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<document id="D6FD9816A671647EB81880C7E451E32E" ID-DOI="10.5852/ejt.2024.957.2651" ID-ISSN="2118-9773" ID-Zenodo-Dep="13773385" ID-ZooBank="9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726582870537" checkinUser="plazi" docAuthor="Selnekovič, Dávid, Jäch, Manfred A. &amp; Kodada, Ján" docDate="2024" docId="038B5001DABFE7F4FDDFF8BAFE572783" docLanguage="en" docName="EJT.2024.957.1-229.pdf" docOrigin="European Journal of Taxonomy 957" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Elmomorphus vietnamensis Selnekovič, Jäch &amp; Kodada, 2024, sp. nov." docType="treatment" docUuid="64F099FB-5D19-4F60-9820-06D6556B9D2E" docUuidSource="ZooBank" docVersion="2" lastPageNumber="188" masterDocId="FFB22879DA08E74FFF9EFFBAFFE32440" masterDocTitle="Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)" masterLastPageNumber="229" masterPageNumber="1" pageNumber="184" updateTime="1726584008522" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="AA1D1B3923A88A8A160F24E51F71BB0A">
<mods:title id="8F8807DC248CFDFA99632CD801692910">Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)</mods:title>
</mods:titleInfo>
<mods:name id="8975A2EAC96A06DFF9F46AA2E4BDEBE3" type="personal">
<mods:role id="37695E75EFA8B10602BF7F5375866C5E">
<mods:roleTerm id="EB503DF3989781FD9ED257F583A307D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7BFBC2F62E6F39C453E4C069F71BE0FB">Selnekovič, Dávid</mods:namePart>
<mods:nameIdentifier id="E4D2D298DA6D263D3A53C6D980B09288" type="LSID">D930D757-D929-4248-A29C-A857E0505345</mods:nameIdentifier>
<mods:affiliation id="B9FF2A0FCD72D8B4647F2C796395F669">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="579F87246DB8E5113AA052669A774FCF" type="email">david.selnekovic@uniba.sk</mods:nameIdentifier>
</mods:name>
<mods:name id="39BEBFA95581F27A0B914BD3E7728B47" type="personal">
<mods:role id="1A36A111EFCACD8AF62B00D51386F3FE">
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<mods:namePart id="3003CBE05E4C4D21327BAB5CAE799EA2">Jäch, Manfred A.</mods:namePart>
<mods:nameIdentifier id="24F0977AC33417AC174934A75B828A7B" type="LSID">D749707A-8823-4110-8D51-BF5AF4E2820F</mods:nameIdentifier>
<mods:affiliation id="6623F88C0EE773BE26E264FD0916C8C6">Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria.</mods:affiliation>
<mods:nameIdentifier id="B0E6E729CD52B49EC2EB26C92C5F00F5" type="email">manfred.jaech@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="20749D28D65B904BA5172CAA80707388">Kodada, Ján</mods:namePart>
<mods:nameIdentifier id="B2B3305C355FAA2464B0C06A0C646F2C" type="LSID">6E88BFBB-8769-44EC-8285-29E357CEC064</mods:nameIdentifier>
<mods:affiliation id="0FBD3BE4929796D80C5A434F7E138434">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
<mods:nameIdentifier id="893E40E54C95F5A1E74F1B20B50E9C06" type="email">jan.kodada@uniba.sk</mods:nameIdentifier>
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<mods:title id="3AEDDD34CC957828B0519784895E5E97">European Journal of Taxonomy</mods:title>
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<mods:part id="F23B7AC3F554357A9EA43F50B185101C">
<mods:date id="1246E1753F032E798B9CAF22124E8957">2024</mods:date>
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<mods:number id="13BB4F6C70E756E661A165299B2F028C">2024-09-11</mods:number>
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<mods:number id="B523B1DBBC6ADF7293B49D92063A4607">957</mods:number>
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<mods:identifier id="AFA08C5CDFF260F00EF63116E9B82636" type="DOI">10.5852/ejt.2024.957.2651</mods:identifier>
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<treatment id="038B5001DABFE7F4FDDFF8BAFE572783" ID-DOI="http://doi.org/10.5281/zenodo.13773662" ID-Zenodo-Dep="13773662" LSID="urn:lsid:zoobank.org:act:64F099FB-5D19-4F60-9820-06D6556B9D2E" httpUri="http://treatment.plazi.org/id/038B5001DABFE7F4FDDFF8BAFE572783" lastPageId="187" lastPageNumber="188" pageId="183" pageNumber="184">
<subSubSection id="C338B29CDABFE7F8FDDFF8BAFB5D237C" pageId="183" pageNumber="184" type="nomenclature">
<paragraph id="8B9DE117DABFE7F8FDDFF8BAFC10235B" blockId="183.[374,1214,1792,1884]" box="[577,1011,1792,1819]" pageId="183" pageNumber="184">
<heading id="D0D5567BDABFE7F8FDDFF8BAFC10235B" box="[577,1011,1792,1819]" centered="true" fontSize="11" level="2" pageId="183" pageNumber="184" reason="2">
