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<document id="45F6FCC208E2822F3004A00608C76209" ID-CLB-Dataset="29403" ID-DOI="10.11646/zootaxa.4462.2.3" ID-GBIF-Dataset="b3b39f39-c823-429d-9354-81c3df099513" ID-ISSN="1175-5326" ID-Zenodo-Dep="1441531" ID-ZooBank="E31D2AE6-275E-4071-B8DB-E2060F3639E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538402810090" checkinUser="plazi" docAuthor="Pereira-Filho, José Moisés B., Saturnino, Regiane &amp; Bonaldo, Alexandre B." docDate="2018" docId="03C63D2E9E3AFFCCB8D432907FFCF871" docLanguage="en" docName="zootaxa.4462.2.3.pdf" docOrigin="Zootaxa 4462 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Attacobius demiguise Pereira-Filho &amp; Saturnino &amp; Bonaldo 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="222" masterDocId="FFFF45569E33FFC7B84330477D46FFD2" masterDocTitle="Five new species and novel descriptions of opposed sexes of four species of the spider genus Attacobius (Araneae: Corinnidae)" masterLastPageNumber="228" masterPageNumber="211" pageNumber="220" updateTime="1698561050674" updateUser="ExternalLinkService">
<mods:mods id="80838A18BB63290966B7DCBD08DABE95" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="62673C255992E2DDB5539DF36DF5A59A">Five new species and novel descriptions of opposed sexes of four species of the spider genus Attacobius (Araneae: Corinnidae)</mods:title>
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<mods:namePart id="484D5B0D2FC84918E15D0900B3FD1563">Pereira-Filho, José Moisés B.</mods:namePart>
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<mods:namePart id="74905D81E1D313AF012D63322D207D25">Saturnino, Regiane</mods:namePart>
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<mods:namePart id="77894055F35959437CADA92764850347">Bonaldo, Alexandre B.</mods:namePart>
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<mods:title id="8E339F5A91C4D94F83631545470054D7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="70857A2185FDDA1BA40404616A93A297">
<mods:date id="23BE48AF615E241AC7467D4F85A566ED">2018</mods:date>
<mods:detail id="714960FB727F1F6BE05DB755E891363D" type="pubDate">
<mods:number id="4D9AED8306BD561CC47777F6032C6CA2">2018-08-27</mods:number>
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<mods:detail id="C6BF50E5E05E70CDD80F8BF0A8F5DE12" type="volume">
<mods:number id="19EBB90041B812594AC134CDE5049049">4462</mods:number>
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<mods:classification id="4277367ABA091FB79BCDE16A5F842028">journal article</mods:classification>
<mods:identifier id="B78104564CCFF311E653E00CF8F8DCB6" type="CLB-Dataset">29403</mods:identifier>
<mods:identifier id="E0114681A37A11495C7F4B978B140449" type="DOI">10.11646/zootaxa.4462.2.3</mods:identifier>
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<mods:identifier id="36FA6203E52C768DBCEE8F77ACB463C6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C63D2E9E3AFFCCB8D432907FFCF871" ID-DOI="http://doi.org/10.5281/zenodo.5978125" ID-GBIF-Taxon="148424874" ID-Zenodo-Dep="5978125" LSID="urn:lsid:plazi:treatment:03C63D2E9E3AFFCCB8D432907FFCF871" httpUri="http://treatment.plazi.org/id/03C63D2E9E3AFFCCB8D432907FFCF871" lastPageId="11" lastPageNumber="222" pageId="9" pageNumber="220">
<subSubSection id="C375DFB39E3AFFCEB8D432907C2CFCC1" pageId="9" pageNumber="220" type="nomenclature">
<paragraph id="8BD08C389E3AFFCEB8D432907CA4FD20" blockId="9.[151,482,727,787]" box="[151,482,727,754]" pageId="9" pageNumber="220">
<heading id="D0983B549E3AFFCEB8D432907CA4FD20" bold="true" box="[151,482,727,754]" fontSize="11" level="1" pageId="9" pageNumber="220" reason="1">
<emphasis id="B91B502A9E3AFFCEB8D432907CA4FD20" bold="true" box="[151,482,727,754]" pageId="9" pageNumber="220">
<taxonomicName id="4C6FF7BB9E3AFFCEB8D432907CDEFD23" ID-CoL="JMCJ" authority="Pereira-Filho &amp; Saturnino &amp; Bonaldo, 2018" authorityName="Pereira-Filho &amp; Saturnino &amp; Bonaldo" authorityYear="2018" box="[151,408,727,753]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="220" phylum="Arthropoda" rank="species" species="demiguise" status="sp. nov.">
