaceen. p. 1 - 280. In: Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf- Urbair. Zoologischer Theil. Zweiter Band. III. Abtheilung. Wien, Kaiserlich-Koniglichen Hof- und Staatsdruckerei, in Commission bei Karl Gerold's Sohn." title="Crustaceen" type="book chapter" volumeTitle="Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf- Urbair. Zoologischer Theil. Zweiter Band. III. Abtheilung. Wien, Kaiserlich-Koniglichen Hof- und Staatsdruckerei, in Commission bei Karl Gerold's Sohn" year="1865">Heller, 1865</bibRefCitation>
: pl. 6, fig. 1;
<bibRefCitation id="EFAB85FC3D5B3B33FF036025FD980EE6" author="Naderloo and Schubart" box="[253,611,1385,1415]" firstAuthor="Naderloo" journalOrPublisher="Zoologischer Anzeiger" pageId="4" pageNumber="5" pagination="33 - 43" part="249" refId="ref3431" refString="Naderloo, R. and Schubart, C. D. 2010. Description of a new species of Parasesarma (Crustacea; Decapoda; Brachyura; Sesarmidae) from the Persian Gulf, based on morphological and genetic characteristics. Zoologischer Anzeiger, 249: 33 - 43." title="Description of a new species of Parasesarma (Crustacea; Decapoda; Brachyura; Sesarmidae) from the Persian Gulf, based on morphological and genetic characteristics" type="journal article" year="2010">Naderloo and Schubart, 2010</bibRefCitation>
: fig. 2A;
<bibRefCitation id="EFAB85FC3D5B3B33FD346025FEA00ECC" author="Rahayu and Ng" firstAuthor="Rahayu" journalOrPublisher="Zootaxa" pageId="4" pageNumber="5" pagination="1 - 22" part="2327" refId="ref3715" refString="Rahayu, D. L. and Ng, P. K. L. 2010. Revision of the Parasesarma plicatum (Latreille, 1803) species-group (Crustacea: Decapoda: Brachyura: Sesarmidae). Zootaxa, 2327: 1 - 22." title="Revision of the Parasesarma plicatum (Latreille, 1803) species-group (Crustacea: Decapoda: Brachyura: Sesarmidae)" type="journal article" year="2010">Rahayu and Ng, 2010</bibRefCitation>
: fig. 1A;
<bibRefCitation id="EFAB85FC3D5B3B33FE3E60DCFCDA0ECC" author="Shahdadi and Schubart" box="[448,801,1424,1453]" firstAuthor="Shahdadi" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="4" pageNumber="5" pagination="517 - 548" part="182" refId="ref3761" refString="Shahdadi, A. and Schubart, C. D. 2017. Taxonomic review of Perisesarma (Decapoda: Brachyura: Sesarmidae) and closely related genera based on morphology and molecular phylogenetics: new classification, two new genera and the questionable phylogenetic value of the epibranchial tooth. Zoological Journal of the Linnean Society, 182: 517 - 548." title="Taxonomic review of Perisesarma (Decapoda: Brachyura: Sesarmidae) and closely related genera based on morphology and molecular phylogenetics: new classification, two new genera and the questionable phylogenetic value of the epibranchial tooth" type="journal article" year="2017">Shahdadi and Schubart, 2017</bibRefCitation>
: figs. 1A, C, 2A, C).
</paragraph>
</subSubSection>
<subSubSection id="C320AB863D5B3B33FF256091FE1B0D80" pageId="4" pageNumber="5" type="distribution">
<paragraph id="8B85F80D3D5B3B33FF256091FE1B0D80" blockId="4.[181,806,153,1761]" pageId="4" pageNumber="5">
<materialsCitation id="3B52F2503D5B3B33FF256091FE210D80" ID-GBIF-Occurrence="2456362799" country="India" location="Maharashtra" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Maharashtra">
<emphasis id="B94E241F3D5B3B33FF256091FDF90E9B" box="[219,514,1501,1530]" italics="true" pageId="4" pageNumber="5">Geographical distribution</emphasis>
.
