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<document id="007287FC69D39149590875C76CD9B595" ID-DOI="10.11646/zootaxa.3722.4.1" ID-GBIF-Dataset="c092e547-503a-41ef-a855-4a12b5d776c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="223434" ID-ZooBank="3A461275-7565-4F6E-9785-12C45E12992E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460692161620" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2013" docId="03DE87B0FFC3FFC2CADAB59FFA1B18F7" docLanguage="en" docName="zt03722p438.pdf" docOrigin="Zootaxa 3722 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrphonothrus seniczaki Colloff, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="410" masterDocId="FFE7FFC8FFC4FFCBCA4DB328FF871A47" masterDocTitle="Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region" masterLastPageNumber="438" masterPageNumber="401" pageNumber="408" updateTime="1698351897157" updateUser="plazi">
<mods:mods id="8818FE9BED77D053480684DAAD048127" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BCEEDF9BD886B969E7F56FC2C7E75C52">
<mods:title id="F9CE716A114BCA68A9BE119468B15383">Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region</mods:title>
</mods:titleInfo>
<mods:name id="796FC468B12CC739633E8511D13D06D1" type="personal">
<mods:role id="4776EAED68F33DDDA83EDB8F6B0C2C52">
<mods:roleTerm id="FB897B68DC5B25C858DE6295FAB04A6F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4DEBF64DB8803EFCA37FAC0198D6C5E7">Colloff, Matthew J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3C725FDC620B15D493A266E001796378">text</mods:typeOfResource>
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<mods:title id="B338E24917D441F985587B26AB4A55A7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B50F30B112F2233807FD7EC0A80B9167">
<mods:date id="9B14A0445AE624575EA7174BD75DC476">2013</mods:date>
<mods:detail id="6A1F78405A75BC40D68D4EA475F1B9B6" type="volume">
<mods:number id="959394BB6DA0038B4E1D288FB018B532">3722</mods:number>
</mods:detail>
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<mods:number id="9042F1838CC5A8548C596076895B67CB">4</mods:number>
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<mods:classification id="334A27775CE238FD30088A4A98F389BD">journal article</mods:classification>
<mods:identifier id="EADC79CF313883F6DFF837C4683A56F6" type="DOI">10.11646/zootaxa.3722.4.1</mods:identifier>
<mods:identifier id="45E77F1950306D18CD03CCB1A688991E" type="GBIF-Dataset">c092e547-503a-41ef-a855-4a12b5d776c4</mods:identifier>
<mods:identifier id="B0ED08028BD86685E214AB5C877559B9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D7101340CB08150CD652665AE7543C71" type="Zenodo-Dep">223434</mods:identifier>
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<treatment id="03DE87B0FFC3FFC2CADAB59FFA1B18F7" ID-DOI="http://doi.org/10.5281/zenodo.6157848" ID-GBIF-Taxon="119580501" ID-Zenodo-Dep="6157848" LSID="urn:lsid:plazi:treatment:03DE87B0FFC3FFC2CADAB59FFA1B18F7" httpUri="http://treatment.plazi.org/id/03DE87B0FFC3FFC2CADAB59FFA1B18F7" lastPageId="9" lastPageNumber="410" pageId="7" pageNumber="408">
<subSubSection id="C36D652DFFC3FFCCCADAB59FFA131D7C" pageId="7" pageNumber="408" type="nomenclature">
<paragraph id="8BC836A6FFC3FFCCCADAB59FFDA01C96" blockId="7.[151,551,1719,1779]" box="[151,551,1719,1745]" pageId="7" pageNumber="408">
<heading id="D08081CAFFC3FFCCCADAB59FFDA01C96" bold="true" box="[151,551,1719,1745]" fontSize="11" level="1" pageId="7" pageNumber="408" reason="1">
<emphasis id="B903EAB4FFC3FFCCCADAB59FFDA01C96" bold="true" box="[151,551,1719,1745]" pageId="7" pageNumber="408">