<emphasis id="B91B502A9E3AFFCEB8D432907CDEFD23" bold="true" box="[151,408,727,753]" italics="true" pageId="9" pageNumber="220">Attacobius demiguise</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228ED519E3AFFCEB9E3329F7CA4FD20" box="[416,482,728,754]" pageId="9" pageNumber="220" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD08C389E3AFFCEB8D432BD7C2CFCC1" blockId="9.[151,482,727,787]" box="[151,362,762,787]" pageId="9" pageNumber="220">
<figureCitation id="135490BD9E3AFFCEB8D432BD7C50FCC1" box="[151,278,762,787]" captionStart-0="FIGURES 11, 12" captionStart-1="FIGURES 13–18" captionStartId-0="10.[151,264,1739,1762]" captionStartId-1="12.[151,264,1738,1761]" captionTargetBox-0="[256,1421,512,1704]" captionTargetBox-1="[187,1423,577,1717]" captionTargetId-0="figure@10.[151,1436,445,1718]" captionTargetId-1="figure@12.[151,1436,541,1729]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionText-0="FIGURES 11, 12. Attacobius demiguise n. sp., left male palp: 11. ventral view; 12. retrolateral view. Abbreviations: AS, apical spur of VL; C, conductor; CRP, cymbial retro-dorsal process; CRT, cymbial retrolateral tubercle; CTP, cymbial transverse process; DL, dorsal lobe of RTA; dMA, distal end of median apophysis; E, embolus; mMA, median extension of median apophysis; pMA proximal end of median apophysis; VL, ventral lobe of RTA." captionText-1="FIGURES 13–18. Attacobius spp., female epigyna: 13–15. A. demiguise n. sp., 13. ventral view, 14. posterior view, 15. dorsal view; 16–18. A. uiriri Bonaldo &amp; Brescovit, 2005, 16. ventral view, 17. posterior view, 18. dorsal view. Abbreviations: CD, copulatory duct; FD, fertilization duct; PS, primary spermathecae; SS, secondary spermathecae." httpUri-0="https://zenodo.org/record/1441543/files/figure.png" httpUri-1="https://zenodo.org/record/1441545/files/figure.png" pageId="9" pageNumber="220">Figs 11–15</figureCitation>
; Map 1
</paragraph>
</subSubSection>
<subSubSection id="C375DFB39E3AFFCEB8D433057E1CFB34" pageId="9" pageNumber="220" type="materials_examined">
<paragraph id="8BD08C389E3AFFCEB8D433057E03FC71" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="220">
<materialsCitation id="3B0786659E3AFFCEB8D433057F2BFCAD" ID-GBIF-Occurrence="1913892237" collectingDate="2009-07-27" collectionCode="UFMG" collectorName="Oliveira &amp; M. D. Miranda" country="Brazil" latitude="-6.143136" location="Altamira" longLatPrecision="1" longitude="-54.33585" pageId="9" pageNumber="220" specimenCode="UFMG 5760" specimenCount="2" specimenCount-male="1" stateProvince="Para" typeStatus="holotype">
<emphasis id="B91B502A9E3AFFCEB8D433057C0CFC89" bold="true" box="[151,330,834,859]" pageId="9" pageNumber="220">
<typeStatus id="54D4329A9E3AFFCEB8D433057D94FC89" box="[151,210,834,859]" pageId="9" pageNumber="220">Type</typeStatus>
material.
</emphasis>
<specimenCount id="9D6947B19E3AFFCEB91533047C2DFC88" box="[342,363,835,858]" pageId="9" pageNumber="220" type="male">♂</specimenCount>
<typeStatus id="54D4329A9E3AFFCEB93B33057C9CFC89" box="[376,474,834,859]" pageId="9" pageNumber="220" type="holotype">holotype</typeStatus>
from
<location id="8EB0DAE39E3AFFCEBA6A33057FD6FC89" LSID="urn:lsid:plazi:treatment:03C63D2E9E3AFFCCB8D432907FFCF871:8EB0DAE39E3AFFCEBA6A33057FD6FC89" box="[553,656,834,859]" country="Brazil" latitude="-6.143136" longLatPrecision="1" longitude="-54.33585" name="Altamira" pageId="9" pageNumber="220" stateProvince="Para">Altamira</location>
,
<collectingRegion id="49AB42DA9E3AFFCEBAE333047F93FC89" box="[672,725,835,859]" country="Brazil" name="Para" pageId="9" pageNumber="220">Pará</collectingRegion>
,
<collectingCountry id="F378CCA89E3AFFCEBAA533057E6BFC89" box="[742,813,834,859]" name="Brazil" pageId="9" pageNumber="220">Brazil</collectingCountry>
,
<geoCoordinate id="EE5BEAFF9E3AFFCEBB7D33057EA7FC89" box="[830,993,834,859]" direction="south" orientation="latitude" pageId="9" pageNumber="220" precision="1" value="-6.143136">06°08'35.29''S</geoCoordinate>
,
<geoCoordinate id="EE5BEAFF9E3AFFCEBBB1330579D8FC89" box="[1010,1182,834,859]" direction="west" orientation="longitude" pageId="9" pageNumber="220" precision="1" value="-54.33585">54°20'09.06''W</geoCoordinate>
, U.