<taxonomicName id="4C229A94DABFE7F8FDDFF8BAFC72235A" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[577,913,1792,1819]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="B9563D05DABFE7F8FDDFF8BAFC72235A" bold="true" box="[577,913,1792,1819]" italics="true" pageId="183" pageNumber="184">Elmomorphus vietnamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDABFE7F8FC07F8BBFC10235B" box="[921,1011,1793,1819]" pageId="183" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B9DE117DABFE7F8FEE8F89BFB5D237C" blockId="183.[374,1214,1792,1884]" box="[374,1214,1825,1852]" pageId="183" pageNumber="184">
<uri id="FFB3ED15DABFE7F8FEE8F89BFB5D237C" box="[374,1214,1825,1852]" pageId="183" pageNumber="184">
urn:lsid:zoobank.org:act:
<uuid id="FF84DBC2DABFE7F8FD08F89BFB5D237C" box="[662,1214,1825,1852]" pageId="183" pageNumber="184">64F099FB-5D19-4F60-9820-06D6556B9D2E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDABFE7F8FD06F8F8FC78231C" box="[664,923,1858,1884]" pageId="183" pageNumber="184" type="description">
<paragraph id="8B9DE117DABFE7F8FD06F8F8FC78231C" blockId="183.[374,1214,1792,1884]" box="[664,923,1858,1884]" pageId="183" pageNumber="184">
<figureCitation id="1319FD92DABFE7F8FD06F8F8FD11231C" box="[664,754,1858,1884]" captionStart="Fig" captionStartId="184.[189,232,1966,1992]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@184.[189,1399,265,1928]" captionTargetPageId="184" captionText="Fig. 90. Elmomorphus vietnamensis sp. nov., holotype, male (NMW), TL: 3.27 mm" figureDoi="http://doi.org/10.5281/zenodo.13773648" httpUri="https://zenodo.org/record/13773648/files/figure.png" pageId="183" pageNumber="184">Figs 90</figureCitation>
,
<figureCitation id="1319FD92DABFE7F8FD60F8F8FCB0231C" box="[766,851,1858,1884]" captionStart="Fig" captionStartId="185.[189,232,1927,1953]" captionTargetBox="[244,1347,276,1877]" captionTargetId="figure-36@185.[229,1357,265,1897]" captionTargetPageId="185" captionText="Fig. 91. A. Elmomorphus schoenmanni sp. nov., holotype (IAECAS), aedeagus, lateral aspect. B. E. vietnamensis sp. nov., paratype from type locality (CKB), aedeagus, lateral aspect. C. Same, ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773650" httpUri="https://zenodo.org/record/13773650/files/figure.png" pageId="183" pageNumber="184">91B–C</figureCitation>
,
<figureCitation id="1319FD92DABFE7F8FCFEF8F8FC78231C" box="[864,923,1858,1884]" captionStart="Fig" captionStartId="222.[189,232,1867,1893]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-44@222.[189,1399,265,1830]" captionTargetPageId="222" captionText="Fig. 114. Distribution maps. A. Elmomorphus globosus sp. nov. B. E. ovalis Kodada, Selnekovič &amp; Jäch sp. nov. C. E. minutus sp. nov. D. E. jii sp. nov. E. E. schoenmanni sp. nov. F. E. vietnamensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773722" httpUri="https://zenodo.org/record/13773722/files/figure.png" pageId="183" pageNumber="184">114F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDABFE7F5FF23F83BFD28265A" lastPageId="186" lastPageNumber="187" pageId="183" pageNumber="184" type="diagnosis">
<paragraph id="8B9DE117DABFE7F8FF23F83BFE2A23DB" blockId="183.[189,457,1921,1947]" box="[189,457,1921,1947]" pageId="183" pageNumber="184">
<heading id="D0D5567BDABFE7F8FF23F83BFE2A23DB" bold="true" box="[189,457,1921,1947]" fontSize="11" level="3" pageId="183" pageNumber="184" reason="3">
<emphasis id="B9563D05DABFE7F8FF23F83BFE2A23DB" bold="true" box="[189,457,1921,1947]" pageId="183" pageNumber="184">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DABFE7F5FF23F817FD28265A" blockId="183.[189,1399,1965,2025]" lastBlockId="186.[187,1399,265,538]" lastPageId="186" lastPageNumber="187" pageId="183" pageNumber="184">
<taxonomicName id="4C229A94DABFE7F8FF23F817FDE22387" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[189,513,1965,1991]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="183" pageNumber="184" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="B9563D05DABFE7F8FF23F817FDE22387" box="[189,513,1965,1991]" italics="true" pageId="183" pageNumber="184">Elmomorphus vietnamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDABFE7F8FD96F814FD802388" box="[520,611,1966,1992]" pageId="183" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1319FD92DABFE7F8FDEDF817FD2A2388" box="[627,713,1965,1992]" captionStart="Fig" captionStartId="184.[189,232,1966,1992]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@184.[189,1399,265,1928]" captionTargetPageId="184" captionText="Fig. 90. Elmomorphus vietnamensis sp. nov., holotype, male (NMW), TL: 3.27 mm" figureDoi="http://doi.org/10.5281/zenodo.13773648" httpUri="https://zenodo.org/record/13773648/files/figure.png" pageId="183" pageNumber="184">Fig. 90</figureCitation>