<taxonomicName id="4C774D25FFC3FFCCCADAB59FFE461C96" box="[151,449,1719,1745]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="seniczaki" status="sp. nov.">
<emphasis id="B903EAB4FFC3FFCCCADAB59FFE461C96" bold="true" box="[151,449,1719,1745]" italics="true" pageId="7" pageNumber="408">Tyrphonothrus seniczaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFC3FFCCCB84B59FFDA01C96" box="[457,551,1719,1745]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFC3FFCCCADAB5F2FF6C1CB4" blockId="7.[151,551,1719,1779]" box="[151,235,1754,1779]" pageId="7" pageNumber="408">
(
<figureCitation id="134C2A23FFC3FFCCCAD2B5F2FF651CB4" box="[159,226,1754,1779]" captionStart="FIGURE 5" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[162,1420,819,1883]" captionTargetId="figure@8.[150,1436,788,1896]" captionTargetPageId="8" captionText="FIGURE 5. Tyrphonothrus seniczaki sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223439/files/figure.png" pageId="7" pageNumber="408">Fig. 5</figureCitation>
)
</paragraph>
<paragraph id="8BC836A6FFC3FFCCCADAB40AFA131D7C" blockId="7.[151,1436,1826,2031]" box="[151,1428,1826,1851]" pageId="7" pageNumber="408">
<emphasis id="B903EAB4FFC3FFCCCADAB40AFEA51D7C" bold="true" box="[151,290,1826,1851]" pageId="7" pageNumber="408">Dimensions</emphasis>
.
<typeStatus id="54CC8804FFC3FFCCCB62B40AFE101D7C" box="[303,407,1826,1851]" pageId="7" pageNumber="408" type="holotype">Holotype</typeStatus>
female length 508 Μm, breadth 278 Μm. Ratio of prodorsum to total length: 0.3 (
<typeStatus id="54CC8804FFC3FFCCCF6DB40AFA0E1D7C" box="[1312,1417,1826,1851]" pageId="7" pageNumber="408" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFC3FFC3CA8AB46EFDC618A7" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408" type="description">
<paragraph id="8BC836A6FFC3FFCCCA8AB46EFB5C1D8D" blockId="7.[151,1436,1826,2031]" pageId="7" pageNumber="408">
<emphasis id="B903EAB4FFC3FFCCCA8AB46EFE991D18" bold="true" box="[199,286,1862,1887]" pageId="7" pageNumber="408">Female</emphasis>
.
<emphasis id="B903EAB4FFC3FFCCCB63B46EFE2C1D18" box="[302,427,1862,1887]" italics="true" pageId="7" pageNumber="408">Prodorsum</emphasis>
. Rostrum rounded. Rostral seta (
<emphasis id="B903EAB4FFC3FFCCC965B46FFCB81D19" box="[808,831,1863,1886]" italics="true" pageId="7" pageNumber="408">ro</emphasis>
) smooth, curved, setiform, thin, 26 Μm long, shorter than mutual distance, emerging from alveolus (
<figureCitation id="134C2A23FFC3FFCCC8E5B442FD6C1DC4" box="[680,747,1898,1923]" captionStart="FIGURE 5" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[162,1420,819,1883]" captionTargetId="figure@8.[150,1436,788,1896]" captionTargetPageId="8" captionText="FIGURE 5. Tyrphonothrus seniczaki sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223439/files/figure.png" pageId="7" pageNumber="408">Fig. 5</figureCitation>
a). Lamellar seta (
<emphasis id="B903EAB4FFC3FFCCC9FBB442FC4D1DC4" box="[950,970,1898,1923]" italics="true" pageId="7" pageNumber="408">le</emphasis>
) smooth, recurved, 20 Μm long, less than half mutual distance, median of apex of broad, incurved, S-shaped prodorsal ridge. Interlamellar seta (
<emphasis id="B903EAB4FFC3FFCCCF51B4A7FAB51DE1" box="[1308,1330,1935,1958]" italics="true" pageId="7" pageNumber="408">in</emphasis>
) smooth, setiform, thin, 108 Μm long, 10 × longer than smooth, minute seta
<emphasis id="B903EAB4FFC3FFCCC932B49BFC1F1D8D" box="[895,920,1971,1994]" italics="true" pageId="7" pageNumber="408">ex</emphasis>
. Prodorsum smooth, porose.