<collectorName id="269AE9EE9E3AFFCEBC9433057875FC89" box="[1239,1331,834,859]" pageId="9" pageNumber="220">Oliveira</collectorName>
&amp;
<collectorName id="269AE9EE9E3AFFCEBD2233047DB0FCAD" pageId="9" pageNumber="220">M.D. Miranda</collectorName>
leg.,
<collectingDate id="EF9553109E3AFFCEB97633207CF8FCAD" box="[309,446,871,895]" pageId="9" pageNumber="220" value="2009-07-27">
<date id="FFD1AAF89E3AFFCEB97633207CF8FCAD" box="[309,446,871,895]" pageId="9" pageNumber="220" value="2009-07-27">27.VII.2009</date>
</collectingDate>
(
<specimenCode id="DBC924439E3AFFCEB98D33207F23FCAD" box="[462,613,870,895]" collectionCode="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="9" pageNumber="220">UFMG 5760</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B0786659E3AFFCEBA3B33207E07FC71" ID-GBIF-Occurrence="1913892235" collectingDate="2009-04-25" collectionCode="UFMG" collectorName="Oliveira &amp; M. D. Miranda" country="Brazil" location="Parauapebas" longitude="-50.56736" pageId="9" pageNumber="220" specimenCode="UFMG 5744" specimenCount="1" specimenCount-female="1" stateProvince="Para" typeStatus="paratype">
<specimenCount id="9D6947B19E3AFFCEBA3B33207FCBFCAC" box="[632,653,871,894]" pageId="9" pageNumber="220" type="female">♀</specimenCount>
<typeStatus id="54D4329A9E3AFFCEBAD633207FB3FCAD" box="[661,757,871,895]" pageId="9" pageNumber="220" type="paratype">paratype</typeStatus>
from
<location id="8EB0DAE39E3AFFCEBB7F33217E8DFCAD" LSID="urn:lsid:plazi:treatment:03C63D2E9E3AFFCCB8D432907FFCF871:8EB0DAE39E3AFFCEBB7F33217E8DFCAD" box="[828,971,870,895]" country="Brazil" longitude="-50.56736" name="Parauapebas" pageId="9" pageNumber="220" stateProvince="Para">Parauapebas</location>
,
<collectingRegion id="49AB42DA9E3AFFCEBB943320794AFCAD" box="[983,1036,871,895]" country="Brazil" name="Para" pageId="9" pageNumber="220">Pará</collectingRegion>
,
<collectingCountry id="F378CCA89E3AFFCEBC543321791DFCAD" box="[1047,1115,870,895]" name="Brazil" pageId="9" pageNumber="220">Brazil</collectingCountry>
0626'12.6''S,
<geoCoordinate id="EE5BEAFF9E3AFFCEBCB9332178DEFCAD" box="[1274,1432,870,895]" direction="west" orientation="longitude" pageId="9" pageNumber="220" precision="1" value="-50.56736">50°34'02.5''W</geoCoordinate>
, U.
<collectorName id="269AE9EE9E3AFFCEB8FA33CD7C53FC71" box="[185,277,906,931]" pageId="9" pageNumber="220">Oliveira</collectorName>
&amp;
<collectorName id="269AE9EE9E3AFFCEB97A33CC7C9CFC71" box="[313,474,906,931]" pageId="9" pageNumber="220">M.D. Miranda</collectorName>
leg.,
<collectingDate id="EF9553109E3AFFCEBA5B33CD7FD2FC71" box="[536,660,906,931]" pageId="9" pageNumber="220" value="2009-04-25">
<date id="FFD1AAF89E3AFFCEBA5B33CD7FD2FC71" box="[536,660,906,931]" pageId="9" pageNumber="220" value="2009-04-25">25.IV.2009</date>
</collectingDate>
(
<specimenCode id="DBC924439E3AFFCEBAE133CC7E7FFC71" box="[674,825,906,931]" collectionCode="UFMG" country="Brazil" httpUri="http://grbio.org/cool/z0ie-wfm6" name="Universidade Federal de Minas Gerais" pageId="9" pageNumber="220">UFMG 5744</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD08C389E3AFFCEB88433E97E1CFB34" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="220">
<emphasis id="B91B502A9E3AFFCEB88433E97C40FC15" bold="true" box="[199,262,942,967]" pageId="9" pageNumber="220">Note.</emphasis>
The male
<typeStatus id="54D4329A9E3AFFCEB9C833E97CABFC15" box="[395,493,942,967]" pageId="9" pageNumber="220" type="holotype">holotype</typeStatus>
is in poor condition, with the abdomen and most legs lost. The two specimens examined were collected nearly
<quantity id="4C9721DD9E3AFFCEBA4233947F10FC39" box="[513,598,978,1003]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="220" unit="km" value="400.0">400 km</quantity>
apart and the sexes are only tentatively associated. Somatic morphology is remarkably stable across
<taxonomicName id="4C6FF7BB9E3AFFCEB9F733B17F6DFBDC" authorityName="Mello-Leitao" authorityYear="1925" box="[436,555,1014,1038]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="220" phylum="Arthropoda" rank="genus">
<emphasis id="B91B502A9E3AFFCEB9F733B17F6DFBDC" box="[436,555,1014,1038]" italics="true" pageId="9" pageNumber="220">Attacobius</emphasis>
</taxonomicName>
species, and the female
<typeStatus id="54D4329A9E3AFFCEBB7C33B07ED9FBDD" box="[831,927,1015,1039]" pageId="9" pageNumber="220" type="paratype">paratype</typeStatus>
