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<document id="D5D1899B69DAB37683587C8F4816CA5D" ID-DOI="10.5852/ejt.2024.919.2407" ID-ISSN="2118-9773" ID-Zenodo-Dep="10559520" ID-ZooBank="72B7907F-F901-4DDD-B2DD-63D03253837E" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1706085863884" checkinUser="plazi" docAuthor="Constant, Jérôme &amp; Pham, Hong Thai" docDate="2024" docId="03DE87B0FFEFFFACFDACFE60A5FDFA17" docLanguage="en" docName="EJT.2024.919.1-87.pdf" docOrigin="European Journal of Taxonomy 919 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2407/10617" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maculergithus Constant &amp; Pham 2016" docType="treatment" docVersion="5" lastPageNumber="25" masterDocId="FFE7FFC8FFF7FFB4FFABFFD4A706FFDA" masterDocTitle="Sixteen issid planthopper species in one day in Dong Son-Ky Thuong Nature Reserve in North Vietnam: Eight new species, one new genus and additional new records (Hemiptera: Fulgoromorpha: Issidae)" masterLastPageNumber="87" masterPageNumber="1" pageNumber="25" updateTime="1706193695395" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7C06B01B99392739F103D8F31AC4A308">Sixteen issid planthopper species in one day in Dong Son-Ky Thuong Nature Reserve in North Vietnam: Eight new species, one new genus and additional new records (Hemiptera: Fulgoromorpha: Issidae)</mods:title>
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<mods:namePart id="CC3B316D2AFEC6133F65CEE62A2DBA75">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="D95B38ACBC7B9F2B089B2613ADAA480D">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
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<mods:namePart id="40DAB8FE2DD05C856C41D105E6EA16A0">Pham, Hong Thai</mods:namePart>
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<mods:affiliation id="1B898AFE755CA897076CC29A8388F58B">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Street, Hanoi, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam.</mods:affiliation>
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<mods:date id="34B6B30303325E7EFE04C45892E87949">2024</mods:date>
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<heading id="D08081CAFFEFFFACFDACFE60A32BFE14" box="[519,1069,435,462]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
Genus
<taxonomicName id="4C774D25FFEFFFACFDF2FE67A32BFE14" ID-CoL="5JTW" authority="Constant &amp; Pham, 2016" authorityName="Constant &amp; Pham" authorityYear="2016" box="[601,1069,435,462]" class="Insecta" family="Issidae" genus="Maculergithus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAB4FFEFFFACFDF2FE67A40DFE14" bold="true" box="[601,779,435,462]" italics="true" pageId="24" pageNumber="25">Maculergithus</emphasis>
<bibRefCitation id="EFE64B57FFEFFFACFCB9FE60A32BFE14" author="Constant J. &amp; Pham H. T." box="[786,1069,436,462]" pageId="24" pageNumber="25" pagination="1 - 16" refId="ref33300" refString="Constant J. &amp; Pham H. T. 2016. Maculergithus, a new subgenus in Gergithus Schumacher, 1915 with two new species from northern Vietnam (Hemiptera: Fulgoromorpha: Issidae). European Journal of Taxonomy 198: 1 - 16. https: // doi. org / 10.5852 / ejt. 2016.198" type="journal article" year="2016">Constant &amp; Pham, 2016</bibRefCitation>
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<subSubSection id="C36D652DFFEFFFACFF16FE21A431FDCA" box="[189,823,501,528]" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="8BC836A6FFEFFFACFF16FE21A431FDCA" blockId="24.[189,1398,501,563]" box="[189,823,501,528]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="AB671188FFEFFFACFF16FE21A431FDCA" box="[189,823,501,528]" pageId="24" pageNumber="25">
<treatmentCitation id="0AD610B7FFEFFFACFF16FE21A434FDCA" author="Constant J. &amp; Pham H. T." box="[189,818,501,528]" page="3" pageId="24" pageNumber="25" year="2016">
<taxonomicName id="4C774D25FFEFFFACFF16FE21A434FDCA" authority="Constant &amp; Pham, 2016: 3" authorityName="Constant &amp; Pham" authorityPageNumber="3" authorityYear="2016" box="[189,818,501,528]" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subGenus" subGenus="Maculergithus">
<emphasis id="B903EAB4FFEFFFACFF16FE21A637FDD5" box="[189,305,501,527]" italics="true" pageId="24" pageNumber="25">Gergithus</emphasis>
(
<emphasis id="B903EAB4FFEFFFACFEEAFE21A6EDFDD5" box="[321,491,501,527]" italics="true" pageId="24" pageNumber="25">Maculergithus</emphasis>
)
<bibRefCitation id="EFE64B57FFEFFFACFE50FE22A434FDCA" author="Constant J. &amp; Pham H. T." box="[507,818,501,528]" pageId="24" pageNumber="25" pagination="1 - 16" refId="ref33300" refString="Constant J. &amp; Pham H. T. 2016. Maculergithus, a new subgenus in Gergithus Schumacher, 1915 with two new species from northern Vietnam (Hemiptera: Fulgoromorpha: Issidae). European Journal of Taxonomy 198: 1 - 16. https: // doi. org / 10.5852 / ejt. 2016.198" type="journal article" year="2016">Constant &amp; Pham, 2016: 3</bibRefCitation>
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.