</paragraph>
<paragraph id="8BC836A6FFC3FFC3CA8AB4FFFDBF1BDE" blockId="7.[151,1436,1826,2031]" lastBlockId="8.[151,1436,151,736]" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408">
<emphasis id="B903EAB4FFC3FFCCCA8AB4FFFEC51DA9" box="[199,322,2007,2030]" italics="true" pageId="7" pageNumber="408">Notogaster</emphasis>
. Anterior notogastral margin straight. Humeral region rounded (
<figureCitation id="134C2A23FFC3FFCCCE57B4FEFBD81DA8" box="[1050,1119,2006,2031]" captionStart="FIGURE 5" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[162,1420,819,1883]" captionTargetId="figure@8.[150,1436,788,1896]" captionTargetPageId="8" captionText="FIGURE 5. Tyrphonothrus seniczaki sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223439/files/figure.png" pageId="7" pageNumber="408">Fig. 5</figureCitation>
a). With 15 pairs of smooth, very thin setae. Setae
<emphasis id="B903EAB4FFCCFFC3CBC3B3B1FE1D1AF7" box="[398,410,153,176]" italics="true" pageId="8" pageNumber="409">e</emphasis>
2,
<emphasis id="B903EAB4FFCCFFC3CBFCB3BFFE381AF7" box="[433,447,151,176]" italics="true" pageId="8" pageNumber="409">h</emphasis>
1 and
<emphasis id="B903EAB4FFCCFFC3CBB2B3BFFD8A1AF7" box="[511,525,151,176]" italics="true" pageId="8" pageNumber="409">h</emphasis>
2 flagelliform apically, much longer than others (58–72 Μm). Setae of the
<emphasis id="B903EAB4FFCCFFC3CF07B3B1FAD11AF7" box="[1354,1366,153,176]" italics="true" pageId="8" pageNumber="409">c</emphasis>
and
<emphasis id="B903EAB4FFCCFFC3CFC3B3BFFA1B1AF7" box="[1422,1436,151,176]" italics="true" pageId="8" pageNumber="409">d</emphasis>
series very short (7–15 Μm);
<emphasis id="B903EAB4FFCCFFC3CBBAB3E8FD841A90" box="[503,515,192,215]" italics="true" pageId="8" pageNumber="409">c</emphasis>
1 positioned on squat tubercle on anterior margin of notogaster,
<emphasis id="B903EAB4FFCCFFC3CF48B396FA941A90" box="[1285,1299,190,215]" italics="true" pageId="8" pageNumber="409">d</emphasis>
1 positioned markedly anteriorly, almost level transversely with bases of
<emphasis id="B903EAB4FFCCFFC3C913B3CEFCED1ABA" box="[862,874,230,253]" italics="true" pageId="8" pageNumber="409">c</emphasis>
3; distance between
<emphasis id="B903EAB4FFCCFFC3CE1AB3CEFBE41ABA" box="[1111,1123,230,253]" italics="true" pageId="8" pageNumber="409">c</emphasis>
1 and
<emphasis id="B903EAB4FFCCFFC3CEE6B3CEFB301ABA" box="[1195,1207,230,253]" italics="true" pageId="8" pageNumber="409">c</emphasis>
2 at least twice that between
<emphasis id="B903EAB4FFCCFFC3CAB3B225FE8D1B63" box="[254,266,269,292]" italics="true" pageId="8" pageNumber="409">c</emphasis>
2 and
<emphasis id="B903EAB4FFCCFFC3CB02B225FEDC1B63" box="[335,347,269,292]" italics="true" pageId="8" pageNumber="409">c</emphasis>
3. Seta
<emphasis id="B903EAB4FFCCFFC3CBE3B225FE3D1B63" box="[430,442,269,292]" italics="true" pageId="8" pageNumber="409">e</emphasis>
1 minute, 4 Μm long;