could also be ascribable to either
<taxonomicName id="4C6FF7BB9E3AFFCEBD5C33B078DAFBDC" authority="Pereira-Filho &amp; Saturnino &amp; Bonaldo, 2018" authorityName="Pereira-Filho &amp; Saturnino &amp; Bonaldo" authorityYear="2018" box="[1311,1436,1014,1038]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="220" phylum="Arthropoda" rank="species" species="carimbo" status="sp. nov.">
<emphasis id="B91B502A9E3AFFCEBD5C33B078DAFBDC" box="[1311,1436,1014,1038]" italics="true" pageId="9" pageNumber="220">A. carimbo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228ED519E3AFFCEB8D4345D7D9FFBE1" box="[151,217,1050,1075]" pageId="9" pageNumber="220" rank="species">
<emphasis id="B91B502A9E3AFFCEB8D4345D7D9FFBE1" bold="true" box="[151,217,1050,1075]" pageId="9" pageNumber="220">n. sp.</emphasis>
</taxonomicNameLabel>
or
<taxonomicName id="4C6FF7BB9E3AFFCEB946345C7F3FFBE1" authority="Bonaldo &amp; Brescovit" authorityName="Bonaldo &amp; Brescovit" authorityYear="2005" box="[261,633,1050,1075]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="220" phylum="Arthropoda" rank="species" species="tucurui">
<emphasis id="B91B502A9E3AFFCEB946345C7C3EFBE0" box="[261,376,1051,1074]" italics="true" pageId="9" pageNumber="220">A. tucurui</emphasis>
Bonaldo &amp; Brescovit
</taxonomicName>
, the two other species known only by males that occur in South and Central
<collectingRegion id="49AB42DA9E3AFFCEB8B334787C60FB85" box="[240,294,1087,1111]" country="Brazil" name="Para" pageId="9" pageNumber="220">Pará</collectingRegion>
. Since phoretic dispersion has been reported in at least two species of the genus (
<bibRefCitation id="EFFEF1C99E3AFFCEBCF6347978D1FB85" box="[1205,1431,1086,1111]" pageId="9" pageNumber="220" refId="ref10318" refString="Ichinose, K., Rinaldi, I. &amp; Forti, L. C. (2004) Winged leaf-cutting ants on nuptial flights used as transport by Attacobius spiders for dispersal. Ecological Entomology, 29, 628 - 631. https: // doi. org / 10.1111 / j. 0307 - 6946.2004.00640. x" type="journal article">
Ichinose
<emphasis id="B91B502A9E3AFFCEBD5F34787808FB84" box="[1308,1358,1086,1110]" italics="true" pageId="9" pageNumber="220">et al</emphasis>
. 2004
</bibRefCitation>
;
<bibRefCitation id="EFFEF1C99E3AFFCEB8D434247C39FBA9" box="[151,383,1122,1147]" pageId="9" pageNumber="220" refId="ref10241" refString="Camargo, R. S., Forti, L. C., De Matos, C. A. O. &amp; Brescovit, A. D. (2015) Phoretic behaviour of Attacobius attarum (Roewer, 1935) (Araneae: Corinnidae: Corinninae) dispersion not associated with predation? Journal of Natural History, 49, 1653 - 1658. https: // doi. org / 10.1080 / 00222933.2015.1005713" type="journal article">
Camargo
<emphasis id="B91B502A9E3AFFCEB94734247C73FBA9" box="[260,309,1123,1147]" italics="true" pageId="9" pageNumber="220">et al</emphasis>
. 2015
</bibRefCitation>
), and these three species appear to inhabit the same phytophysiognomy, geographic proximity seems to be a poor criterion. This problem will only be solved when more samples from the region came to light. On the other hand, from a strict nomenclatural point of view, proposing this doubtful association is preferable than making available a fourth, possibly unnecessary specific name.
</paragraph>
</subSubSection>
<subSubSection id="C375DFB39E3AFFCEB88434B579FFFAA5" pageId="9" pageNumber="220" type="etymology">
<paragraph id="8BD08C389E3AFFCEB88434B579FFFAA5" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="220">
<emphasis id="B91B502A9E3AFFCEB88434B57C08FAD9" bold="true" box="[199,334,1266,1291]" pageId="9" pageNumber="220">Etymology.</emphasis>
The specific name is a noun in apposition taken from a character from Harry Potter's Universe by J.K. Rowling. The Demiguise is described in the Magizoology compendium “Fantastic Beasts and Where to Find Them” as being capable of making itself invisible when in danger. The name alludes to the behaviour of some
<taxonomicName id="4C6FF7BB9E3AFFCEB8D435197C48FAA4" authorityName="Mello-Leitao" authorityYear="1925" box="[151,270,1374,1398]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="220" phylum="Arthropoda" rank="genus">
<emphasis id="B91B502A9E3AFFCEB8D435197C48FAA4" box="[151,270,1374,1398]" italics="true" pageId="9" pageNumber="220">Attacobius</emphasis>
</taxonomicName>
species, which appear to become invisible to ants while climbing onto their bodies.