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<typeStatus id="54CC8804FFEFFFACFC97FE22A470FDCA" box="[828,886,502,528]" pageId="24" pageNumber="25">Type</typeStatus>
species:
<taxonomicName id="4C774D25FFEFFFACFC74FE21A503FDE8" ID-CoL="3FWF4" authority="Constant &amp; Pham, 2016" authorityName="Constant &amp; Pham" authorityYear="2016" class="Insecta" family="Issidae" genus="Gergithus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tamdao" subGenus="Maculergithus">
<emphasis id="B903EAB4FFEFFFACFC74FE21A355FDD5" box="[991,1107,501,527]" italics="true" pageId="24" pageNumber="25">Gergithus</emphasis>
(
<emphasis id="B903EAB4FFEFFFACFBCFFE21A208FDD5" box="[1124,1294,501,527]" italics="true" pageId="24" pageNumber="25">Maculergithus</emphasis>
)
<emphasis id="B903EAB4FFEFFFACFAB5FE21A270FDD5" box="[1310,1398,501,527]" italics="true" pageId="24" pageNumber="25">tamdao</emphasis>
<bibRefCitation id="EFE64B57FFEFFFACFF41FDCDA503FDE8" author="Constant J. &amp; Pham H. T." box="[234,517,536,563]" pageId="24" pageNumber="25" pagination="1 - 16" refId="ref33300" refString="Constant J. &amp; Pham H. T. 2016. Maculergithus, a new subgenus in Gergithus Schumacher, 1915 with two new species from northern Vietnam (Hemiptera: Fulgoromorpha: Issidae). European Journal of Taxonomy 198: 1 - 16. https: // doi. org / 10.5852 / ejt. 2016.198" type="journal article" year="2016">Constant &amp; Pham, 2016</bibRefCitation>
</taxonomicName>
by original designation.
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<paragraph id="8BC836A6FFEFFFACFF16FD8EA4AAFDAE" blockId="24.[189,940,602,629]" box="[189,940,602,629]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="AB671188FFEFFFACFF16FD8EA4AAFDAE" box="[189,940,602,629]" pageId="24" pageNumber="25">
<taxonomicName id="4C774D25FFEFFFACFF16FD8EA661FDAE" ID-CoL="5JTW" authorityName="Constant &amp; Pham" authorityYear="2016" box="[189,359,602,628]" class="Insecta" family="Issidae" genus="Maculergithus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAB4FFEFFFACFF16FD8EA661FDAE" box="[189,359,602,628]" italics="true" pageId="24" pageNumber="25">Maculergithus</emphasis>
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<treatmentCitation id="0AD610B7FFEFFFACFE2FFD8EA59CFDAE" author="Gnezdilov V. M." box="[388,666,602,628]" page="1344" pageId="24" pageNumber="25" year="2017">
<bibRefCitation id="EFE64B57FFEFFFACFE2FFD8EA59CFDAE" author="Gnezdilov V. M." box="[388,666,602,628]" pageId="24" pageNumber="25" pagination="1338 - 1352" refId="ref34143" refString="Gnezdilov V. M. 2017 b. Addenda to the revisions of the genera Gergithus Stal and Hemisphaerius Schaum (Hemiptera, Auchenorrhyncha, Fulgoroidea: Issidae). Entomological Review 97 (9): 1338 - 1352. https: // doi. org / 10.1134 / S 0013873817090123" type="journal article" year="2017">Gnezdilov 2017b: 1344</bibRefCitation>
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(raised to genus level).