) is characterised by having the dorsal plastron confined to small areas on the frontoclypeus around the antennal insertions. The elytral striae are composed of large, deeply impressed punctures. The pronotal surface is smooth, not microreticulate as in
<taxonomicName id="4C229A94DAB2E7F5FB30FEB0FA942563" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1198,1399,265,291]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="schillhammeri" status="sp. nov.">
<emphasis id="B9563D05DAB2E7F5FB30FEB0FA942563" box="[1198,1399,265,291]" italics="true" pageId="186" pageNumber="187">E. schillhammeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAB2E7F5FF23FE97FEFB2507" box="[189,280,301,327]" pageId="186" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C229A94DAB2E7F5FECDFE97FE3F2506" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[339,476,300,326]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="globosus" status="sp. nov.">
<emphasis id="B9563D05DAB2E7F5FECDFE97FE3F2506" box="[339,476,300,326]" italics="true" pageId="186" pageNumber="187">E. globosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAB2E7F5FE7AFE97FDA32507" box="[484,576,301,327]" pageId="186" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
The absence of microgranules on the pronotal disc and elytra and the absence of long erect setae on the labrum, prosternal process, and metaventrite in males separate this species from
<taxonomicName id="4C229A94DAB2E7F5FEC1FECEFE2A25CD" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[351,457,371,397]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="ovalis" status="sp. nov.">
<emphasis id="B9563D05DAB2E7F5FEC1FECEFE2A25CD" box="[351,457,371,397]" italics="true" pageId="186" pageNumber="187">E. ovalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAB2E7F5FE4AFEC9FDD225CD" box="[468,561,371,397]" pageId="186" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C229A94DAB2E7F5FDEEFECEFD1325CD" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[624,752,372,397]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B9563D05DAB2E7F5FDEEFECEFD1325CD" box="[624,752,372,397]" italics="true" pageId="186" pageNumber="187">E. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAB2E7F5FD65FEC9FCBB25CD" box="[763,856,371,397]" pageId="186" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
The surface of the vertex is smooth and not microreticulate as in
<taxonomicName id="4C229A94DAB2E7F5FE28FE2DFE0C25F0" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[438,495,407,432]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="jii" status="sp. nov.">
<emphasis id="B9563D05DAB2E7F5FE28FE2DFE0C25F0" box="[438,495,407,432]" italics="true" pageId="186" pageNumber="187">E. jii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAB2E7F5FE69FE2DFDB025F1" box="[503,595,407,433]" pageId="186" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
The microreticulation of the elytra is formed by irregular polygonal meshes, whereas in
<taxonomicName id="4C229A94DAB2E7F5FE3DFE00FDBE2593" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[419,605,441,467]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="186" pageNumber="187" phylum="Arthropoda" rank="species" species="schoenmanni" status="sp. nov.">
<emphasis id="B9563D05DAB2E7F5FE3DFE00FDBE2593" box="[419,605,441,467]" italics="true" pageId="186" pageNumber="187">E. schoenmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAB2E7F5FDFFFE00FD592594" box="[609,698,442,468]" pageId="186" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
it is formed by transverse lines. The pronotum is moderately convex, with the anterior angles moderately deflexed. The phallobase and parameres are rather long and slender (
<figureCitation id="1319FD92DAB2E7F5FEBCFDBAFE4E265A" box="[290,429,512,538]" captionStart="Fig" captionStartId="185.[189,232,1927,1953]" captionTargetBox="[244,1347,276,1877]" captionTargetId="figure-36@185.[229,1357,265,1897]" captionTargetPageId="185" captionText="Fig. 91. A. Elmomorphus schoenmanni sp. nov., holotype (IAECAS), aedeagus, lateral aspect. B. E. vietnamensis sp. nov., paratype from type locality (CKB), aedeagus, lateral aspect. C. Same, ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773650" httpUri="https://zenodo.org/record/13773650/files/figure.png" pageId="186" pageNumber="187">Fig. 91B–C</figureCitation>
) (PhL/PrL: 2.02 ± 0.13).