<emphasis id="B903EAB4FFCCFFC3C8FBB225FD391B62" box="[694,702,269,293]" italics="true" pageId="8" pageNumber="409">f</emphasis>
2,
<emphasis id="B903EAB4FFCCFFC3C89BB224FD631B62" box="[726,740,268,293]" italics="true" pageId="8" pageNumber="409">h</emphasis>
3 and
<emphasis id="B903EAB4FFCCFFC3C969B225FCB51B63" box="[804,818,269,292]" italics="true" pageId="8" pageNumber="409">p</emphasis>
1 setiform, subequal (20–26 Μm long). Short, acutelydiagonal ridges between
<emphasis id="B903EAB4FFCCFFC3CBF6B21BFE4E1B0B" box="[443,457,307,332]" italics="true" pageId="8" pageNumber="409">h</emphasis>
2 and
<emphasis id="B903EAB4FFCCFFC3C842B21BFD9A1B0B" box="[527,541,307,332]" italics="true" pageId="8" pageNumber="409">h</emphasis>
1; otherwise ridges absent. Notogastral margins slightly narrowing anteriorly, broadest between
<emphasis id="B903EAB4FFCCFFC3CB25B273FEF31B35" box="[360,372,347,370]" italics="true" pageId="8" pageNumber="409">e</emphasis>
2 and
<emphasis id="B903EAB4FFCCFFC3CBF4B272FE461B35" box="[441,449,346,370]" italics="true" pageId="8" pageNumber="409">f</emphasis>
2. Setae
<emphasis id="B903EAB4FFCCFFC3C86CB273FDA81B35" box="[545,559,347,370]" italics="true" pageId="8" pageNumber="409">p</emphasis>
3 markedly anterior of
<emphasis id="B903EAB4FFCCFFC3C975B273FCC11B35" box="[824,838,347,370]" italics="true" pageId="8" pageNumber="409">p</emphasis>
2. Ratio of length to breadth 1.2. Caudal margin Ushaped, rounded. Notogaster smooth.
</paragraph>
<paragraph id="8BC836A6FFCCFFC3CA8AB28DFB931842" blockId="8.[151,1436,151,736]" pageId="8" pageNumber="409">
<emphasis id="B903EAB4FFCCFFC3CA8AB28DFED11BFB" box="[199,342,421,444]" italics="true" pageId="8" pageNumber="409">Coxisternum</emphasis>
. Epimeral setal formula 3-1-2-3. Epimeral setae smooth, setiform, minute;
<emphasis id="B903EAB4FFCCFFC3CEFEB28DFB481BFA" box="[1203,1231,421,445]" italics="true" pageId="8" pageNumber="409">3a</emphasis>
,
<emphasis id="B903EAB4FFCCFFC3CE92B28DFAA21BFA" box="[1247,1317,421,445]" italics="true" pageId="8" pageNumber="409">3c, 4a</emphasis>
–
<emphasis id="B903EAB4FFCCFFC3CF7EB28EFAB81BFA" box="[1331,1343,422,445]" italics="true" pageId="8" pageNumber="409">c</emphasis>
slightly longer than others (9–12 Μm);
<emphasis id="B903EAB4FFCCFFC3CBBAB2E0FD941BA6" box="[503,531,456,481]" italics="true" pageId="8" pageNumber="409">3b</emphasis>
absent. Epimeral plates I rounded anterolaterally; all plates separated medially (
<figureCitation id="134C2A23FFCCFFC3CAD2B2C4FF651842" box="[159,226,492,517]" captionStart="FIGURE 5" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[162,1420,819,1883]" captionTargetId="figure@8.[150,1436,788,1896]" captionTargetPageId="8" captionText="FIGURE 5. Tyrphonothrus seniczaki sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223439/files/figure.png" pageId="8" pageNumber="409">Fig. 5</figureCitation>
b). Apodeme IV straight, transverse. Epimeral plate IV rounded laterally.
