270 lines
32 KiB
XML
270 lines
32 KiB
XML
<document id="677E510C412070812E5D84AAFFC8F3EC" ID-CLB-Dataset="45654" ID-DOI="10.5281/zenodo.279712" ID-GBIF-Dataset="e43a19ff-986e-4000-b2d4-ae3ae60a753f" ID-ISSN="1175-5326" ID-Zenodo-Dep="279712" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460427389018" checkinUser="plazi" docAuthor="Akhmetova, Lilia & Frolov, Andrey" docDate="2012" docId="03C65D478D16FFF0FF04FA5095D0FF3D" docLanguage="en" docName="zt03159p068.pdf" docOrigin="Zootaxa 3159" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Aphodius (Melinopterus) gissaricus Akhmetova & Frolov, new species" docType="treatment" docVersion="8" lastPageNumber="68" masterDocId="FFFF253F8D16FFF3FF93FD46952FFFEC" masterDocTitle="A new scarab species, Aphodius gissaricus (Coleoptera: Scarabaeidae: Aphodiinae), from the Pamir-Alay Mountains in Tajikistan" masterLastPageNumber="68" masterPageNumber="65" pageNumber="65" updateTime="1698330905583" updateUser="ExternalLinkService">
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<mods:titleInfo id="3A55A887054AF50337DE745AEE13064B">
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<mods:title id="B3F3396E2450C2B45152A90B60E1A3CC">A new scarab species, Aphodius gissaricus (Coleoptera: Scarabaeidae: Aphodiinae), from the Pamir-Alay Mountains in Tajikistan</mods:title>
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<mods:namePart id="48F854675D1A0EE4F9BE9545ACF5DBBB">Akhmetova, Lilia</mods:namePart>
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</mods:name>
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<mods:roleTerm id="409BC0CE8075C9118B83CFE3D565A765">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BFF23BBF593C80E4623049DD837D05B3">Frolov, Andrey</mods:namePart>
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<mods:typeOfResource id="3EFDAAFBD955F3D5C303E37DF2F9874A">text</mods:typeOfResource>
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<mods:title id="9B55A70F3ECE2436A6B4D32E65A0359F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3AEC21F870D65F8F5174FCF0D9507C1C">
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<mods:date id="CFB847E6A8CE4C3F40B8743E36456177">2012</mods:date>
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<mods:number id="9441D250C476A9EC120D4719A7022B56">3159</mods:number>
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<mods:classification id="BD179573668DC7898FF183B88B887DA0">journal article</mods:classification>
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<treatment id="03C65D478D16FFF0FF04FA5095D0FF3D" ID-DOI="http://doi.org/10.5281/zenodo.6174407" ID-GBIF-Taxon="119428946" ID-Zenodo-Dep="6174407" LSID="urn:lsid:plazi:treatment:03C65D478D16FFF0FF04FA5095D0FF3D" httpUri="http://treatment.plazi.org/id/03C65D478D16FFF0FF04FA5095D0FF3D" lastPageId="3" lastPageNumber="68" pageId="0" pageNumber="65">
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<subSubSection id="C375BFDA8D16FFF3FF04FA50946CF8BC" pageId="0" pageNumber="65" type="nomenclature">
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<paragraph id="8BD0EC518D16FFF3FF04FA5096DAF8DC" blockId="0.[151,1013,1814,1872]" box="[151,1013,1814,1840]" pageId="0" pageNumber="65">
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<heading id="D0985B3D8D16FFF3FF04FA5096DAF8DC" bold="true" box="[151,1013,1814,1840]" fontSize="11" level="1" pageId="0" pageNumber="65" reason="1">
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<emphasis id="B91B30438D16FFF3FF04FA5096DAF8DC" bold="true" box="[151,1013,1814,1840]" pageId="0" pageNumber="65">
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<taxonomicName id="4C6F97D28D16FFF3FF04FA509677F8DC" authority="Akhmetova & Frolov" authorityName="Akhmetova & Frolov" box="[151,856,1814,1840]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="gissaricus" status="sp. nov." subGenus="Melinopterus">
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<emphasis id="B91B30438D16FFF3FF04FA509424F8DC" bold="true" box="[151,267,1814,1840]" italics="true" pageId="0" pageNumber="65">Aphodius</emphasis>
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(
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<emphasis id="B91B30438D16FFF3FE8FFA509492F8DC" bold="true" box="[284,445,1814,1840]" italics="true" pageId="0" pageNumber="65">Melinopterus</emphasis>
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)
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<emphasis id="B91B30438D16FFF3FE5DFA509769F8DC" bold="true" box="[462,582,1814,1840]" italics="true" pageId="0" pageNumber="65">gissaricus</emphasis>