</paragraph>
</subSubSection>
<caption id="DF5DB19FDAB0E7F7FF23F814FB712389" ID-DOI="http://doi.org/10.5281/zenodo.13773648" ID-Zenodo-Dep="13773648" box="[189,1170,1966,1993]" httpUri="https://zenodo.org/record/13773648/files/figure.png" pageId="184" pageNumber="185" startId="184.[189,232,1966,1992]" targetBox="[189,1399,265,1928]" targetPageId="184">
<paragraph id="8B9DE117DAB0E7F7FF23F814FB712389" blockId="184.[189,1170,1966,1993]" box="[189,1170,1966,1993]" pageId="184" pageNumber="185">
<emphasis id="B9563D05DAB0E7F7FF23F814FEF92388" bold="true" box="[189,282,1966,1992]" pageId="184" pageNumber="185">Fig. 90.</emphasis>
<taxonomicName id="4C229A94DAB0E7F7FEBEF814FD872388" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[288,612,1966,1992]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="184" pageNumber="185" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="B9563D05DAB0E7F7FEBEF814FD872388" box="[288,612,1966,1992]" italics="true" pageId="184" pageNumber="185">Elmomorphus vietnamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAB0E7F7FDF5F815FD2B2389" box="[619,712,1967,1993]" pageId="184" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male (NMW), TL: 3.27 mm
</paragraph>
</caption>
<caption id="DF5DB19FDAB1E7F6FF23F83DFDA823A8" ID-DOI="http://doi.org/10.5281/zenodo.13773650" ID-Zenodo-Dep="13773650" httpUri="https://zenodo.org/record/13773650/files/figure.png" pageId="185" pageNumber="186" startId="185.[189,232,1927,1953]" targetBox="[244,1347,276,1877]" targetPageId="185">
<paragraph id="8B9DE117DAB1E7F6FF23F83DFDA823A8" blockId="185.[189,1399,1927,2024]" pageId="185" pageNumber="186">
<emphasis id="B9563D05DAB1E7F6FF23F83DFEAB23E1" bold="true" box="[189,328,1927,1953]" pageId="185" pageNumber="186">Fig. 91. A</emphasis>
.
<taxonomicName id="4C229A94DAB1E7F6FEFEF83DFD5323E1" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[352,688,1927,1953]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="schoenmanni" status="sp. nov.">
<emphasis id="B9563D05DAB1E7F6FEFEF83DFD5323E1" box="[352,688,1927,1953]" italics="true" pageId="185" pageNumber="186">Elmomorphus schoenmanni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAB1E7F6FD5FF832FCC423E2" box="[705,807,1928,1954]" pageId="185" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (IAECAS), aedeagus, lateral aspect.
<emphasis id="B9563D05DAB1E7F6FF23F810FF322384" bold="true" box="[189,209,1962,1988]" pageId="185" pageNumber="186">B</emphasis>
.
<taxonomicName id="4C229A94DAB1E7F6FF7DF811FE432384" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[227,416,1963,1988]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="185" pageNumber="186" phylum="Arthropoda" rank="species" species="vietnamensis" status="sp. nov.">
<emphasis id="B9563D05DAB1E7F6FF7DF811FE432384" box="[227,416,1963,1988]" italics="true" pageId="185" pageNumber="186">E. vietnamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDAB1E7F6FE35F811FDEE2385" box="[427,525,1963,1989]" pageId="185" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
, paratype from type locality (CKB), aedeagus, lateral aspect.
<emphasis id="B9563D05DAB1E7F6FA98F810FAF82384" bold="true" box="[1286,1307,1962,1988]" pageId="185" pageNumber="186">C</emphasis>
. Same, ventral aspect. Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C338B29CDAB2E7F5FF23FDF8FB8326CA" box="[189,1120,578,651]" pageId="186" pageNumber="187" type="etymology">
<paragraph id="8B9DE117DAB2E7F5FF23FDF8FEA7261C" blockId="186.[189,324,578,604]" box="[189,324,578,604]" pageId="186" pageNumber="187">
<heading id="D0D5567BDAB2E7F5FF23FDF8FEA7261C" bold="true" box="[189,324,578,604]" fontSize="11" level="3" pageId="186" pageNumber="187" reason="3">
<emphasis id="B9563D05DAB2E7F5FF23FDF8FEA7261C" bold="true" box="[189,324,578,604]" pageId="186" pageNumber="187">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF23FDCAFB8326CA" blockId="186.[189,1120,624,651]" box="[189,1120,624,651]" pageId="186" pageNumber="187">
The epithet is a Latin adjective in the nominative singular referring to