<taxonomicName id="4C3A838E3D5B3B33FDEE6092FCDE0E9B" baseAuthorityName="Davie" baseAuthorityYear="2003" box="[528,805,1501,1530]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bengalense">
<emphasis id="B94E241F3D5B3B33FDEE6092FCDE0E9B" box="[528,805,1501,1530]" italics="true" pageId="4" pageNumber="5">Parasesarma bengalense</emphasis>
</taxonomicName>
is currently known from
<collectingCountry id="F32DB89D3D5B3B33FE1E634FFDE50D41" box="[480,542,1539,1568]" name="India" pageId="4" pageNumber="5" value="India">India</collectingCountry>
(
<collectingRegion id="49FE36EF3D5B3B33FDD0634FFD370D41" box="[558,716,1539,1568]" country="India" name="Maharashtra" pageId="4" pageNumber="5">Maharashtra</collectingRegion>
,
<collectingRegion id="49FE36EF3D5B3B33FD29634FFCDF0D41" box="[727,804,1539,1568]" country="India" name="Kerala" pageId="4" pageNumber="5">Kerala</collectingRegion>
and possibly
<collectingRegion id="49FE36EF3D5B3B33FEAF6366FDE80D26" box="[337,531,1578,1607]" country="India" name="Andhra Pradesh" pageId="4" pageNumber="5">Andhra Pradesh</collectingRegion>
states) (present study;
<bibRefCitation id="EFAB85FC3D5B3B33FF4B631CFE610D0C" author="Bouillon" box="[181,410,1616,1645]" etAl="et al." firstAuthor="Bouillon" journalOrPublisher="Marine Ecology Progress Series" pageId="4" pageNumber="5" pagination="77 - 88" part="278" refId="ref2843" refString="Bouillon, S.; Moens, T.; Overmeer, I.; Koedam, N. and Dehairs, F. 2004. Resource utilization patterns of epifauna from mangrove forests with contrasting inputs of local versus imported organic matter. Marine Ecology Progress Series, 278: 77 - 88." title="Resource utilization patterns of epifauna from mangrove forests with contrasting inputs of local versus imported organic matter" type="journal article" year="2004">
Bouillon
<emphasis id="B94E241F3D5B3B33FEE0631DFEB20D0C" box="[286,329,1616,1645]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
., 2004
</bibRefCitation>
),
<collectingCountry id="F32DB89D3D5B3B33FE52631CFDEC0D0C" box="[428,535,1616,1645]" name="Sri Lanka" pageId="4" pageNumber="5" value="Sri Lanka">Sri Lanka</collectingCountry>
(Southern and Western provinces) (
<bibRefCitation id="EFAB85FC3D5B3B33FEB3633BFE0B0DF5" author="Davie" box="[333,496,1655,1684]" firstAuthor="Davie" journalOrPublisher="Raffles Bulletin of Zoology" pageId="4" pageNumber="5" pagination="387 - 391" part="51" refId="ref2963" refString="Davie, P. J. F. 2003. A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal. Raffles Bulletin of Zoology, 51: 387 - 391." title="A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal" type="journal article" year="2003">Davie, 2003</bibRefCitation>
), and
<collectingCountry id="F32DB89D3D5B3B33FDC0633BFD570DF5" box="[574,684,1655,1684]" name="Thailand" pageId="4" pageNumber="5" value="Thailand">Thailand</collectingCountry>
(
<collectingRegion id="49FE36EF3D5B3B33FD40633BFCDE0DF5" box="[702,805,1655,1684]" country="Thailand" name="Ranong" pageId="4" pageNumber="5">Ranong</collectingRegion>
and
<collectingRegion id="49FE36EF3D5B3B33FF1463D1FEBB0DDB" box="[234,320,1693,1722]" country="Thailand" name="Phuket" pageId="4" pageNumber="5">Phuket</collectingRegion>
provinces) (
<bibRefCitation id="EFAB85FC3D5B3B33FE2763D1FD8F0DDB" author="Davie" box="[473,628,1693,1722]" firstAuthor="Davie" journalOrPublisher="Raffles Bulletin of Zoology" pageId="4" pageNumber="5" pagination="387 - 391" part="51" refId="ref2963" refString="Davie, P. J. F. 2003. A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal. Raffles Bulletin of Zoology, 51: 387 - 391." title="A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal" type="journal article" year="2003">Davie, 2003</bibRefCitation>
;
<bibRefCitation id="EFAB85FC3D5B3B33FD8163D1FE940D80" author="Shahdadi and Schubart" firstAuthor="Shahdadi" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="4" pageNumber="5" pagination="517 - 548" part="182" refId="ref3761" refString="Shahdadi, A. and Schubart, C. D. 2017. Taxonomic review of Perisesarma (Decapoda: Brachyura: Sesarmidae) and closely related genera based on morphology and molecular phylogenetics: new classification, two new genera and the questionable phylogenetic value of the epibranchial tooth. Zoological Journal of the Linnean Society, 182: 517 - 548." title="Taxonomic review of Perisesarma (Decapoda: Brachyura: Sesarmidae) and closely related genera based on morphology and molecular phylogenetics: new classification, two new genera and the questionable phylogenetic value of the epibranchial tooth" type="journal article" year="2017">Shahdadi and Schubart, 2017</bibRefCitation>
) (
<figureCitation id="1301E4883D5B3B33FE756388FE370D80" box="[395,460,1732,1761]" captionStart="Figure 1" captionStartId="1.[181,250,1853,1876]" captionTargetBox="[300,1344,1034,1816]" captionTargetId="figure@1.[300,1348,1021,1816]" captionTargetPageId="1" captionText="Figure 1. Map showing distribution of Parasesarma bengalense(Davie,2003).1, Malgund (Maharashtra, India); 2, Coringa (Andhra Pradesh, India); 3, Palayi Kadavu (Kerala, India); 4, Valiyavattom (Kerala, India); 5, Ranong (Thailand); 6, Phuket (Thailand); 7, Colombo (Sri Lanka); 8, Galle (Sri Lanka). Circle, present records; diamond, unconfirmed record; square, past records; triangle, type locality." figureDoi="http://doi.org/10.5281/zenodo.3550117" httpUri="https://zenodo.org/record/3550117/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>