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Akhmetova & Frolov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2288D388D16FFF3FCF6FA5096DAF8DC" box="[869,1013,1814,1840]" pageId="0" pageNumber="65" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD0EC518D16FFF3FF04FA7F946CF8BC" blockId="0.[151,1013,1814,1872]" box="[151,323,1849,1872]" pageId="0" pageNumber="65">
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<figureCitation id="1354F0D48D16FFF3FF04FA7F9405F8BC" box="[151,298,1849,1872]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1879,1903]" captionTargetBox="[151,1427,272,1849]" captionTargetId="figure@1.[151,1436,263,1855]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Aphodius. A. gissaricus, holotype, male (1, 7); paratype, male (2, 3, 5); A. scuticollis male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae." httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="0" pageNumber="65">Figs 1–3, 5, 7</figureCitation>
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,
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<figureCitation id="1354F0D48D16FFF3FEA5FA7F946CF8BC" box="[310,323,1849,1872]" captionStart="FIGURE 9" captionStartId="2.[151,255,1988,2012]" captionTargetBox="[151,1436,1328,1964]" captionTargetId="figure@2.[151,1436,1328,1964]" captionTargetPageId="2" captionText="FIGURE 9. Distribution map of Aphodius gissaricus (▲) and A. scuticollis (■)." httpUri="https://zenodo.org/record/279714/files/figure.png" pageId="0" pageNumber="65">9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C375BFDA8D16FFF2FF04FA389765FF3D" lastPageId="1" lastPageNumber="66" pageId="0" pageNumber="65" type="materials_examined">
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<paragraph id="8BD0EC518D16FFF2FF04FA389765FF3D" blockId="0.[151,1436,1918,2010]" lastBlockId="1.[151,1436,152,209]" lastPageId="1" lastPageNumber="66" pageId="0" pageNumber="65">
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<emphasis id="B91B30438D16FFF3FF04FA389412F87A" bold="true" box="[151,317,1918,1942]" pageId="0" pageNumber="65">
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<typeStatus id="54D452F38D16FFF3FF04FA3895E0F87A" box="[151,207,1918,1942]" pageId="0" pageNumber="65">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54D452F38D16FFF3FED7FA389485F879" box="[324,426,1918,1941]" pageId="0" pageNumber="65" type="holotype">Holotype</typeStatus>
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, male with the label “
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<collectingCountry id="F378ACC18D16FFF3FD06FA389630F879" box="[661,799,1918,1941]" name="Tajikistan" pageId="0" pageNumber="65">Tadzhikistan</collectingCountry>
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1984 Hissar mts., ca
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<quantity id="4C9741B48D16FFF3FB99FA38917BF879" box="[1034,1108,1918,1941]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="0" pageNumber="65" unit="m" value="3300.0">3300m</quantity>
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<collectionCode id="ED7E74948D16FFF3FBC8FA389198F879" box="[1115,1207,1918,1941]" pageId="0" pageNumber="65">ANZOB</collectionCode>
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pass, 8.8 David Král lgt.” (
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<collectionCode id="ED7E74948D16FFF3FF40FAE7940AF854" box="[211,293,1953,1976]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="0" pageNumber="65">NMPC</collectionCode>
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). Eleven
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<typeStatus id="54D452F38D16FFF3FE15FAE794DEF854" box="[390,497,1953,1976]" pageId="0" pageNumber="65" type="paratype">paratypes</typeStatus>
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:
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<specimenCount id="9D6927D88D16FFF3FE69FAE79764F854" box="[506,587,1953,1976]" pageId="0" pageNumber="65" type="male">7 males</specimenCount>
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||
with the same data as the
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<typeStatus id="54D452F38D16FFF3FCF2FAE79691F854" box="[865,958,1953,1976]" pageId="0" pageNumber="65" type="holotype">holotype</typeStatus>
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(