</paragraph>
</subSubSection>
<subSubSection id="C375DFB39E3AFFCEB88435C57C8BF99D" pageId="9" pageNumber="220" type="diagnosis">
<paragraph id="8BD08C389E3AFFCEB88435C57C8BF99D" blockId="9.[151,1437,834,2011]" pageId="9" pageNumber="220">
<emphasis id="B91B502A9E3AFFCEB88435C57C07FA49" bold="true" box="[199,321,1410,1435]" pageId="9" pageNumber="220">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C6FF7BB9E3AFFCEB9F335C47FE6FA49" authority="Pereira-Filho &amp; Saturnino &amp; Bonaldo, 2018" authorityName="Pereira-Filho &amp; Saturnino &amp; Bonaldo" authorityYear="2018" box="[432,672,1411,1435]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="220" phylum="Arthropoda" rank="species" species="demiguise" status="sp. nov.">
<emphasis id="B91B502A9E3AFFCEB9F335C47FE6FA49" box="[432,672,1411,1435]" italics="true" pageId="9" pageNumber="220">Attacobius demiguise</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228ED519E3AFFCEBAE535C57FA2FA49" box="[678,740,1410,1435]" pageId="9" pageNumber="220" rank="species">
<emphasis id="B91B502A9E3AFFCEBAE535C57FA2FA49" bold="true" box="[678,740,1410,1435]" pageId="9" pageNumber="220">n. sp.</emphasis>
</taxonomicNameLabel>
share with those of
<taxonomicName id="4C6FF7BB9E3AFFCEBB8735C4797EFA49" authority="Pereira-Filho &amp; Saturnino &amp; Bonaldo, 2018" authorityName="Pereira-Filho &amp; Saturnino &amp; Bonaldo" authorityYear="2018" box="[964,1080,1411,1435]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="220" phylum="Arthropoda" rank="species" species="thalitae" status="sp. nov.">
<emphasis id="B91B502A9E3AFFCEBB8735C4797EFA49" box="[964,1080,1411,1435]" italics="true" pageId="9" pageNumber="220">A. thalitae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228ED519E3AFFCEBC7D35C5793AFA49" box="[1086,1148,1410,1435]" pageId="9" pageNumber="220" rank="species">
<emphasis id="B91B502A9E3AFFCEBC7D35C5793AFA49" bold="true" box="[1086,1148,1410,1435]" pageId="9" pageNumber="220">n. sp.</emphasis>
</taxonomicNameLabel>
the short proximal end of median apophysis and the absence of the tegular retrolateral groove (
<figureCitation id="135490BD9E3AFFCEBBE035E1795EFA6D" box="[931,1048,1446,1471]" captionStart="FIGURES 9, 10" captionStartId="8.[151,264,1546,1569]" captionTargetBox="[162,1420,263,1508]" captionTargetId="figure@8.[151,1436,193,1525]" captionTargetPageId="8" captionText="FIGURES 9, 10. Attacobius thalitae n. sp., left male palp: 9. ventral view; 10. retrolateral view. Abbreviations: C, conductor; CPP, Cymbial prolateral process; CRP, cymbial retro-dorsal process; CTP, cymbial transverse process; dMA, distal end of median apophysis; E, embolus; mMA, median extension of median apophysis; pMA proximal end of median apophysis; PTK, prolateral ventral tegular keel." httpUri="https://zenodo.org/record/1441541/files/figure.png" pageId="9" pageNumber="220">Figs 9, 10</figureCitation>
). They are promptly distinguished from males of that species by the large, cup-shaped, cymbial retro-dorsal process and by the absence of tegular keels (
<figureCitation id="135490BD9E3AFFCEB8A235A97C21F9D5" box="[225,359,1518,1543]" captionStart="FIGURES 11, 12" captionStartId="10.[151,264,1739,1762]" captionTargetBox="[256,1421,512,1704]" captionTargetId="figure@10.[151,1436,445,1718]" captionTargetPageId="10" captionText="FIGURES 11, 12. Attacobius demiguise n. sp., left male palp: 11. ventral view; 12. retrolateral view. Abbreviations: AS, apical spur of VL; C, conductor; CRP, cymbial retro-dorsal process; CRT, cymbial retrolateral tubercle; CTP, cymbial transverse process; DL, dorsal lobe of RTA; dMA, distal end of median apophysis; E, embolus; mMA, median extension of median apophysis; pMA proximal end of median apophysis; VL, ventral lobe of RTA." httpUri="https://zenodo.org/record/1441543/files/figure.png" pageId="9" pageNumber="220">Figs 11, 12</figureCitation>
). Females are similar to those of
<taxonomicName id="4C6FF7BB9E3AFFCEBAA835A87E0EF9D4" authorityName="Bonaldo &amp; Brescovit" authorityYear="2005" box="[747,840,1519,1542]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="220" phylum="Arthropoda" rank="species" species="uiriri">
<emphasis id="B91B502A9E3AFFCEBAA835A87E0EF9D4" box="[747,840,1519,1542]" italics="true" pageId="9" pageNumber="220">A. uiriri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6FF7BB9E3AFFCEBBC035A87EAFF9D4" authorityName="Bonaldo &amp; Brescovit" authorityYear="2005" box="[899,1001,1518,1542]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="220" phylum="Arthropoda" rank="species" species="blakei">