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<heading id="D08081CAFFEFFFACFF16FD4FA630FD6F" bold="true" box="[189,310,667,693]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B903EAB4FFEFFFACFF16FD4FA630FD6F" bold="true" box="[189,310,667,693]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FD1EA244FD3E" blockId="24.[189,1399,714,1269]" box="[189,1346,714,741]" pageId="24" pageNumber="25">
The characters defining the genus were given by
<bibRefCitation id="EFE64B57FFEFFFACFD57FD1EA324FD3E" author="Constant J. &amp; Pham H. T." box="[764,1058,714,740]" pageId="24" pageNumber="25" pagination="1 - 16" refId="ref33300" refString="Constant J. &amp; Pham H. T. 2016. Maculergithus, a new subgenus in Gergithus Schumacher, 1915 with two new species from northern Vietnam (Hemiptera: Fulgoromorpha: Issidae). European Journal of Taxonomy 198: 1 - 16. https: // doi. org / 10.5852 / ejt. 2016.198" type="journal article" year="2016">Constant &amp; Pham (2016)</bibRefCitation>
and
<bibRefCitation id="EFE64B57FFEFFFACFBF0FD1EA239FD3E" author="Gnezdilov V. M." box="[1115,1343,714,740]" pageId="24" pageNumber="25" pagination="1338 - 1352" refId="ref34143" refString="Gnezdilov V. M. 2017 b. Addenda to the revisions of the genera Gergithus Stal and Hemisphaerius Schaum (Hemiptera, Auchenorrhyncha, Fulgoroidea: Issidae). Entomological Review 97 (9): 1338 - 1352. https: // doi. org / 10.1134 / S 0013873817090123" type="journal article" year="2017">Gnezdilov (2017b)</bibRefCitation>
:
</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FD39A56BFCDD" blockId="24.[189,1399,714,1269]" box="[189,621,749,775]" pageId="24" pageNumber="25">(1) the black-brown ground colour;</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FCC5A5DDFC94" blockId="24.[189,1399,714,1269]" pageId="24" pageNumber="25">(2) the tegmina without humeral hump or projection of the costal margin under eye, and with numerous yellow spots separated from one another;</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FC83A64CFC4F" blockId="24.[189,1399,714,1269]" pageId="24" pageNumber="25">(3) the frons wide, widened above clypeus, with a broad yellow transverse band above frontoclypeal suture;</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FC49A530FC6D" blockId="24.[189,1399,714,1269]" box="[189,566,925,951]" pageId="24" pageNumber="25">(4) the clypeus without carina;</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FC14A461FC00" blockId="24.[189,1399,714,1269]" box="[189,871,960,987]" pageId="24" pageNumber="25">(5) the vertex at least 4 times as wide as long in midline;</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FC30A489FC24" blockId="24.[189,1399,714,1269]" box="[189,911,996,1022]" pageId="24" pageNumber="25">(6) the mesonotum with a yellow spot on each lateral angle;</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FBD3A5D4FBFB" blockId="24.[189,1399,714,1269]" box="[189,722,1031,1057]" pageId="24" pageNumber="25">(7) the hind wing well developed, unilobed;</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FBFEA6EAFBB2" blockId="24.[189,1399,714,1269]" pageId="24" pageNumber="25">(8) the posterior margin of the pygofer of the male genitalia strongly rounded posterad in lateral view and without process;</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FBA5A6C7FB74" blockId="24.[189,1399,714,1269]" pageId="24" pageNumber="25">(9) the anal tube of the male elongate, wide, truncate apically and with apicolateral angles projecting posteroventrally;</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FB63A69FFB2F" blockId="24.[189,1399,714,1269]" pageId="24" pageNumber="25">(10) the asymmetrical aedeagus with ventral pair of short hooks and left lateral lobe of periandrium with large process.</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFEFFFACFF16FACFA5FDFA17" pageId="24" pageNumber="25" type="taxon_list">
<paragraph id="8BC836A6FFEFFFACFF16FACFA68AFAEF" blockId="24.[189,396,1307,1333]" box="[189,396,1307,1333]" pageId="24" pageNumber="25">
<heading id="D08081CAFFEFFFACFF16FACFA68AFAEF" bold="true" box="[189,396,1307,1333]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B903EAB4FFEFFFACFF16FACFA68AFAEF" bold="true" box="[189,396,1307,1333]" pageId="24" pageNumber="25">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FA9EA450FABE" blockId="24.[189,923,1354,1486]" box="[189,854,1354,1380]" pageId="24" pageNumber="25">