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<specimenCount id="9D6927D88D16FFF3FC5DFAE7917FF854" box="[974,1104,1953,1976]" pageId="0" pageNumber="65" type="generic">5 specimens</specimenCount>
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–
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<collectionCode id="ED7E74948D16FFF3FBF9FAE79197F854" box="[1130,1208,1953,1976]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="0" pageNumber="65">NMPC</collectionCode>
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,
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<specimenCount id="9D6927D88D16FFF3FB50FAE7906AF854" box="[1219,1349,1953,1976]" pageId="0" pageNumber="65" type="generic">2 specimens</specimenCount>
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||
–
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<collectionCode id="ED7E74948D16FFF3FACCFAE790BFF854" box="[1375,1424,1953,1976]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="0" pageNumber="65">ZIN</collectionCode>
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);
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<specimenCount id="9D6927D88D16FFF3FF04FA8595C3F836" box="[151,236,1987,2010]" pageId="0" pageNumber="65" type="male">3 males</specimenCount>
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||
, “Asia c.
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<collectingCountry id="F378ACC18D16FFF3FEC1FA8594E7F836" box="[338,456,1987,2010]" name="Tajikistan" pageId="0" pageNumber="65">Tadjikistan</collectingCountry>
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||
<collectionCode id="ED7E74948D16FFF3FE5CFA85973DF836" box="[463,530,1987,2010]" pageId="0" pageNumber="65">USSR</collectionCode>
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||
<date id="FFD1CA918D16FFF3FD8BFA85975AF836" box="[536,629,1987,2010]" pageId="0" pageNumber="65" value="1986-04-05">5.4.1986</date>
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||
Alaj – Romit
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||
<quantity id="4C9741B48D16FFF3FC9FFA859672F836" box="[780,861,1987,2010]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="65" unit="m" value="2000.0">2000 m</quantity>
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||
T. Růžička lgt.” (
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<specimenCount id="9D6927D88D16FFF3FB8AFA8591B2F836" box="[1049,1181,1987,2010]" pageId="0" pageNumber="65" type="generic">2 specimens</specimenCount>
|
||
–
|
||
<collectionCode id="ED7E74948D16FFF3FB24FA85902AF836" box="[1207,1285,1987,2010]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="0" pageNumber="65">NMPC</collectionCode>
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||
,
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||
<specimenCount id="9D6927D88D16FFF3FA83FA8590A6F836" box="[1296,1417,1987,2010]" pageId="0" pageNumber="65" type="generic">1 specimen</specimenCount>
|
||
–
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||
<collectionCode id="ED7E74948D17FFF2FF04FDDE95E7FF43" box="[151,200,152,175]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="66">ZIN</collectionCode>
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||
);
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||
<specimenCount id="9D6927D88D17FFF2FF49FDDE940BFF43" box="[218,292,152,175]" pageId="1" pageNumber="66" type="male">1 male</specimenCount>
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, “
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<collectingCountry id="F378ACC18D17FFF2FEA4FDDE94EBFF43" box="[311,452,152,175]" name="Tajikistan" pageId="1" pageNumber="66">Tadzhikistan</collectingCountry>
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||
, yugo-zap. sklon khr. Aktau [south-western slope of Aktau Ridge],
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||
<quantity id="4C9741B48D17FFF2FB1CFDDE91CDFF43" box="[1167,1250,152,175]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="66" unit="m" value="2000.0">2000 m</quantity>
|
||
.
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<date id="FFD1CA918D17FFF2FB78FDDE904EFF43" box="[1259,1377,152,175]" pageId="1" pageNumber="66" value="1984-04-13">13.IV.1984</date>
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T. N. Vereschagina” [in Cyrillic letters] (
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<collectionCode id="ED7E74948D17FFF2FD9FFDFC9711FF3D" box="[524,574,186,209]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="66">ZIN</collectionCode>
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||
).