<collectingCountry id="F335A187DAB2E7F5FC6BFDCAFBBF26CA" box="[1013,1116,624,650]" name="Vietnam" pageId="186" pageNumber="187">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDAB2E7F5FF23FD08FD782079" pageId="186" pageNumber="187" type="materials_examined">
<paragraph id="8B9DE117DAB2E7F5FF23FD08FE8F268B" blockId="186.[189,364,689,716]" box="[189,364,689,716]" pageId="186" pageNumber="187">
<heading id="D0D5567BDAB2E7F5FF23FD08FE8F268B" bold="true" box="[189,364,689,716]" fontSize="11" level="3" pageId="186" pageNumber="187" reason="3">
<emphasis id="B9563D05DAB2E7F5FF23FD08FE8F268B" bold="true" box="[189,364,689,716]" pageId="186" pageNumber="187">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF74FD5AFEBF26BA" blockId="186.[189,1399,736,833]" box="[234,348,736,762]" pageId="186" pageNumber="187">
<emphasis id="B9563D05DAB2E7F5FF74FD5AFEBF26BA" bold="true" box="[234,348,736,762]" pageId="186" pageNumber="187">
<typeStatus id="54995FB5DAB2E7F5FF74FD5AFEBF26BA" box="[234,348,736,762]" pageId="186" pageNumber="187" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF23FCBEFE4F2701" blockId="186.[189,1399,736,833]" pageId="186" pageNumber="187">
<materialsCitation id="3B4AEB4ADAB2E7F5FF23FCBEFE4F2701" collectingDate="1996-06-08" collectionCode="NMW" collectorName="Dembicky" country="Vietnam" location="Vinh Phuc Province" pageId="186" pageNumber="187" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc" typeStatus="holotype">
<collectingCountry id="F335A187DAB2E7F5FF23FCBEFEA7275E" box="[189,324,772,798]" name="Vietnam" pageId="186" pageNumber="187">VIETNAM</collectingCountry>
–
<emphasis id="B9563D05DAB2E7F5FEF9FCB9FD86275D" bold="true" box="[359,613,770,797]" pageId="186" pageNumber="187">
<collectingRegion id="49E62FF5DAB2E7F5FEF9FCB9FD86275D" box="[359,613,770,797]" country="Vietnam" name="Vinh Phuc" pageId="186" pageNumber="187">Vinh Phúc Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D242A9EDAB2E7F5FD1AFCBEFD7E275E" box="[644,669,772,798]" count="1" pageId="186" pageNumber="187" type="male">♂</specimenCount>
; “N-VIETNAM: Tam Dao (2) 1.-
<date id="FF9CC7D7DAB2E7F5FBB5FCB9FB7F275D" box="[1067,1180,771,797]" pageId="186" pageNumber="187" value="1996-06-08">
<collectingDate id="EFD83E3FDAB2E7F5FBB5FCB9FB7F275D" box="[1067,1180,771,797]" pageId="186" pageNumber="187" value="1996-06-08">8.6.1996</collectingDate>
</date>
leg.
<collectorName id="26D784C1DAB2E7F5FB45FCB9FAB6275D" box="[1243,1365,771,797]" pageId="186" pageNumber="187">Dembicky</collectorName>
&amp; Pacholatko”;
<collectionCode id="ED3379D2DAB2E7F5FEC2FC9DFE4B2701" box="[348,424,807,833]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="186" pageNumber="187">NMW</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF74FCD2FE8427C2" blockId="186.[189,1399,872,969]" box="[234,359,872,898]" pageId="186" pageNumber="187">
<emphasis id="B9563D05DAB2E7F5FF74FCD2FE8427C2" bold="true" box="[234,359,872,898]" pageId="186" pageNumber="187">
<typeStatus id="54995FB5DAB2E7F5FF74FCD2FE8427C2" box="[234,359,872,898]" pageId="186" pageNumber="187" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF23FC36FEEE2789" blockId="186.[189,1399,872,969]" pageId="186" pageNumber="187">
<materialsCitation id="3B4AEB4ADAB2E7F5FF23FC36FEEA2789" collectingDate="1996-06-08" collectionCode="CKB, NMW" collectorName="Dembicky" country="Vietnam" location="Vinh Phuc Province" pageId="186" pageNumber="187" specimenCount="16" specimenCount-female="4" specimenCount-male="8" stateProvince="Vinh Phuc" typeStatus="paratype">
<collectingCountry id="F335A187DAB2E7F5FF23FC36FEA727E6" box="[189,324,908,934]" name="Vietnam" pageId="186" pageNumber="187">VIETNAM</collectingCountry>
–
<emphasis id="B9563D05DAB2E7F5FEF9FC31FD8027E5" bold="true" box="[359,611,906,933]" pageId="186" pageNumber="187">
<collectingRegion id="49E62FF5DAB2E7F5FEF9FC31FD8027E5" box="[359,611,906,933]" country="Vietnam" name="Vinh Phuc" pageId="186" pageNumber="187">Vinh Phúc Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D242A9EDAB2E7F5FDE0FC36FD2727E6" box="[638,708,908,934]" count="8" pageId="186" pageNumber="187" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="9D242A9EDAB2E7F5FD4CFC36FCF427E6" box="[722,791,908,934]" count="4" pageId="186" pageNumber="187" type="female">4 ♀♀</specimenCount>