<emphasis id="B91B502A9E3AFFCEBBC035A87EAFF9D4" box="[899,1001,1518,1542]" italics="true" pageId="9" pageNumber="220">A. blakei</emphasis>
</taxonomicName>
(
<figureCitation id="135490BD9E3AFFCEBBB935A979C6F9D5" box="[1018,1152,1518,1543]" captionStart="FIGURES 13–18" captionStartId="12.[151,264,1738,1761]" captionTargetBox="[187,1423,577,1717]" captionTargetId="figure@12.[151,1436,541,1729]" captionTargetPageId="12" captionText="FIGURES 13–18. Attacobius spp., female epigyna: 13–15. A. demiguise n. sp., 13. ventral view, 14. posterior view, 15. dorsal view; 16–18. A. uiriri Bonaldo &amp; Brescovit, 2005, 16. ventral view, 17. posterior view, 18. dorsal view. Abbreviations: CD, copulatory duct; FD, fertilization duct; PS, primary spermathecae; SS, secondary spermathecae." httpUri="https://zenodo.org/record/1441545/files/figure.png" pageId="9" pageNumber="220">Figs 16, 18</figureCitation>
,
<figureCitation id="135490BD9E3AFFCEBCCD35A87991F9D5" box="[1166,1239,1519,1543]" captionStart="FIGURES 19–24" captionStartId="14.[151,264,1601,1624]" captionTargetBox="[187,1436,373,1568]" captionTargetId="figure@14.[151,1436,373,1580]" captionTargetPageId="14" captionText="FIGURES 19–24. Attacobius spp., female epigyna: 19–21. A. blakei Bonaldo &amp; Brescovit, 2005, 19. ventral view, 20. posterior view, 21. dorsal view; 22–24. A. tremembe n. sp., 22. ventral view, 23. posterior view, 24. dorsal view. Abbreviations: CD, copulatory duct; EVP, epigynal ventral plate; FD, fertilization duct; PS, primary spermathecae; SS, secondary spermathecae." httpUri="https://zenodo.org/record/1441547/files/figure.png" pageId="9" pageNumber="220">19, 21</figureCitation>
) by the medially interrupted epigynal ventral plate, but can be recognized by the presence of small, globular secondary spermathecae (
<figureCitation id="135490BD9E3AFFCEB97D36717C86F99D" box="[318,448,1590,1615]" captionStart="FIGURES 13–18" captionStartId="12.[151,264,1738,1761]" captionTargetBox="[187,1423,577,1717]" captionTargetId="figure@12.[151,1436,541,1729]" captionTargetPageId="12" captionText="FIGURES 13–18. Attacobius spp., female epigyna: 13–15. A. demiguise n. sp., 13. ventral view, 14. posterior view, 15. dorsal view; 16–18. A. uiriri Bonaldo &amp; Brescovit, 2005, 16. ventral view, 17. posterior view, 18. dorsal view. Abbreviations: CD, copulatory duct; FD, fertilization duct; PS, primary spermathecae; SS, secondary spermathecae." httpUri="https://zenodo.org/record/1441545/files/figure.png" pageId="9" pageNumber="220">Figs 13–15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C375DFB39E3AFFCCB884361D7C37FE27" lastPageId="11" lastPageNumber="222" pageId="9" pageNumber="220" type="description">
<paragraph id="8BD08C389E3AFFCDB884361D7ECBFE5A" blockId="9.[151,1437,834,2011]" lastBlockId="10.[151,1436,151,392]" lastPageId="10" lastPageNumber="221" pageId="9" pageNumber="220">
<emphasis id="B91B502A9E3AFFCEB884361D7C1FF9A1" bold="true" box="[199,345,1626,1651]" pageId="9" pageNumber="220">Description.</emphasis>
<emphasis id="B91B502A9E3AFFCEB921361C7CDAF9A1" box="[354,412,1627,1651]" italics="true" pageId="9" pageNumber="220">Male</emphasis>
(
<typeStatus id="54D4329A9E3AFFCEB9ED361D7F52F9A1" box="[430,532,1626,1651]" pageId="9" pageNumber="220" type="holotype">holotype</typeStatus>
, UFMG 5760). Carapace pale yellow, cephalic region slightly darker, anterior median eye area with black spot divided by longitudinal gap; chelicerae pale yellow with fangs brown; endites and labium pale yellow, with white apex; sternum pale yellow with brown margins; legs: femur and patella yellow, tibiae, metatarsus and tarsus dark yellow; abdomen lost. Total length (?). Carapace 1.58 long, 1.28 wide, 0.30 high. Clypeus 0.12 high. Eye diameters and interdistances: AME 0.10, ALE 0.09, PME 0.10, PLE 0.09; AME–AME 0.07, AME–ALE 0.02, PME–PME 0.13, PME–PLE 0.04, ALE–PLE 0.01. MOQ length: 0.22, front width: 0.25, back width: 0.26. Chelicerae 0.55 long, with 3 promarginal and 2 retromarginal teeth. Sternum 0.87 long, 0.91 wide. Abdomen lost. Leg measurements: I: femur 1.32/ patella 0.54/ tibia 1.19/ metatarsus 1.24/ tarsus 0.90/ total 5.19. II: 1.16/ 0.53/ 1.12/ 1.09/ 0.87/ 4.77. III: lost. IV: 1.40/ 0.52/ 1.19/ 1.39/ lost/ (?). Leg spination: I: femur d1- 1-0, p0, r0, v0; tibia d0, p0, r0,
<date id="FFD1AAF89E3AFFCEB9BE37D87F75F865" box="[509,563,1951,1975]" pageId="9" pageNumber="220">v2-2</date>