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF10BCD98D17FFF2FF04FA119422F85F" httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="1" pageNumber="66" targetBox="[151,1427,272,1849]" targetPageId="1">
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||
<paragraph id="8BD0EC518D17FFF2FF04FA119422F85F" blockId="1.[151,1437,1879,1971]" pageId="1" pageNumber="66">
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||
<emphasis id="B91B30438D17FFF2FF04FA11946DF883" bold="true" box="[151,322,1879,1903]" pageId="1" pageNumber="66">FIGURES 1–8.</emphasis>
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<taxonomicName id="4C6F97D28D17FFF2FEDBFA1E9482F883" box="[328,429,1880,1903]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B91B30438D17FFF2FEDBFA1E9482F883" box="[328,429,1880,1903]" italics="true" pageId="1" pageNumber="66">Aphodius</emphasis>
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||
</taxonomicName>
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||
.
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||
<taxonomicName id="4C6F97D28D17FFF2FE29FA1E976CF883" box="[442,579,1880,1903]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="66" phylum="Arthropoda" rank="species" species="gissaricus">
|
||
<emphasis id="B91B30438D17FFF2FE29FA1E976CF883" box="[442,579,1880,1903]" italics="true" pageId="1" pageNumber="66">A. gissaricus</emphasis>
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||
</taxonomicName>
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||
, holotype, male (1, 7); paratype, male (2, 3, 5);
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<taxonomicName id="4C6F97D28D17FFF2FBADFA1E91E8F883" box="[1086,1223,1880,1903]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="66" phylum="Arthropoda" rank="species" species="scuticollis">
|
||
<emphasis id="B91B30438D17FFF2FBADFA1E91E8F883" box="[1086,1223,1880,1903]" italics="true" pageId="1" pageNumber="66">A. scuticollis</emphasis>
|
||
</taxonomicName>
|
||
male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae.
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||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C375BFDA8D14FFF1FF54FDD194B6FC39" pageId="2" pageNumber="67" type="description">
|
||
<paragraph id="8BD0EC518D14FFF1FF54FDD191EBFED9" blockId="2.[151,1437,151,1284]" pageId="2" pageNumber="67">
|
||
<emphasis id="B91B30438D14FFF1FF54FDD1947DFF43" bold="true" box="[199,338,151,175]" pageId="2" pageNumber="67">Description.</emphasis>
|
||
<typeStatus id="54D452F38D14FFF1FECFFDDE94EDFF43" box="[348,450,152,175]" pageId="2" pageNumber="67" type="holotype">Holotype</typeStatus>
|
||
, male (
|
||
<figureCitation id="1354F0D48D14FFF1FD85FDDE9776FF43" box="[534,601,152,175]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1879,1903]" captionTargetBox="[151,1427,272,1849]" captionTargetId="figure@1.[151,1436,263,1855]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Aphodius. A. gissaricus, holotype, male (1, 7); paratype, male (2, 3, 5); A. scuticollis male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae." httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="2" pageNumber="67">Fig. 1</figureCitation>
|
||
). Body length 5.0 mm. Head shiny, with dark brown disc and pale brown anterior margin and sides of clypeus, densely punctate (punctures separated by approximately their diameter). Clypeus with feeble sinuation anteriorly, rounded laterally. Genae rounded, weakly separated from lateral margin of clypeus, distinctly protruding past eyes, with a few long setae. Frontoclypeal suture visible as fine line, interrupted medially. Head disc without tubercles. Eye approximately as wide as distance between eye and gula in ventral view.