</paragraph>
<paragraph id="8BC836A6FFCCFFC3CA8AB138FED918FB" blockId="8.[151,1436,151,736]" pageId="8" pageNumber="409">
<emphasis id="B903EAB4FFCCFFC3CA8AB138FE1E186E" box="[199,409,528,553]" italics="true" pageId="8" pageNumber="409">Anogenital Region</emphasis>
. With six pairs of smooth, fine, flagelliform genital setae, sub-equal in length (12–18 Μm),
<emphasis id="B903EAB4FFCCFFC3CADAB11EFF22180A" box="[151,165,566,589]" italics="true" pageId="8" pageNumber="409">g</emphasis>
1-3 closely spaced,
<emphasis id="B903EAB4FFCCFFC3CB3EB11EFE06180A" box="[371,385,566,589]" italics="true" pageId="8" pageNumber="409">g</emphasis>
4-6 spaced more broadly (
<figureCitation id="134C2A23FFCCFFC3C8DAB11CFD5D180A" box="[663,730,564,589]" captionStart="FIGURE 5" captionStartId="8.[151,250,1918,1940]" captionTargetBox="[162,1420,819,1883]" captionTargetId="figure@8.[150,1436,788,1896]" captionTargetPageId="8" captionText="FIGURE 5. Tyrphonothrus seniczaki sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223439/files/figure.png" pageId="8" pageNumber="409">Fig. 5</figureCitation>
b). Genital plates sub-rectangular, rounded apically; each plate 97 Μm long, 72 Μm broad. Posterior margin of genital plate transverse, overlapping adanal plate. Each adanal plate 134 Μm long, 33 Μm broad; three pairs of smooth, short, setiform adanal setae, 11–14 Μm long. One pair of anal setae 14 Μm long.
</paragraph>
<paragraph id="8BC836A6FFCCFFC3CA8AB1E0FDC618A7" blockId="8.[151,1436,151,736]" box="[199,577,711,736]" pageId="8" pageNumber="409">
<emphasis id="B903EAB4FFCCFFC3CA8AB1E0FF7B1898" box="[199,252,712,735]" italics="true" pageId="8" pageNumber="409">Legs</emphasis>
. Pre-tarsi heterotridactylous.
</paragraph>
</subSubSection>
<caption id="DF08662EFFCCFFC3CADAB456FC731DD3" box="[151,1012,1918,1940]" httpUri="https://zenodo.org/record/223439/files/figure.png" pageId="8" pageNumber="409" targetBox="[162,1420,819,1883]" targetPageId="8">
<paragraph id="8BC836A6FFCCFFC3CADAB456FC731DD3" blockId="8.[151,1012,1918,1940]" box="[151,1012,1918,1940]" pageId="8" pageNumber="409">
<emphasis id="B903EAB4FFCCFFC3CADAB456FE941DD3" bold="true" box="[151,275,1918,1940]" pageId="8" pageNumber="409">FIGURE 5.</emphasis>
<taxonomicName id="4C774D25FFCCFFC3CB54B456FD961DD3" box="[281,529,1918,1940]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="seniczaki" status="sp. nov.">
<emphasis id="B903EAB4FFCCFFC3CB54B456FD961DD3" box="[281,529,1918,1940]" italics="true" pageId="8" pageNumber="409">Tyrphonothrus seniczaki</emphasis>
</taxonomicName>
<emphasis id="B903EAB4FFCCFFC3C85AB457FDE01DD3" bold="true" box="[535,615,1919,1940]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A23057CFFFCCFFC3C85AB457FDE01DD3" box="[535,615,1919,1940]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female; a) dorsal; b) ventral.
</paragraph>
</caption>
<subSubSection id="C36D652DFFCCFFC2CA8AB493FE841A92" lastPageId="9" lastPageNumber="410" pageId="8" pageNumber="409" type="materials_examined">
<paragraph id="8BC836A6FFCCFFC3CA8AB493FA1B1D94" blockId="8.[199,1436,1979,2004]" box="[199,1436,1979,2004]" pageId="8" pageNumber="409">
<emphasis id="B903EAB4FFCCFFC3CA8AB493FE351D93" bold="true" box="[199,434,1979,2004]" pageId="8" pageNumber="409">Material examined.</emphasis>
Wet moss, Cumberland Memorial Scenic Reserve, Victoria,
<geoCoordinate id="EE435061FFCCFFC3CE12B494FB341D93" box="[1119,1203,1980,2004]" direction="south" orientation="latitude" pageId="8" pageNumber="409" precision="925" value="-37.55">37º33'S</geoCoordinate>
<geoCoordinate id="EE435061FFCCFFC3CEF1B493FAA61D93" box="[1212,1313,1979,2004]" direction="east" orientation="longitude" pageId="8" pageNumber="409" precision="925" value="145.86667">145º52'E</geoCoordinate>
, coll. R.E.