,
<specimenCount id="9D242A9EDAB2E7F5FCB8FC36FC8727E6" box="[806,868,908,934]" count="4" pageId="186" pageNumber="187" type="generic">4 ex.</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED3379D2DAB2E7F5FAADFC36FA9127E6" box="[1331,1394,908,934]" pageId="186" pageNumber="187">CKB</collectionCode>
,
<collectionCode id="ED3379D2DAB2E7F5FF23FC15FEEA2789" box="[189,265,943,969]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="186" pageNumber="187">NMW</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF23FC4AFEBE204A" blockId="186.[189,349,1008,1034]" box="[189,349,1008,1034]" pageId="186" pageNumber="187">
<heading id="D0D5567BDAB2E7F5FF23FC4AFEBE204A" bold="true" box="[189,349,1008,1034]" fontSize="11" level="3" pageId="186" pageNumber="187" reason="3">
<emphasis id="B9563D05DAB2E7F5FF23FC4AFEBE204A" bold="true" box="[189,349,1008,1034]" pageId="186" pageNumber="187">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF23FBA5FD782079" blockId="186.[189,667,1055,1081]" box="[189,667,1055,1081]" pageId="186" pageNumber="187">
<collectingCountry id="F335A187DAB2E7F5FF23FBA5FEC02079" box="[189,291,1055,1081]" name="Vietnam" pageId="186" pageNumber="187">Vietnam</collectingCountry>
,
<collectingRegion id="49E62FF5DAB2E7F5FEB0FBA5FDFF2079" box="[302,540,1055,1081]" country="Vietnam" name="Vinh Phuc" pageId="186" pageNumber="187">Vinh Phúc Province</collectingRegion>
, Tam Dao.
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDAB2E7F4FF23FBDAFA942717" lastPageId="187" lastPageNumber="188" pageId="186" pageNumber="187" type="description">
<paragraph id="8B9DE117DAB2E7F5FF23FBDAFEAD203A" blockId="186.[189,334,1120,1146]" box="[189,334,1120,1146]" pageId="186" pageNumber="187">
<heading id="D0D5567BDAB2E7F5FF23FBDAFEAD203A" bold="true" box="[189,334,1120,1146]" fontSize="11" level="3" pageId="186" pageNumber="187" reason="3">
<emphasis id="B9563D05DAB2E7F5FF23FBDAFEAD203A" bold="true" box="[189,334,1120,1146]" pageId="186" pageNumber="187">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF23FB35FC502176" blockId="186.[189,1399,1167,1335]" pageId="186" pageNumber="187">
Measurements (mm): TL:
<specimenCount id="9D242A9EDAB2E7F5FE6CFB35FDFD20E9" box="[498,542,1167,1193]" count="1" pageId="186" pageNumber="187" type="male">♂♂</specimenCount>
3.14–3.34 (3.27± 0.06, n=9),
<specimenCount id="9D242A9EDAB2E7F5FC1BFB35FC5220E9" box="[901,945,1167,1193]" count="1" pageId="186" pageNumber="187" type="female">♀♀</specimenCount>
3.36–3.47 (3.41± 0.05, n=4); PL:
<specimenCount id="9D242A9EDAB2E7F5FAD4FB35FA9520E9" box="[1354,1398,1167,1193]" count="1" pageId="186" pageNumber="187" type="male">♂♂</specimenCount>
0.75–0.85 (0.80 ± 0.03, n= 9),
<specimenCount id="9D242A9EDAB2E7F5FDAAFB09FD83208D" box="[564,608,1203,1229]" count="1" pageId="186" pageNumber="187" type="female">♀♀</specimenCount>
0.80–0.85 (0.82 ±0.02, n=4); PW:
<specimenCount id="9D242A9EDAB2E7F5FBB6FB09FBB7208D" box="[1064,1108,1203,1229]" count="1" pageId="186" pageNumber="187" type="male">♂♂</specimenCount>
1.48–1.61 (1.55± 0.04, n= 9),
<specimenCount id="9D242A9EDAB2E7F5FE8AFB6CFEA320B0" box="[276,320,1238,1264]" count="1" pageId="186" pageNumber="187" type="female">♀♀</specimenCount>
1.55–1.66 (1.59± 0.04, n =4); EL:
<specimenCount id="9D242A9EDAB2E7F5FC95FB6CFCD420B0" box="[779,823,1238,1264]" count="1" pageId="186" pageNumber="187" type="male">♂♂</specimenCount>
2.39–2.52 (2.47± 0.04, n =9),
<specimenCount id="9D242A9EDAB2E7F5FB5AFB6CFB1320B0" box="[1220,1264,1238,1264]" count="1" pageId="186" pageNumber="187" type="female">♀♀</specimenCount>
2.55–2.63 (2.59 ± 0.03, n =4); EW:
<specimenCount id="9D242A9EDAB2E7F5FE73FB43FDFA2153" box="[493,537,1273,1299]" count="1" pageId="186" pageNumber="187" type="male">♂♂</specimenCount>
1.74–1.92 (1.85± 0.05, n= 9),
<specimenCount id="9D242A9EDAB2E7F5FC0BFB43FC222153" box="[917,961,1273,1299]" count="1" pageId="186" pageNumber="187" type="female">♀♀</specimenCount>
1.85–1.95 (1.92± 0.04, n= 4); PhL: 0.72–0.83 (0.78± 0.03, n =9); PrL: 0.35–0.41 (0.39 ±0.02, n =9).