|
||
</paragraph>
|
||
<paragraph id="8BD0EC518D14FFF1FF54FC0691BFFE76" blockId="2.[151,1437,151,1284]" pageId="2" pageNumber="67">Pronotum wider than elytral base, shiny, pale brown on disc and pale yellow laterally. Pronotum apical margin without border, lateral margins and base with thin border. Disc with sparse, fine punctation (punctures separated by 1–4 times their diameter). Basal angles of pronotum rounded; sides with sparse, short, yellow setae.</paragraph>
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||
<paragraph id="8BD0EC518D14FFF1FF54FCE396E1FE50" blockId="2.[151,1437,151,1284]" box="[199,974,421,444]" pageId="2" pageNumber="67">Scutellum narrow, triangular, shiny, pale brown with brown lateral sides.</paragraph>
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||
<paragraph id="8BD0EC518D14FFF1FF54FC809030FE13" blockId="2.[151,1437,151,1284]" pageId="2" pageNumber="67">Elytra pale yellow with brown suture. Humeral teeth absent. Elytral intervals feebly convex, finely and sparsely punctate. Sides and apices of elytra without setae. Striae deep, punctures on striae larger than width of striae.</paragraph>
|
||
<paragraph id="8BD0EC518D14FFF1FF54FF4F946CFDAE" blockId="2.[151,1437,151,1284]" pageId="2" pageNumber="67">Ventral side of body pale yellow except abdomen brown. Legs pale yellow. Perimeter of metasternal disc with dense, long, pale setae.</paragraph>
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<paragraph id="8BD0EC518D14FFF1FF54FF0B959BFD24" blockId="2.[151,1437,151,1284]" pageId="2" pageNumber="67">
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Outer teeth of protibia long and slender. Spur of protibia slightly curved downward and rounded apically. Lower spur of mesotibia acute apically, more than half as long as upper spur. Metatarsomere 1 as long as upper spur of metatibia and metatarsomeres 2–3 combined. Adjacent apical setae of mesotibiae and metatibiae distinctly unequal in length (
|
||
<figureCitation id="1354F0D48D14FFF1FAE2FFD69588FD24" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1879,1903]" captionTargetBox="[151,1427,272,1849]" captionTargetId="figure@1.[151,1436,263,1855]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Aphodius. A. gissaricus, holotype, male (1, 7); paratype, male (2, 3, 5); A. scuticollis male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae." httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="2" pageNumber="67">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD0EC518D14FFF1FF54FF959678FD06" blockId="2.[151,1437,151,1284]" box="[199,855,723,746]" pageId="2" pageNumber="67">Parameres feebly curved, tapering and slightly acute apically.</paragraph>
|
||
<paragraph id="8BD0EC518D14FFF1FF54FFB294B7FCE0" blockId="2.[151,1437,151,1284]" box="[199,408,756,780]" pageId="2" pageNumber="67">
|
||
<emphasis id="B91B30438D14FFF1FF54FFB2940EFCE0" bold="true" box="[199,289,756,780]" pageId="2" pageNumber="67">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BD0EC518D14FFF1FF54FE5094B6FC39" blockId="2.[151,1437,151,1284]" pageId="2" pageNumber="67">
|
||
<emphasis id="B91B30438D14FFF1FF54FE509469FCC2" bold="true" box="[199,326,790,814]" pageId="2" pageNumber="67">Variability.</emphasis>
|
||
Body length of the
|
||
<typeStatus id="54D452F38D14FFF1FDB4FE5097A2FCC1" box="[551,653,790,813]" pageId="2" pageNumber="67" type="paratype">paratypes</typeStatus>
|
||
varies from 3.5–5.0 mm. Specimens from Romit are darker than those from Anzob Pass and Aktau Mountains. The coloration of elytra varies: one
|
||
<typeStatus id="54D452F38D14FFF1FC5CFE7E9104FCA3" box="[975,1067,824,847]" pageId="2" pageNumber="67" type="paratype">paratype</typeStatus>
|
||
has a dark, obscure elytral pattern (
|
||
<figureCitation id="1354F0D48D14FFF1FF0CFE1F95CFFC9C" box="[159,224,857,880]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1879,1903]" captionTargetBox="[151,1427,272,1849]" captionTargetId="figure@1.[151,1436,263,1855]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Aphodius. A. gissaricus, holotype, male (1, 7); paratype, male (2, 3, 5); A. scuticollis male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae." httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="2" pageNumber="67">Fig. 3</figureCitation>