-2; metatarsus d0, p0, r0,
<date id="FFD1AAF89E3AFFCEBB1137D87ECEF865" box="[850,904,1951,1975]" pageId="9" pageNumber="220">v2-2</date>
-0. II: femur d1-1-0, p0, r0, v0; tibia d0, p0, r0,
<date id="FFD1AAF89E3AFFCEB8D437847D8BF809" box="[151,205,1987,2011]" pageId="9" pageNumber="220">v2-2</date>
-2; metatarsus d0, p0, r0, v0. III: lost. IV: femur d0, p0, r0-0-1, v0; tibia d0, p0, r0, v0; metatarsus lost. Palp: RTA with two lobes, ventral lobe small; apical spur sclerotized, with narrow base arising sub-apically from ventral lobe, dorsal lobe sub-rectangular, oriented dorsally, with small apical digitiform process; cymbial prolateral process separated from cymbial transversal process; cymbial transversal process small, nearly straight, without basal prong; cymbial retrolateral tubercle large, conical, pointed ventro-distally; cymbial retro-dorsal process large, cupshaped; both prolateral and ventral tegular keels absent; tegular retrolateral groove absent; distal extension of median apophysis blunt, not projected distally; median extension of median apophysis smooth, composed of two pieces, proximal extension of median apophysis short (
<figureCitation id="135490BD9E39FFCDBABC31287E39FE5A" box="[767,895,367,392]" captionStart="FIGURES 11, 12" captionStartId="10.[151,264,1739,1762]" captionTargetBox="[256,1421,512,1704]" captionTargetId="figure@10.[151,1436,445,1718]" captionTargetPageId="10" captionText="FIGURES 11, 12. Attacobius demiguise n. sp., left male palp: 11. ventral view; 12. retrolateral view. Abbreviations: AS, apical spur of VL; C, conductor; CRP, cymbial retro-dorsal process; CRT, cymbial retrolateral tubercle; CTP, cymbial transverse process; DL, dorsal lobe of RTA; dMA, distal end of median apophysis; E, embolus; mMA, median extension of median apophysis; pMA proximal end of median apophysis; VL, ventral lobe of RTA." httpUri="https://zenodo.org/record/1441543/files/figure.png" pageId="10" pageNumber="221">Figs 11, 12</figureCitation>
).
</paragraph>
<caption id="DF10DCB09E39FFCDB8D4368C7EE9F8EF" httpUri="https://zenodo.org/record/1441543/files/figure.png" pageId="10" pageNumber="221" startId="10.[151,264,1739,1762]" targetBox="[256,1421,512,1704]" targetPageId="10">
<paragraph id="8BD08C389E39FFCDB8D4368C7EE9F8EF" blockId="10.[151,1436,1739,1853]" pageId="10" pageNumber="221">
<emphasis id="B91B502A9E39FFCDB8D4368C7C17F932" bold="true" box="[151,337,1739,1762]" pageId="10" pageNumber="221">FIGURES 11, 12.</emphasis>
<taxonomicName id="4C6FF7BB9E39FFCDB914368C7F76F932" authority="Pereira-Filho &amp; Saturnino &amp; Bonaldo, 2018" authorityName="Pereira-Filho &amp; Saturnino &amp; Bonaldo" authorityYear="2018" box="[343,560,1739,1760]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="221" phylum="Arthropoda" rank="species" species="demiguise" status="sp. nov.">
<emphasis id="B91B502A9E39FFCDB914368C7F76F932" box="[343,560,1739,1760]" italics="true" pageId="10" pageNumber="221">Attacobius demiguise</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A228ED519E39FFCDBA76368B7F28F933" box="[565,622,1740,1761]" pageId="10" pageNumber="221" rank="species">
<emphasis id="B91B502A9E39FFCDBA76368B7F28F933" bold="true" box="[565,622,1740,1761]" pageId="10" pageNumber="221">n. sp.</emphasis>
</taxonomicNameLabel>
, left male palp: 11. ventral view; 12. retrolateral view. Abbreviations: AS, apical spur of VL; C, conductor; CRP, cymbial retro-dorsal process; CRT, cymbial retrolateral tubercle; CTP, cymbial transverse process; DL, dorsal lobe of RTA; dMA, distal end of median apophysis; E, embolus; mMA, median extension of median apophysis; pMA proximal end of median apophysis; VL, ventral lobe of RTA.
</paragraph>
</caption>
<paragraph id="8BD08C389E39FFCDB884372D78DBF83D" blockId="10.[151,1437,1898,2031]" pageId="10" pageNumber="221">
<emphasis id="B91B502A9E39FFCDB884372D7C5AF850" box="[199,284,1898,1922]" italics="true" pageId="10" pageNumber="221">Female</emphasis>
(
<typeStatus id="54D4329A9E39FFCDB968372C7CD4F851" box="[299,402,1899,1923]" pageId="10" pageNumber="221" type="paratype">Paratype</typeStatus>
, UFMG 5744). Colouration as in male, except carapace with cephalic region dark yellow; abdomen pale grey. Total length 4.09. Carapace 1.76 long, 1.49 wide, 0.48 high. Clypeus 0.1 high. Eye diameters and interdistances: AME 0.11, ALE 0.11, PME 0.09, PLE 0.11; AME–AME 0.09, AME–ALE contiguous, PME– PME 0.15, PME–PLE 0.06, ALE–PLE contiguous. MOQ length: 0.25, front width: 0.27, back width: 0.30.