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF23FAE7FB132245" blockId="186.[189,1398,1373,1541]" pageId="186" pageNumber="187">
Body broadly oval, widest before midlength of elytra; dorsum moderately convex (
<figureCitation id="1319FD92DAB2E7F5FB2FFAE7FAEE2138" box="[1201,1293,1373,1400]" captionStart="Fig" captionStartId="184.[189,232,1966,1992]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@184.[189,1399,265,1928]" captionTargetPageId="184" captionText="Fig. 90. Elmomorphus vietnamensis sp. nov., holotype, male (NMW), TL: 3.27 mm" figureDoi="http://doi.org/10.5281/zenodo.13773648" httpUri="https://zenodo.org/record/13773648/files/figure.png" pageId="186" pageNumber="187">Fig. 90</figureCitation>
). Colour dark brown to black, mouthparts and antennae reddish brown; trochanters, tarsi, and distal portions of femora usually dark brown. Pubescence on head, pronotum and elytra consisting of very short and sparse yellowish setae. Plastron on dorsum confined to small areas around antennal insertions; on venter confined to median part of prosternum, prosternal process, and median part of metaventrite.
</paragraph>
<paragraph id="8B9DE117DAB2E7F5FF23F996FE8E22B7" blockId="186.[189,1399,1580,1783]" pageId="186" pageNumber="187">Head surface with weak microreticulation in anterior portion and with minute microgranules in lateral portions around eyes; round punctures smaller than eye facets, separated by 1.0–1.5 × puncture diameter. Plastron confined to small areas around antennal insertions. Anterior margin of clypeus straight. Labrum microreticulate on exposed portion, with minute round setiferous punctures; setae of equal length in both sexes. Eyes slightly protruding from head outline, interfacetal setae short. Antennae 10-segmented, densely setose.</paragraph>
<paragraph id="8B9DE117DAB2E7F4FF23F8A4FB95266A" blockId="186.[189,1399,1822,2025]" lastBlockId="187.[189,1399,265,554]" lastPageId="187" lastPageNumber="188" pageId="186" pageNumber="187">
Pronotum transverse, PW/PL:
<specimenCount id="9D242A9EDAB2E7F5FDACF8A4FDBD2378" box="[562,606,1822,1848]" count="1" pageId="186" pageNumber="187" type="male">♂♂</specimenCount>
1.88–2.00 (1.95± 0.04, n=9),
<specimenCount id="9D242A9EDAB2E7F5FC45F8A4FBE42378" box="[987,1031,1822,1848]" count="1" pageId="186" pageNumber="187" type="female">♀♀</specimenCount>
1.90–2.00 (1.95± 0.04, n= 4). Surface smooth, with round setiferous punctures and rather inapparent microgranules on lateral portions; plastron absent. Anterior rim as wide as two eye facets; anterior angles strongly deflexed, produced anteriorly; lateral pronotal sides weakly rounded. Hypomeron widest behind middle, covered with plastron. Prosternum covered with plastron, except on prosternal process and narrow triangular area before prosternal process. Lateral edges of prosternal process divergent, straight; posterior edge rounded; lateral portions broad, flat, separated from median portion by distinct, deeply impressed narrow ridges; median keel flat; prosternal process without groups of long setae in males. Metaventrite covered with plastron, except for posteriorly widened, convex median part; without groups of long setae in males. Elytra broad, strongly convex, widest before middle, EL/EW:
<specimenCount id="9D242A9EDAB3E7F4FC32FED6FC3B25C6" box="[940,984,364,390]" count="1" pageId="187" pageNumber="188" type="male">♂♂</specimenCount>
1.30–1.39 (1.34 ±0.03, n= 9),
<specimenCount id="9D242A9EDAB3E7F4FAD4FED6FA9525C6" box="[1354,1398,364,390]" count="1" pageId="187" pageNumber="188" type="female">♀♀</specimenCount>
1.32–1.38 (1.35± 0.02, n =4). Surface with distinct microreticulation formed by irregular meshes, and small punctures scattered over entire surface; nine deep longitudinal striae with large round punctures, separated by 0.5–1.0 × puncture diameter; plastron absent. Tibiae slightly curved, PTL/PL:
<specimenCount id="9D242A9EDAB3E7F4FB61FE75FAC825A9" box="[1279,1323,463,489]" count="1" pageId="187" pageNumber="188" type="male">♂♂</specimenCount>
0.94– 1.03 (0.97 ±0.03, n=9),
<specimenCount id="9D242A9EDAB3E7F4FE78FE55FDF12649" box="[486,530,495,521]" count="1" pageId="187" pageNumber="188" type="female">♀♀</specimenCount>
0.88–0.98 (0.92 ± 0.04, n= 4). Protarsus about 0.7× as long as protibia; terminal protarsomeres slightly shorter than three previous tarsomeres combined.