<taxonomicName id="4C774D25FFEFFFACFF16FA9EA450FABE" authority="(Constant &amp; Pham, 2016)" baseAuthorityName="Constant &amp; Pham" baseAuthorityYear="2016" box="[189,854,1354,1380]" class="Insecta" family="Issidae" genus="Maculergithus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="luteomaculatus">
<emphasis id="B903EAB4FFEFFFACFF16FA9EA527FABE" box="[189,545,1354,1380]" italics="true" pageId="24" pageNumber="25">Maculergithus luteomaculatus</emphasis>
(
<bibRefCitation id="EFE64B57FFEFFFACFD9AFA9EA448FABE" author="Constant J. &amp; Pham H. T." box="[561,846,1354,1380]" pageId="24" pageNumber="25" pagination="1 - 16" refId="ref33300" refString="Constant J. &amp; Pham H. T. 2016. Maculergithus, a new subgenus in Gergithus Schumacher, 1915 with two new species from northern Vietnam (Hemiptera: Fulgoromorpha: Issidae). European Journal of Taxonomy 198: 1 - 16. https: // doi. org / 10.5852 / ejt. 2016.198" type="journal article" year="2016">Constant &amp; Pham, 2016</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FAB9A474FA5D" blockId="24.[189,923,1354,1486]" box="[189,882,1389,1416]" pageId="24" pageNumber="25">
<taxonomicName id="4C774D25FFEFFFACFF16FAB9A474FA5D" authority="(Che, Zhang &amp; Wang, 2007)" baseAuthorityName="Che, Zhang &amp; Wang" baseAuthorityYear="2007" box="[189,882,1389,1416]" class="Insecta" family="Issidae" genus="Maculergithus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="multipunctatus">
<emphasis id="B903EAB4FFEFFFACFF16FAB9A51AFA5D" box="[189,540,1389,1415]" italics="true" pageId="24" pageNumber="25">Maculergithus multipunctatus</emphasis>
(
<bibRefCitation id="EFE64B57FFEFFFACFD87FAB9A46CFA5D" author="Che Y. &amp; Zhang Y. - L. &amp; Wang Y. - L." box="[556,874,1389,1416]" pageId="24" pageNumber="25" pagination="611 - 627" refId="ref32906" refString="Che Y. - L, Zhang Y. - L. &amp; Wang Y. - L. 2007. Seven new species and one new record of Gergithus Stal (Hemiptera: Fulgoroidea: Issidae) from China. Proceedings of the Entomological Society of Washington 109 (3): 611 - 627." type="journal article" year="2007">Che, Zhang &amp; Wang, 2007</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FA44A49DFA71" blockId="24.[189,923,1354,1486]" box="[189,923,1424,1451]" pageId="24" pageNumber="25">
<taxonomicName id="4C774D25FFEFFFACFF16FA44A49DFA71" authority="(Che, Zhang &amp; Wang, 2007)" baseAuthorityName="Che, Zhang &amp; Wang" baseAuthorityYear="2007" box="[189,923,1424,1451]" class="Insecta" family="Issidae" genus="Maculergithus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="quinquemaculatus">
<emphasis id="B903EAB4FFEFFFACFF16FA44A543FA70" box="[189,581,1424,1450]" italics="true" pageId="24" pageNumber="25">Maculergithus quinquemaculatus</emphasis>
(
<bibRefCitation id="EFE64B57FFEFFFACFDFEFA44A495FA71" author="Che Y. &amp; Zhang Y. - L. &amp; Wang Y. - L." box="[597,915,1424,1451]" pageId="24" pageNumber="25" pagination="611 - 627" refId="ref32906" refString="Che Y. - L, Zhang Y. - L. &amp; Wang Y. - L. 2007. Seven new species and one new record of Gergithus Stal (Hemiptera: Fulgoroidea: Issidae) from China. Proceedings of the Entomological Society of Washington 109 (3): 611 - 627." type="journal article" year="2007">Che, Zhang &amp; Wang, 2007</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BC836A6FFEFFFACFF16FA60A5FDFA17" blockId="24.[189,923,1354,1486]" box="[189,763,1459,1486]" pageId="24" pageNumber="25">
<taxonomicName id="4C774D25FFEFFFACFF16FA60A5FDFA17" authority="(Constant &amp; Pham, 2016)" baseAuthorityName="Constant &amp; Pham" baseAuthorityYear="2016" box="[189,763,1459,1486]" class="Insecta" family="Issidae" genus="Maculergithus" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="tamdao">
<emphasis id="B903EAB4FFEFFFACFF16FA60A6C0FA14" box="[189,454,1460,1486]" italics="true" pageId="24" pageNumber="25">Maculergithus tamdao</emphasis>
(
<bibRefCitation id="EFE64B57FFEFFFACFE7EFA60A5F5FA17" author="Constant J. &amp; Pham H. T." box="[469,755,1459,1486]" pageId="24" pageNumber="25" pagination="1 - 16" refId="ref33300" refString="Constant J. &amp; Pham H. T. 2016. Maculergithus, a new subgenus in Gergithus Schumacher, 1915 with two new species from northern Vietnam (Hemiptera: Fulgoromorpha: Issidae). European Journal of Taxonomy 198: 1 - 16. https: // doi. org / 10.5852 / ejt. 2016.198" type="journal article" year="2016">Constant &amp; Pham, 2016</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>