</paragraph>
<paragraph id="8BC836A6FFCDFFC2CADAB3BFFE841A92" blockId="9.[151,1436,151,693]" pageId="9" pageNumber="410">
Rougley,
<date id="FFC91066FFCDFFC2CB4CB3BFFEFB1AF7" box="[257,380,151,176]" pageId="9" pageNumber="410" value="1996-03-12">12.iii.1996</date>
.
<typeStatus id="54CC8804FFCDFFC2CBCAB3BFFE6A1AF7" box="[391,493,151,176]" pageId="9" pageNumber="410" type="holotype">Holotype</typeStatus>
deposited in the Australian National Insect Collection, CSIRO Ecosystem Sciences, Canberra.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFCDFFC2CA8AB3F7FBB01B07" pageId="9" pageNumber="410" type="etymology">
<paragraph id="8BC836A6FFCDFFC2CA8AB3F7FBB01B07" blockId="9.[151,1436,151,693]" pageId="9" pageNumber="410">
<emphasis id="B903EAB4FFCDFFC2CA8AB3F7FEC91ABF" bold="true" box="[199,334,223,248]" pageId="9" pageNumber="410">Etymology.</emphasis>
This species is named in honour of Professor Stanisław Seniczak (Department of Ecology, University of Technology and Life Sciences, Bydgoszcz,
<collectingCountry id="F3607636FFCDFFC2C965B22CFCFF1B5A" box="[808,888,260,285]" name="Poland" pageId="9" pageNumber="410">Poland</collectingCountry>
) in recognition of his contribution to acarology, particularly the study of the taxonomy, morphology and ontogeny of the
<taxonomicName id="4C774D25FFCDFFC2C988B20FFBB51B07" box="[965,1074,295,320]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oribatida" pageId="9" pageNumber="410" phylum="Arthropoda" rank="order">Oribatida</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFCDFFC2CA8AB264FA1B18F7" pageId="9" pageNumber="410" type="discussion">
<paragraph id="8BC836A6FFCDFFC2CA8AB264FB521BBE" blockId="9.[151,1436,151,693]" pageId="9" pageNumber="410">
<emphasis id="B903EAB4FFCDFFC2CA8AB264FEB31B22" bold="true" box="[199,308,332,357]" pageId="9" pageNumber="410">Remarks</emphasis>
.
<taxonomicName id="4C774D25FFCDFFC2CB05B264FDE61B22" box="[328,609,332,357]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="seniczaki" status="sp. nov.">
<emphasis id="B903EAB4FFCDFFC2CB05B264FDE61B22" box="[328,609,332,357]" italics="true" pageId="9" pageNumber="410">Tyrphonothrus seniczaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFCDFFC2C823B264FD491B23" box="[622,718,332,357]" pageId="9" pageNumber="410" rank="species">
<emphasis id="B903EAB4FFCDFFC2C823B264FD4F1B22" bold="true" box="[622,712,332,357]" pageId="9" pageNumber="410">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from other members of the genus by the following combination of characters:1) the very short lamellar setae; 2) the very long interlamellar setae, 10 × longer than
<emphasis id="B903EAB4FFCDFFC2CF36B259FA131BCF" box="[1403,1428,369,392]" italics="true" pageId="9" pageNumber="410">ex</emphasis>
; 3) setae
<emphasis id="B903EAB4FFCDFFC2CAB2B2BDFE8C1BEB" box="[255,267,405,428]" italics="true" pageId="9" pageNumber="410">c</emphasis>
1 on the anterior margin of the notogaster; 4) setae
<emphasis id="B903EAB4FFCDFFC2C93AB2BCFC021BEA" box="[887,901,404,429]" italics="true" pageId="9" pageNumber="410">d</emphasis>
1 positioned markedly anteriorly, almost level transversely with bases of
<emphasis id="B903EAB4FFCDFFC2CBF0B294FE4E1B94" box="[445,457,444,467]" italics="true" pageId="9" pageNumber="410">c</emphasis>
3; 5) the minute setae
<emphasis id="B903EAB4FFCDFFC2C8F5B294FD431B94" box="[696,708,444,467]" italics="true" pageId="9" pageNumber="410">e</emphasis>
1; 6) the thin, flagelliform setae
<emphasis id="B903EAB4FFCDFFC2CE6EB294FBA81B94" box="[1059,1071,444,467]" italics="true" pageId="9" pageNumber="410">e</emphasis>
2,
<emphasis id="B903EAB4FFCDFFC2CE0FB292FBD71B94" box="[1090,1104,442,467]" italics="true" pageId="9" pageNumber="410">h</emphasis>
1 and
<emphasis id="B903EAB4FFCDFFC2CEC1B292FB1D1B94" box="[1164,1178,442,467]" italics="true" pageId="9" pageNumber="410">h</emphasis>
2; 7) the short, diagonal lateral notogastral ridges; 8) the epimeral setal formula 3-1-2-3; 9) with six pairs of genital setae.