</paragraph>
<paragraph id="8B9DE117DAB3E7F4FF23FDF5FAAC26AC" blockId="187.[189,1399,591,749]" pageId="187" pageNumber="188">
Ventrites covered with plastron, except for middle of ventrite 1 between two admedian keels. Ventrite
<quantity id="4CDA4CF2DAB3E7F4FF23FDD5FF0F26C9" box="[189,236,623,650]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="187" pageNumber="188" unit="in" value="5.0">5 in</quantity>
males with minute, often inapparent triangular excision at apex; in females arcuate at apex with short longitudinal keel. Aedeagus (
<figureCitation id="1319FD92DAB3E7F4FDCCFD2AFD0326EB" box="[594,736,656,683]" captionStart="Fig" captionStartId="185.[189,232,1927,1953]" captionTargetBox="[244,1347,276,1877]" captionTargetId="figure-36@185.[229,1357,265,1897]" captionTargetPageId="185" captionText="Fig. 91. A. Elmomorphus schoenmanni sp. nov., holotype (IAECAS), aedeagus, lateral aspect. B. E. vietnamensis sp. nov., paratype from type locality (CKB), aedeagus, lateral aspect. C. Same, ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773650" httpUri="https://zenodo.org/record/13773650/files/figure.png" pageId="187" pageNumber="188">Fig. 91B–C</figureCitation>
); phallobase rather long, proximally expanded, PhL/PrL: 1.76–2.29 (2.02± 0.13, n =9); parameres long, slightly curved ventrad in distal third, apices pointed in lateral view; penis slender; sclerotised fibula slender. Bursa copulatrix without spiny microsclerites.
</paragraph>
<paragraph id="8B9DE117DAB3E7F4FF23FCABFDD4276B" blockId="187.[189,567,785,811]" box="[189,567,785,811]" pageId="187" pageNumber="188">
<heading id="D0D5567BDAB3E7F4FF23FCABFDD4276B" bold="true" box="[189,567,785,811]" fontSize="11" level="3" pageId="187" pageNumber="188" reason="3">
<emphasis id="B9563D05DAB3E7F4FF23FCABFDD4276B" bold="true" box="[189,567,785,811]" pageId="187" pageNumber="188">Secondary sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAB3E7F4FF23FC87FA942717" blockId="187.[189,1399,829,856]" box="[189,1399,829,856]" pageId="187" pageNumber="188">Male ventrite 5 with minute triangular excision at apex, in female rounded with short longitudinal keel.</paragraph>
</subSubSection>
<subSubSection id="C338B29CDAB3E7F4FF23FCC6FE572783" box="[189,436,892,963]" pageId="187" pageNumber="188" type="distribution">
<paragraph id="8B9DE117DAB3E7F4FF23FCC6FEB527D6" blockId="187.[189,342,892,918]" box="[189,342,892,918]" pageId="187" pageNumber="188">
<heading id="D0D5567BDAB3E7F4FF23FCC6FEB527D6" bold="true" box="[189,342,892,918]" fontSize="11" level="3" pageId="187" pageNumber="188" reason="3">
<emphasis id="B9563D05DAB3E7F4FF23FCC6FEB527D6" bold="true" box="[189,342,892,918]" pageId="187" pageNumber="188">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DAB3E7F4FF23FC13FE572783" blockId="187.[189,436,937,963]" box="[189,436,937,963]" pageId="187" pageNumber="188">
<collectingCountry id="F335A187DAB3E7F4FF23FC13FEC32783" box="[189,288,937,963]" name="Vietnam" pageId="187" pageNumber="188">Vietnam</collectingCountry>
(
<figureCitation id="1319FD92DAB3E7F4FEAEFC13FE442783" box="[304,423,937,963]" captionStart="Fig" captionStartId="222.[189,232,1867,1893]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-44@222.[189,1399,265,1830]" captionTargetPageId="222" captionText="Fig. 114. Distribution maps. A. Elmomorphus globosus sp. nov. B. E. ovalis Kodada, Selnekovič &amp; Jäch sp. nov. C. E. minutus sp. nov. D. E. jii sp. nov. E. E. schoenmanni sp. nov. F. E. vietnamensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773722" httpUri="https://zenodo.org/record/13773722/files/figure.png" pageId="187" pageNumber="188">Fig. 114F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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