</paragraph>
<paragraph id="8BD08C389E38FFCCB8D430D07C37FE27" blockId="11.[151,1437,151,573]" pageId="11" pageNumber="222">
Chelicerae 0.64 long, with 3 promarginal and 2 retromarginal teeth. Sternum 0.94 long, 0.93 wide. Leg measurements: I: femur 1.64/ patella 0.63/ tibia 1.47/ metatarsus 1.58/ tarsus 1.10/ total 6.42. II: 1.54/ 0.59/ 1.32/ 1.50/ lost / (?). III: 1.50/ 0.56/ 1.26/ 1.57/ 1.16/ 6.05. IV: 1.73/ 0.60/ 1.52/ 1.89/ lost / (?). Leg spination: I: femur d1-1-0, p0-0-1, r0, v0; tibia d0, p0, r0,
<date id="FFD1AAF89E38FFCCBA0C31437FC3FECE" box="[591,645,260,284]" pageId="11" pageNumber="222">v3-2</date>
-2; metatarsus d0, p0, r0,
<date id="FFD1AAF89E38FFCCBBE731437E9CFECE" box="[932,986,260,284]" pageId="11" pageNumber="222">v2-2</date>
-0. II: femur d1-1-0, p0, r0, v0; tibia d0, p0, r0,
<date id="FFD1AAF89E38FFCCB8A6316F7C5DFE92" box="[229,283,296,320]" pageId="11" pageNumber="222">v3-2</date>
-2; metatarsus d0, p0, r0,
<date id="FFD1AAF89E38FFCCBA76316F7F2DFE92" box="[565,619,296,320]" pageId="11" pageNumber="222">v2-2</date>
-0. III: femur d1-1-0, p0, r0, v0; tibia d0, p0, r0, v0; metatarsus d0, p0, r0, v0. IV: femur d1-1-0, p0, r0-0-1, v0; tibia d0, p0, r0, v0; metatarsus d0, p0, r0, v0. Epigynum: ventral surface medially excavated, ventral plate medially interrupted; posterior transverse septum present; dorsal extension of epigynal posterior plate reduced; copulatory openings ventral, medially placed; copulatory ducts translucent; spermathecae separated from each other; primary spermathecae large, sub-quadrangular; secondary spermathecae small (
<figureCitation id="135490BD9E38FFCCB8A1319B7C22FE27" box="[226,356,476,501]" captionStart="FIGURES 13–18" captionStartId="12.[151,264,1738,1761]" captionTargetBox="[187,1423,577,1717]" captionTargetId="figure@12.[151,1436,541,1729]" captionTargetPageId="12" captionText="FIGURES 13–18. Attacobius spp., female epigyna: 13–15. A. demiguise n. sp., 13. ventral view, 14. posterior view, 15. dorsal view; 16–18. A. uiriri Bonaldo &amp; Brescovit, 2005, 16. ventral view, 17. posterior view, 18. dorsal view. Abbreviations: CD, copulatory duct; FD, fertilization duct; PS, primary spermathecae; SS, secondary spermathecae." httpUri="https://zenodo.org/record/1441545/files/figure.png" pageId="11" pageNumber="222">Figs 13–15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C375DFB39E38FFCCB88431B879F6FDCA" box="[199,1200,511,536]" pageId="11" pageNumber="222" type="distribution">
<paragraph id="8BD08C389E38FFCCB88431B879F6FDCA" blockId="11.[151,1437,151,573]" box="[199,1200,511,536]" pageId="11" pageNumber="222">
<emphasis id="B91B502A9E38FFCCB88431B87C26FDCA" bold="true" box="[199,352,511,536]" pageId="11" pageNumber="222">Distribution.</emphasis>
Known only from Altamira and Parauapebas, state of
<collectingRegion id="49AB42DA9E38FFCCBB8032477EBEFDCA" box="[963,1016,512,536]" country="Brazil" name="Para" pageId="11" pageNumber="222">Pará</collectingRegion>
,
<collectingCountry id="F378CCA89E38FFCCBC4031B87901FDCA" box="[1027,1095,511,536]" name="Brazil" pageId="11" pageNumber="222">Brazil</collectingCountry>
(Map 1).
</paragraph>
</subSubSection>
<subSubSection id="C375DFB39E38FFCCB88432637FFCF871" pageId="11" pageNumber="222" type="materials_examined">
<paragraph id="8BD08C389E38FFCCB88432637F24FDEF" blockId="11.[151,1437,151,573]" box="[199,610,548,573]" pageId="11" pageNumber="222">
<emphasis id="B91B502A9E38FFCCB88432637CF6FDEF" bold="true" box="[199,432,548,573]" pageId="11" pageNumber="222">Material examined.</emphasis>
Only the types.
</paragraph>
<paragraph id="8BD08C389E38FFCCB8D437C97FFCF871" blockId="11.[151,698,1933,1955]" box="[151,698,1933,1955]" pageId="11" pageNumber="222">
<emphasis id="B91B502A9E38FFCCB8D437C97DACF871" bold="true" box="[151,234,1934,1955]" pageId="11" pageNumber="222">MAP 1.</emphasis>
Distribution records of species of
<taxonomicName id="4C6FF7BB9E38FFCCBA0B37C97FF2F871" authorityName="Mello-Leitao" authorityYear="1925" box="[584,692,1934,1955]" class="Arachnida" family="Corinnidae" genus="Attacobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B91B502A9E38FFCCBA0B37C97FF2F871" box="[584,692,1934,1955]" italics="true" pageId="11" pageNumber="222">Attacobius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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