</paragraph>
<paragraph id="8BC836A6FFCDFFC2CA8AB12DFA1B18F7" blockId="9.[151,1436,151,693]" pageId="9" pageNumber="410">
<taxonomicName id="4C774D25FFCDFFC2CA8AB12DFE5F1859" box="[199,472,517,542]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="seniczaki" status="sp. nov.">
<emphasis id="B903EAB4FFCDFFC2CA8AB12DFE5F1859" box="[199,472,517,542]" italics="true" pageId="9" pageNumber="410">Tyrphonothrus seniczaki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFCDFFC2CB92B12DFDBF185A" box="[479,568,517,542]" pageId="9" pageNumber="410" rank="species">
<emphasis id="B903EAB4FFCDFFC2CB92B12DFDB61859" bold="true" box="[479,561,517,542]" pageId="9" pageNumber="410">sp. nov</emphasis>
.
</taxonomicNameLabel>
is morphologically most similar to
<taxonomicName id="4C774D25FFCDFFC2C98BB12EFF481805" authority="Yamamoto, Kuriki &amp; Aoki, 1993" authorityName="Yamamoto, Kuriki &amp; Aoki" authorityYear="1993" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="azumaensis">
<emphasis id="B903EAB4FFCDFFC2C98BB12EFBE5185A" box="[966,1122,518,541]" italics="true" pageId="9" pageNumber="410">T. azumaensis</emphasis>
Yamamoto, Kuriki &amp; Aoki, 1993
</taxonomicName>
and
<taxonomicName id="4C774D25FFCDFFC2CB44B102FDEE1805" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" box="[265,617,553,578]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="longirostrum">
<emphasis id="B903EAB4FFCDFFC2CB44B102FE301805" box="[265,439,553,578]" italics="true" pageId="9" pageNumber="410">T. longirostrum</emphasis>
Hammer, 1966
</taxonomicName>
a in that all three species have the posterior M-shaped notogastral ridge complex reduced to only its acutely-sloping lateral elements (bearing setae
<emphasis id="B903EAB4FFCDFFC2CE41B165FB9D1821" box="[1036,1050,589,614]" italics="true" pageId="9" pageNumber="410">h</emphasis>
1) and the longitudinal ridges are absent. These species have seven pairs of genital and two pairs of anal setae compared with six pairs and one pair in
<emphasis id="B903EAB4FFCDFFC2CAFEB1B1FEBB18F6" box="[179,316,664,689]" italics="true" pageId="9" pageNumber="410">
<taxonomicName id="4C774D25FFCDFFC2CAFEB1B1FEBF18F6" box="[179,312,664,689]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="seniczaki">T. seniczaki</taxonomicName>
.
</emphasis>
In the latter species setae of the
<emphasis id="B903EAB4FFCDFFC2C8E9B1B1FD3718F7" box="[676,688,665,688]" italics="true" pageId="9" pageNumber="410">c</emphasis>
and
<emphasis id="B903EAB4FFCDFFC2C8ABB1B0FD7318F6" box="[742,756,664,689]" italics="true" pageId="9" pageNumber="410">d</emphasis>
series and
<emphasis id="B903EAB4FFCDFFC2C93DB1B1FCFB18F7" box="[880,892,665,688]" italics="true" pageId="9" pageNumber="410">e</emphasis>
1 are considerably shorter than in the former two.
</paragraph>
</subSubSection>
</treatment>
</document>
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