|
||
), while the
|
||
<typeStatus id="54D452F38D14FFF1FEF2FE1F9491FC9C" box="[353,446,857,880]" pageId="2" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
has almost uniformly pale yellow elytra (
|
||
<figureCitation id="1354F0D48D14FFF1FC10FE1F96ECFC9C" box="[899,963,857,880]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1879,1903]" captionTargetBox="[151,1427,272,1849]" captionTargetId="figure@1.[151,1436,263,1855]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Aphodius. A. gissaricus, holotype, male (1, 7); paratype, male (2, 3, 5); A. scuticollis male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae." httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="2" pageNumber="67">Fig. 1</figureCitation>
|
||
). An elytral pattern with a few longitudinal maculae (similar to that in
|
||
<taxonomicName id="4C6F97D28D14FFF1FE2DFE3D9766FC7E" box="[446,585,891,914]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="scuticollis">
|
||
<emphasis id="B91B30438D14FFF1FE2DFE3D9766FC7E" box="[446,585,891,914]" italics="true" pageId="2" pageNumber="67">A. scuticollis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="1354F0D48D14FFF1FDCBFE3D97B6FC7E" box="[600,665,891,914]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1879,1903]" captionTargetBox="[151,1427,272,1849]" captionTargetId="figure@1.[151,1436,263,1855]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Aphodius. A. gissaricus, holotype, male (1, 7); paratype, male (2, 3, 5); A. scuticollis male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae." httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="2" pageNumber="67">Fig. 4</figureCitation>
|
||
) is found in most of other
|
||
<typeStatus id="54D452F38D14FFF1FC52FE3D9105FC7E" box="[961,1066,891,914]" pageId="2" pageNumber="67" type="paratype">paratypes</typeStatus>
|
||
, but the maculae are only slightly darker than the background and the pattern is feebly distinct. The sides and apices of the elytra of a few
|
||
<typeStatus id="54D452F38D14FFF1FA93FEDB9049FC58" box="[1280,1382,925,948]" pageId="2" pageNumber="67" type="paratype">paratypes</typeStatus>
|
||
bear short, sparse, pale setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375BFDA8D14FFF1FF54FE99960FFB73" pageId="2" pageNumber="67" type="diagnosis">
|
||
<paragraph id="8BD0EC518D14FFF1FF54FE9997E2FBD6" blockId="2.[151,1437,151,1284]" pageId="2" pageNumber="67">
|
||
<emphasis id="B91B30438D14FFF1FF54FE999413FC1B" bold="true" box="[199,316,991,1015]" pageId="2" pageNumber="67">Diagnosis.</emphasis>
|
||
The species is similar to
|
||
<taxonomicName id="4C6F97D28D14FFF1FDDDFEA696ABFC1B" authority="Semenov, 1898" authorityName="Semenov" authorityYear="1898" box="[590,900,992,1015]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="scuticollis">
|
||
<emphasis id="B91B30438D14FFF1FDDDFEA697F7FC1B" box="[590,728,992,1015]" italics="true" pageId="2" pageNumber="67">A. scuticollis</emphasis>
|
||
Semenov, 1898
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1354F0D48D14FFF1FC00FEA6913AFC1B" box="[915,1045,992,1015]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1879,1903]" captionTargetBox="[151,1427,272,1849]" captionTargetId="figure@1.[151,1436,263,1855]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Aphodius. A. gissaricus, holotype, male (1, 7); paratype, male (2, 3, 5); A. scuticollis male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae." httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="2" pageNumber="67">Figs. 4, 6, 8</figureCitation>
|
||
), but differs from it in lacking acute process on the apex of metatibia in males (
|
||
<figureCitation id="1354F0D48D14FFF1FDCCF9479795FBF4" box="[607,698,1025,1048]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1879,1903]" captionTargetBox="[151,1427,272,1849]" captionTargetId="figure@1.[151,1436,263,1855]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Aphodius. A. gissaricus, holotype, male (1, 7); paratype, male (2, 3, 5); A. scuticollis male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae." httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="2" pageNumber="67">Fig. 7, 8</figureCitation>
|
||
), in having less convex body, and in having slightly longer and more slender parameres with less curved apices (
|
||
<figureCitation id="1354F0D48D14FFF1FDF7F96597EFFBD6" box="[612,704,1059,1082]" captionStart="FIGURES 1 – 8" captionStartId="1.[151,269,1879,1903]" captionTargetBox="[151,1427,272,1849]" captionTargetId="figure@1.[151,1436,263,1855]" captionTargetPageId="1" captionText="FIGURES 1 – 8. Aphodius. A. gissaricus, holotype, male (1, 7); paratype, male (2, 3, 5); A. scuticollis male (4, 6, 8). Figs. 1, 3, 4 — body in dorsal view; 2 — body in ventral view; 5, 6, — aedeagus in dorsal and lateral view; 7, 8 — apices of metatibiae." httpUri="https://zenodo.org/record/279713/files/figure.png" pageId="2" pageNumber="67">Fig. 5, 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD0EC518D14FFF1FF54F903960FFB73" blockId="2.[151,1437,151,1284]" pageId="2" pageNumber="67">
|
||
In the key to Palaearctic
|
||
<taxonomicName id="4C6F97D28D14FFF1FE48F903962FFBB0" authority="Balthasar 1964" authorityName="Balthasar" authorityYear="1964" box="[475,768,1093,1116]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91B30438D14FFF1FE48F903976FFBB0" box="[475,576,1093,1116]" italics="true" pageId="2" pageNumber="67">Aphodius</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFE91A08D14FFF1FDC3F90397D6FBB0" author="Balthasar" box="[592,761,1093,1116]" pageId="2" pageNumber="67" refString="Balthasar, V. (1964) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera Lamellicornia. Aphodiidae. Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague, 3, 652 pp." type="book" year="1964">Balthasar 1964</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6F97D28D14FFF1FC9EF90396B6FBB0" box="[781,921,1093,1116]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="gissaricus">
|
||
<emphasis id="B91B30438D14FFF1FC9EF90396B6FBB0" box="[781,921,1093,1116]" italics="true" pageId="2" pageNumber="67">A. gissaricus</emphasis>
|
||
</taxonomicName>
|
||
will key to the couplet with
|
||
<taxonomicName id="4C6F97D28D14FFF1FB4FF903940FFB91" authority="Reitter" authorityName="Reitter" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="stolzi" subGenus="Melinopterus">
|
||
<emphasis id="B91B30438D14FFF1FB4FF90391C3FBB0" box="[1244,1260,1093,1116]" italics="true" pageId="2" pageNumber="67">A</emphasis>
|
||
. (
|
||
<emphasis id="B91B30438D14FFF1FA96F90390BCFBB0" box="[1285,1427,1093,1116]" italics="true" pageId="2" pageNumber="67">Melinopterus</emphasis>
|
||
)
|
||
<emphasis id="B91B30438D14FFF1FF04F92095FFFB91" box="[151,208,1126,1149]" italics="true" pageId="2" pageNumber="67">stolzi</emphasis>
|
||
Reitter
|
||
</taxonomicName>
|
||
but can be easily separated from it by slender and acute apices of parameres. These two species also have widely separated ranges:
|
||
<taxonomicName id="4C6F97D28D14FFF1FE36F9CE94D4FB73" box="[421,507,1160,1183]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="stolzi">
|
||
<emphasis id="B91B30438D14FFF1FE36F9CE94D4FB73" box="[421,507,1160,1183]" italics="true" pageId="2" pageNumber="67">A. stolzi</emphasis>
|
||
</taxonomicName>
|
||
occurs in southern Europe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C375BFDA8D14FFF1FF54F9EF9119FAE8" pageId="2" pageNumber="67" type="distribution">
|
||
<paragraph id="8BD0EC518D14FFF1FF54F9EF9119FAE8" blockId="2.[151,1437,151,1284]" pageId="2" pageNumber="67">
|
||
<emphasis id="B91B30438D14FFF1FF54F9EF9476FB2D" bold="true" box="[199,345,1193,1217]" pageId="2" pageNumber="67">Distribution.</emphasis>
|
||
<taxonomicName id="4C6F97D28D14FFF1FEF3F9EC971EFB2D" box="[352,561,1194,1217]" class="Insecta" family="Scarabaeidae" genus="Aphodus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="gissaricus">
|
||
<emphasis id="B91B30438D14FFF1FEF3F9EC971EFB2D" box="[352,561,1194,1217]" italics="true" pageId="2" pageNumber="67">Aphodus gissaricus</emphasis>
|
||
</taxonomicName>
|
||
is currently known from three localities in the Pamir-Alay mountain system: two in Gissar Range (Anzob Pass and Romit) and one in Aktau Mountains.
|
||
<taxonomicName id="4C6F97D28D14FFF1FC18F98D914DFB0E" box="[907,1122,1227,1250]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="scuticollis">
|
||
<emphasis id="B91B30438D14FFF1FC18F98D914DFB0E" box="[907,1122,1227,1250]" italics="true" pageId="2" pageNumber="67">Aphodius scuticollis</emphasis>
|
||
</taxonomicName>
|
||
has a wider distribution from the Kopet-Dag Range on the southern border of
|
||
<collectingCountry id="F378ACC18D14FFF1FD09F9AB9602FAE8" box="[666,813,1261,1284]" name="Turkmenistan" pageId="2" pageNumber="67">Turkmenistan</collectingCountry>
|
||
to South Siberia (
|
||
<figureCitation id="1354F0D48D14FFF1FC7EF9AB9106FAE8" box="[1005,1065,1261,1284]" captionStart="FIGURE 9" captionStartId="2.[151,255,1988,2012]" captionTargetBox="[151,1436,1328,1964]" captionTargetId="figure@2.[151,1436,1328,1964]" captionTargetPageId="2" captionText="FIGURE 9. Distribution map of Aphodius gissaricus (▲) and A. scuticollis (■)." httpUri="https://zenodo.org/record/279714/files/figure.png" pageId="2" pageNumber="67">Fig 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF10BCD98D14FFF1FF04FA8296DFF830" box="[151,1008,1988,2012]" httpUri="https://zenodo.org/record/279714/files/figure.png" pageId="2" pageNumber="67" targetBox="[151,1436,1328,1964]" targetPageId="2">
|
||
<paragraph id="8BD0EC518D14FFF1FF04FA8296DFF830" blockId="2.[151,1008,1988,2012]" box="[151,1008,1988,2012]" pageId="2" pageNumber="67">
|
||
<emphasis id="B91B30438D14FFF1FF04FA829436F830" bold="true" box="[151,281,1988,2012]" pageId="2" pageNumber="67">FIGURE 9.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName id="4C6F97D28D14FFF1FE6AFA8397FFF830" box="[505,720,1989,2012]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="gissaricus">
|
||
<emphasis id="B91B30438D14FFF1FE6AFA8397FFF830" box="[505,720,1989,2012]" italics="true" pageId="2" pageNumber="67">Aphodius gissaricus</emphasis>
|
||
</taxonomicName>
|
||
(▲) and
|
||
<taxonomicName id="4C6F97D28D14FFF1FCA4FA8396EFF830" box="[823,960,1989,2012]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="scuticollis">
|
||
<emphasis id="B91B30438D14FFF1FCA4FA8396EFF830" box="[823,960,1989,2012]" italics="true" pageId="2" pageNumber="67">A. scuticollis</emphasis>
|
||
</taxonomicName>
|
||
(■).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C375BFDA8D15FFF0FF54FDD195D0FF3D" pageId="3" pageNumber="68" type="etymology">
|
||
<paragraph id="8BD0EC518D15FFF0FF54FDD195D0FF3D" blockId="3.[151,1435,151,209]" pageId="3" pageNumber="68">
|
||
<emphasis id="B91B30438D15FFF0FF54FDD19468FF43" bold="true" box="[199,327,151,175]" pageId="3" pageNumber="68">Etymology.</emphasis>
|
||
The name of the new species is derived from Gissar Mountains where the larger part of
|
||
<typeStatus id="54D452F38D15FFF0FB69FDDE9007FF43" box="[1274,1320,152,175]" pageId="3" pageNumber="68">type</typeStatus>
|
||
series was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |