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<document id="733B74AA08972DB73A13514658CF82AA" ID-DOI="10.11646/zootaxa.5026.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10100344" ID-ZooBank="07E0C922-6B0F-4916-85C2-AD95146A8F1E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629979906524" checkinUser="plazi" docAuthor="Neita-Moreno, Jhon C." docDate="2021" docId="03932D70B629C37CFF7B097EA13DD86F" docLanguage="en" docName="zootaxa.5026.1.1.pdf" docOrigin="Zootaxa 5026 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclocephala brevipennis Endrodi 1964" docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="FFAA5508B623C377FFEC0C06A02ADC12" masterDocTitle="A review of the black species of Cyclocephala Dejean (Coleoptera: Scarabaeidae Dynastinae) from Colombia" masterLastPageNumber="58" masterPageNumber="1" pageNumber="11" updateTime="1699603307102" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="031910F18F13291B2659BCD99B6A0AB5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B85C3ADDC24F5CAAAFEEC526E97326B7">
<mods:title id="D83A94B49685BD595CC7E8697A9A2556">A review of the black species of Cyclocephala Dejean (Coleoptera: Scarabaeidae Dynastinae) from Colombia</mods:title>
</mods:titleInfo>
<mods:name id="E0CB1F7DC34BCCFFD85B1FE148E4FBFB" type="personal">
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<mods:roleTerm id="0E6D0EF36FC090F37ADD22516E60C0D1">Author</mods:roleTerm>
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<mods:namePart id="F56CFD725DBEA8D775FB6A699994C1F3">Neita-Moreno, Jhon C.</mods:namePart>
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<mods:title id="42300EB0D2DC3F60382363C261E374C0">Zootaxa</mods:title>
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<mods:date id="04C8209EFE84FFD8F066B1004EF11067">2021</mods:date>
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<mods:number id="D5FCE2D05E7CD938828CCDDED99CD5C2">2021-08-24</mods:number>
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<mods:number id="13FC3E6A12AB241770BE375D6F219FA5">5026</mods:number>
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<mods:classification id="9482D003D4C3E6005DD538C917BCE0B6">journal article</mods:classification>
<mods:identifier id="233B77045F71C50E931F1922B2BA5DB5" type="DOI">10.11646/zootaxa.5026.1.1</mods:identifier>
<mods:identifier id="82EDD0274E4B527EB3766773B7F007B8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C88A686FD161BCCB92FD3C5F44D38324" type="Zenodo-Dep">10100344</mods:identifier>
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<treatment id="03932D70B629C37CFF7B097EA13DD86F" LSID="urn:lsid:plazi:treatment:03932D70B629C37CFF7B097EA13DD86F" httpUri="http://treatment.plazi.org/id/03932D70B629C37CFF7B097EA13DD86F" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="C320CFEDB629C37DFF7B097EA2ADD980" box="[151,647,1400,1427]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8B859C66B629C37DFF7B097EA2ADD980" blockId="10.[151,1259,1400,1463]" box="[151,647,1400,1427]" pageId="10" pageNumber="11">
<heading id="D0CD2B0AB629C37DFF7B097EA2ADD980" bold="true" box="[151,647,1400,1427]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="4C3AE7E5B629C37DFF7B097EA2ADD980" authority="Endrodi, 1964" authorityName="Endrodi" authorityYear="1964" box="[151,647,1400,1427]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="B94E4074B629C37DFF7B097EA2ADD980" bold="true" box="[151,647,1400,1427]" pageId="10" pageNumber="11">
<emphasis id="B94E4074B629C37DFF7B097EA1E7D981" bold="true" box="[151,461,1400,1427]" italics="true" pageId="10" pageNumber="11">Cyclocephala brevipennis</emphasis>
<bibRefCitation id="EFABE197B629C37DFE39097EA2ADD980" author="Endrodi, S." box="[469,647,1400,1426]" pageId="10" pageNumber="11" pagination="433 - 470" refId="ref23004" refString="Endrodi, S. (1964) Eine Reihe von neuen Cyclocephala - Arten (Col., Melolonthidae, Dynastinae). Folia Entomologica Hungarica, 17, 433 - 470." type="journal article" year="1964">Endrödi, 1964</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB629C37DFF7B099AA4C1D9A5" box="[151,1259,1436,1463]" pageId="10" pageNumber="11" type="description">
<paragraph id="8B859C66B629C37DFF7B099AA4C1D9A5" blockId="10.[151,1259,1400,1463]" box="[151,1259,1436,1463]" pageId="10" pageNumber="11">
(
<figureCitation id="130180E3B629C37DFF73099AA10FD9A5" box="[159,293,1436,1463]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[308,1279,181,1919]" captionTargetId="figure-15@13.[308,1279,181,1919]" captionTargetPageId="13" captionText="FIGURE 2A–D. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis male; B, Cyclocephala brevipennis female; C, Cyclocephala fasciolata male; D, Cyclocephala santandereana (holotype). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100348" httpUri="https://zenodo.org/record/10100348/files/figure.png" pageId="10" pageNumber="11">Figs. 2A–B</figureCitation>
;
<figureCitation id="130180E3B629C37DFEDC099BA15CD9A5" box="[304,374,1437,1463]" captionStart="FIGURE 9" captionStartId="20.[151,250,1912,1936]" captionTargetBox="[151,1429,188,1878]" captionTargetId="figure-29@20.[151,1436,181,1888]" captionTargetPageId="20" captionText="FIGURE 9A–D. Adult mandibles and maxillae. A, Cyclocephala brevipennis mandible (ventral view); B, Cyclocephala brevipennis maxilla (ventral view); C, Cyclocephala fasciolata mandible (dorsal view); D, Cyclocephala fasciolata maxilla (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.10100366" httpUri="https://zenodo.org/record/10100366/files/figure.png" pageId="10" pageNumber="11">9A–B</figureCitation>
;
<figureCitation id="130180E3B629C37DFE6E099BA19FD9A5" box="[386,437,1437,1463]" captionStart="FIGURE 12" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[264,1334,202,1911]" captionTargetId="figure-15@23.[253,1334,181,1919]" captionTargetPageId="23" captionText="FIGURE 12A–F. Adult labia. A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala fasciolata; D, Cyclocephala brevipennis; E, Cyclocephala ligyrina; F, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100372" httpUri="https://zenodo.org/record/10100372/files/figure.png" pageId="10" pageNumber="11">12D</figureCitation>
;
<figureCitation id="130180E3B629C37DFE2D099BA225D9A5" box="[449,527,1437,1463]" captionStart="FIGURE 13" captionStartId="24.[151,250,1821,1845]" captionTargetBox="[153,1423,181,1788]" captionTargetId="figure-29@24.[151,1436,181,1798]" captionTargetPageId="24" captionText="FIGURE 13A–F. Adult epipharynxes. A, Cyclocephala marqueti ventral view; B, Cyclocephala marqueti dorsal view; C, Cyclocephala proxima ventral view; D, Cyclocephala proxima dorsal view; E, Cyclocephala brevipennis ventral view; F, Cyclocephala brevipennis dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10100377" httpUri="https://zenodo.org/record/10100377/files/figure.png" pageId="10" pageNumber="11">13E–F</figureCitation>
;
<figureCitation id="130180E3B629C37DFDF7099AA266D9A5" box="[539,588,1436,1463]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[219,1372,189,1907]" captionTargetId="figure-29@26.[206,1381,181,1919]" captionTargetPageId="26" captionText="FIGURE 15A–J. Adult male large claw (caudal view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100382" httpUri="https://zenodo.org/record/10100382/files/figure.png" pageId="10" pageNumber="11">15C</figureCitation>
;
<figureCitation id="130180E3B629C37DFDB4099AA2A0D9A4" box="[600,650,1436,1462]" captionStart="FIGURE 16" captionStartId="27.[151,250,1676,1700]" captionTargetBox="[167,1426,192,1639]" captionTargetId="figure-15@27.[151,1436,181,1652]" captionTargetPageId="27" captionText="FIGURE 16A–J. Adult male large claw (lateral view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100384" httpUri="https://zenodo.org/record/10100384/files/figure.png" pageId="10" pageNumber="11">16C</figureCitation>
;
<figureCitation id="130180E3B629C37DFD79099BA2EDD9A5" box="[661,711,1437,1463]" captionStart="FIGURE 17" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[178,1404,181,1911]" captionTargetId="figure-29@28.[169,1418,181,1919]" captionTargetPageId="28" captionText="FIGURE 17A–J. Protarsomere 5 (caudal view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100386" httpUri="https://zenodo.org/record/10100386/files/figure.png" pageId="10" pageNumber="11">17C</figureCitation>
;
<figureCitation id="130180E3B629C37DFD3F099BA30FD9A5" box="[723,805,1437,1463]" captionStart="FIGURE 18" captionStartId="29.[151,250,1846,1870]" captionTargetBox="[161,1428,191,1813]" captionTargetId="figure-15@29.[151,1436,181,1822]" captionTargetPageId="29" captionText="FIGURE 18A–M. Protibiae (dorsal view). A, Cyclocephala marqueti (male); B, Cyclocephala proxima (male); C, Cyclocephala proxima (female); D, Cyclocephala brevipennis (male); E, Cyclocephala brevipennis (female); F, Cyclocephala fasciolata; G, Cyclocephala santandereana (holotype); H, Cyclocephala dalensi; I, Cyclocephala kaszabi; J, Cyclocephala ligyrina; K, Cyclocephala rufa; L, Cyclocephala arhuacana (holotype); M, Cyclocephala arhuacana (allotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100388" httpUri="https://zenodo.org/record/10100388/files/figure.png" pageId="10" pageNumber="11">18D, E</figureCitation>
;
<figureCitation id="130180E3B629C37DFCDD099BA34ED9A5" box="[817,868,1437,1463]" captionStart="FIGURE 20" captionStartId="31.[151,250,1913,1937]" captionTargetBox="[196,1389,161,1875]" captionTargetId="figure-15@31.[185,1402,151,1890]" captionTargetPageId="31" captionText="FIGURE 20A–G. Female epipleura (ventral view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100394" httpUri="https://zenodo.org/record/10100394/files/figure.png" pageId="10" pageNumber="11">20D</figureCitation>
;
<figureCitation id="130180E3B629C37DFC9C099BA38ED9A5" box="[880,932,1437,1463]" captionStart="FIGURE 22" captionStartId="33.[151,250,1736,1760]" captionTargetBox="[155,1423,191,1701]" captionTargetId="figure-15@33.[151,1436,181,1712]" captionTargetPageId="33" captionText="FIGURE 22A–D. Wing. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis; B, C, D, Cyclocephala fasciolata." figureDoi="http://doi.org/10.5281/zenodo.10100398" httpUri="https://zenodo.org/record/10100398/files/figure.png" pageId="10" pageNumber="11">22A</figureCitation>
;
<figureCitation id="130180E3B629C37DFC43099BA3C8D9A5" box="[943,994,1437,1463]" captionStart="FIGURE 24" captionStartId="35.[151,250,1840,1864]" captionTargetBox="[152,1423,181,1807]" captionTargetId="figure-15@35.[151,1436,181,1816]" captionTargetPageId="35" captionText="FIGURE 24A–H. Pygidia (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina." figureDoi="http://doi.org/10.5281/zenodo.10100402" httpUri="https://zenodo.org/record/10100402/files/figure.png" pageId="10" pageNumber="11">24D</figureCitation>
;
<figureCitation id="130180E3B629C37DFC02099AA40AD9A5" box="[1006,1056,1436,1463]" captionStart="FIGURE 25" captionStartId="39.[151,250,1962,1986]" captionTargetBox="[154,1436,777,1932]" captionTargetId="figure-280@39.[151,1436,777,1939]" captionTargetPageId="39" captionText="FIGURE 25A–D. Pygidia and heads (dorsal view). A, Cyclocephala rufa pygidium; B, Cyclocephala arhuacana (holotype) pygidium; C, Cyclocephala brevipennis head (dorsal view); D, Cyclocephala santandereana (holotype) head (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10100404" httpUri="https://zenodo.org/record/10100404/files/figure.png" pageId="10" pageNumber="11">25C</figureCitation>
;
<figureCitation id="130180E3B629C37DFBC0099BA455D9A5" box="[1068,1151,1437,1463]" captionStart="FIGURE 27" captionStartId="41.[151,250,1885,1909]" captionTargetBox="[205,1382,190,1855]" captionTargetId="figure-15@41.[193,1395,181,1861]" captionTargetPageId="41" captionText="FIGURE 27A–F. Parameres and phallobases, Cyclocephala fasciolata species group. A, Cyclocephala fasciolata parameres, caudal view; B, Cyclocephala fasciolata phallobase, lateral view; C, Cyclocephala brevipennis parameres, caudal view; D, Cyclocephala brevipennis phallobase, lateral view; E, Cyclocephala santandereana parameres, caudal view; F, Cyclocephala santandereana phallobase, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10100409" httpUri="https://zenodo.org/record/10100409/files/figure.png" pageId="10" pageNumber="11">27C, D</figureCitation>
; 32F; 39)
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB629C37DFF7B09E2A167DAE9" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8B859C66B629C37DFF7B09E2A167DAE9" blockId="10.[151,1437,1508,2039]" pageId="10" pageNumber="11">
<emphasis id="B94E4074B629C37DFF7B09E2A120D9EC" bold="true" box="[151,266,1508,1534]" pageId="10" pageNumber="11">Diagnosis</emphasis>
.
<taxonomicName id="4C3AE7E5B629C37DFEF609E2A215D9EC" authorityName="Endrodi" authorityYear="1964" box="[282,575,1508,1534]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="B94E4074B629C37DFEF609E2A215D9EC" box="[282,575,1508,1534]" italics="true" pageId="10" pageNumber="11">Cyclocephala brevipennis</emphasis>
</taxonomicName>
can be distinguished by the glabrous frons and clypeus (
<figureCitation id="130180E3B629C37DFB2209E2A51ED9ED" box="[1230,1332,1508,1535]" captionStart="FIGURE 25" captionStartId="39.[151,250,1962,1986]" captionTargetBox="[154,1436,777,1932]" captionTargetId="figure-280@39.[151,1436,777,1939]" captionTargetPageId="39" captionText="FIGURE 25A–D. Pygidia and heads (dorsal view). A, Cyclocephala rufa pygidium; B, Cyclocephala arhuacana (holotype) pygidium; C, Cyclocephala brevipennis head (dorsal view); D, Cyclocephala santandereana (holotype) head (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10100404" httpUri="https://zenodo.org/record/10100404/files/figure.png" pageId="10" pageNumber="11">Fig. 25C</figureCitation>
) (similar to
<taxonomicName id="4C3AE7E5B629C37DFF590A0FA16EDA30" authorityName="Bates" authorityYear="1888" box="[181,324,1544,1571]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciolata">
<emphasis id="B94E4074B629C37DFF590A0FA16EDA30" box="[181,324,1544,1571]" italics="true" pageId="10" pageNumber="11">C. fasciolata</emphasis>
</taxonomicName>
,
<figureCitation id="130180E3B629C37DFEBF0A0EA180DA31" box="[339,426,1544,1571]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[308,1279,181,1919]" captionTargetId="figure-15@13.[308,1279,181,1919]" captionTargetPageId="13" captionText="FIGURE 2A–D. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis male; B, Cyclocephala brevipennis female; C, Cyclocephala fasciolata male; D, Cyclocephala santandereana (holotype). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100348" httpUri="https://zenodo.org/record/10100348/files/figure.png" pageId="10" pageNumber="11">Fig. 2C</figureCitation>
, but with setae present in
<taxonomicName id="4C3AE7E5B629C37DFD3D0A0FA3B2DA30" authorityName="Neita-Moreno" authorityYear="2021" box="[721,920,1544,1571]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="santandereana">
<emphasis id="B94E4074B629C37DFD3D0A0FA3B2DA30" box="[721,920,1544,1571]" italics="true" pageId="10" pageNumber="11">C. santandereana</emphasis>
</taxonomicName>
<figureCitation id="130180E3B629C37DFC4C0A0EA422DA31" box="[928,1032,1544,1571]" captionStart="FIGURE 25" captionStartId="39.[151,250,1962,1986]" captionTargetBox="[154,1436,777,1932]" captionTargetId="figure-280@39.[151,1436,777,1939]" captionTargetPageId="39" captionText="FIGURE 25A–D. Pygidia and heads (dorsal view). A, Cyclocephala rufa pygidium; B, Cyclocephala arhuacana (holotype) pygidium; C, Cyclocephala brevipennis head (dorsal view); D, Cyclocephala santandereana (holotype) head (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.10100404" httpUri="https://zenodo.org/record/10100404/files/figure.png" pageId="10" pageNumber="11">Fig. 25D</figureCitation>
); pygidium black (similar to
<taxonomicName id="4C3AE7E5B629C37DFABD0A0FA0CFDA54" authorityName="Bates" authorityYear="1888" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciolata">
<emphasis id="B94E4074B629C37DFABD0A0FA0CFDA54" italics="true" pageId="10" pageNumber="11">C. fasciolata</emphasis>
</taxonomicName>
), with punctures stronger, denser and with slender, long setae (
<figureCitation id="130180E3B629C37DFC460A2AA439DA55" box="[938,1043,1580,1607]" captionStart="FIGURE 24" captionStartId="35.[151,250,1840,1864]" captionTargetBox="[152,1423,181,1807]" captionTargetId="figure-15@35.[151,1436,181,1816]" captionTargetPageId="35" captionText="FIGURE 24A–H. Pygidia (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina." figureDoi="http://doi.org/10.5281/zenodo.10100402" httpUri="https://zenodo.org/record/10100402/files/figure.png" pageId="10" pageNumber="11">Fig. 24D</figureCitation>
), while the pygidium of
<taxonomicName id="4C3AE7E5B629C37DFACB0A2BA0D8DA78" authorityName="Neita-Moreno" authorityYear="2021" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="santandereana">
<emphasis id="B94E4074B629C37DFACB0A2BA0D8DA78" italics="true" pageId="10" pageNumber="11">C. santandereana</emphasis>
</taxonomicName>
is testaceous, with sparse punctures, and with slender, long setae (
<figureCitation id="130180E3B629C37DFC310A56A469DA79" box="[989,1091,1616,1643]" captionStart="FIGURE 24" captionStartId="35.[151,250,1840,1864]" captionTargetBox="[152,1423,181,1807]" captionTargetId="figure-15@35.[151,1436,181,1816]" captionTargetPageId="35" captionText="FIGURE 24A–H. Pygidia (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina." figureDoi="http://doi.org/10.5281/zenodo.10100402" httpUri="https://zenodo.org/record/10100402/files/figure.png" pageId="10" pageNumber="11">Fig. 24E</figureCitation>
). The surface of the pronotum, elytra, and metasternum has sparse punctures (dense in
<taxonomicName id="4C3AE7E5B629C37DFCE30A73A3B4DA9C" authorityName="Bates" authorityYear="1888" box="[783,926,1652,1679]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciolata">
<emphasis id="B94E4074B629C37DFCE30A73A3B4DA9C" box="[783,926,1652,1679]" italics="true" pageId="10" pageNumber="11">C. fasciolata</emphasis>
</taxonomicName>
dense); the surface of the pygidium is finely scabrous with ocellate punctures (in
<taxonomicName id="4C3AE7E5B629C37DFDDB0A9FA2EDDAA0" authorityName="Bates" authorityYear="1888" box="[567,711,1688,1715]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fasciolata">
<emphasis id="B94E4074B629C37DFDDB0A9FA2EDDAA0" box="[567,711,1688,1715]" italics="true" pageId="10" pageNumber="11">C. fasciolata</emphasis>
</taxonomicName>
the surface is rugopunctate, and the punctures are not ocellate). The external edge at the base of protarsomere 5 lacks a small concavity (
<figureCitation id="130180E3B629C37DFC360ABAA46ADAC5" box="[986,1088,1724,1751]" captionStart="FIGURE 17" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[178,1404,181,1911]" captionTargetId="figure-29@28.[169,1418,181,1919]" captionTargetPageId="28" captionText="FIGURE 17A–J. Protarsomere 5 (caudal view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100386" httpUri="https://zenodo.org/record/10100386/files/figure.png" pageId="10" pageNumber="11">Fig. 17C</figureCitation>
) (with a small concavity in
<taxonomicName id="4C3AE7E5B629C37DFA6F0ABBA116DAE8" authorityName="Neita-Moreno" authorityYear="2021" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="santandereana">
<emphasis id="B94E4074B629C37DFA6F0ABBA116DAE8" italics="true" pageId="10" pageNumber="11">C. santandereana</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB629C37CFF2B0B02A187DF07" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
<paragraph id="8B859C66B629C37CFF2B0B02A16ADDE7" blockId="10.[151,1437,1508,2039]" lastBlockId="11.[151,1437,151,1149]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B94E4074B629C37DFF2B0B02A190DB0D" bold="true" box="[199,442,1796,1823]" pageId="10" pageNumber="11">Redescription. Male</emphasis>
(
<figureCitation id="130180E3B629C37DFE260B02A20CDB0D" box="[458,550,1796,1823]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[308,1279,181,1919]" captionTargetId="figure-15@13.[308,1279,181,1919]" captionTargetPageId="13" captionText="FIGURE 2A–D. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis male; B, Cyclocephala brevipennis female; C, Cyclocephala fasciolata male; D, Cyclocephala santandereana (holotype). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100348" httpUri="https://zenodo.org/record/10100348/files/figure.png" pageId="10" pageNumber="11">Fig. 2A</figureCitation>
). Length 19.4–20.0 mm; width
<quantity id="4CC23183B629C37DFC620B03A40DDB0D" box="[910,1063,1797,1823]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.1" metricValueMin="8.9" pageId="10" pageNumber="11" unit="mm" value="9.5" valueMax="10.1" valueMin="8.9">8.9–10.1 mm</quantity>
. Color black except each elytron and pronotum with transverse, slightly oblique, orange band not reaching lateral or sutural margins; band occasionally broken in 4 spots or nearly obsolete. Some specimens have elytra almost completely reddish orange.
<emphasis id="B94E4074B629C37DFABF0B4BA5B6DB75" bold="true" box="[1363,1436,1869,1895]" pageId="10" pageNumber="11">Head:</emphasis>
Surface of frons and clypeus glabrous, with punctures moderate in size, becoming denser and smaller at clypeal apex. Clypeal apex broadly, weakly rounded, narrowly reflexed. Mandibles apically rounded, internal face with a groove (
<figureCitation id="130180E3B629C37DFF1A0BBEA178DBC1" box="[246,338,1976,2003]" captionStart="FIGURE 9" captionStartId="20.[151,250,1912,1936]" captionTargetBox="[151,1429,188,1878]" captionTargetId="figure-29@20.[151,1436,181,1888]" captionTargetPageId="20" captionText="FIGURE 9A–D. Adult mandibles and maxillae. A, Cyclocephala brevipennis mandible (ventral view); B, Cyclocephala brevipennis maxilla (ventral view); C, Cyclocephala fasciolata mandible (dorsal view); D, Cyclocephala fasciolata maxilla (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.10100366" httpUri="https://zenodo.org/record/10100366/files/figure.png" pageId="10" pageNumber="11">Fig. 9A</figureCitation>
); labium densely setose, paraglossa undeveloped, apex widely notched (
<figureCitation id="130180E3B629C37DFB610BBEA4DDDBC1" box="[1165,1271,1976,2003]" captionStart="FIGURE 12" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[264,1334,202,1911]" captionTargetId="figure-15@23.[253,1334,181,1919]" captionTargetPageId="23" captionText="FIGURE 12A–F. Adult labia. A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala fasciolata; D, Cyclocephala brevipennis; E, Cyclocephala ligyrina; F, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100372" httpUri="https://zenodo.org/record/10100372/files/figure.png" pageId="10" pageNumber="11">Fig. 12D</figureCitation>
); maxilla with galea developed, with 6 teeth (
<figureCitation id="130180E3B629C37DFE110BDAA273DBE5" box="[509,601,2012,2039]" captionStart="FIGURE 9" captionStartId="20.[151,250,1912,1936]" captionTargetBox="[151,1429,188,1878]" captionTargetId="figure-29@20.[151,1436,181,1888]" captionTargetPageId="20" captionText="FIGURE 9A–D. Adult mandibles and maxillae. A, Cyclocephala brevipennis mandible (ventral view); B, Cyclocephala brevipennis maxilla (ventral view); C, Cyclocephala fasciolata mandible (dorsal view); D, Cyclocephala fasciolata maxilla (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.10100366" httpUri="https://zenodo.org/record/10100366/files/figure.png" pageId="10" pageNumber="11">Fig. 9B</figureCitation>
). Epipharynx rectangular, densely setose, apex projecting at center (
<figureCitation id="130180E3B629C37CFA830BDAA0CCDCA3" captionStart="FIGURE 13" captionStartId="24.[151,250,1821,1845]" captionTargetBox="[153,1423,181,1788]" captionTargetId="figure-29@24.[151,1436,181,1798]" captionTargetPageId="24" captionText="FIGURE 13A–F. Adult epipharynxes. A, Cyclocephala marqueti ventral view; B, Cyclocephala marqueti dorsal view; C, Cyclocephala proxima ventral view; D, Cyclocephala proxima dorsal view; E, Cyclocephala brevipennis ventral view; F, Cyclocephala brevipennis dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10100377" httpUri="https://zenodo.org/record/10100377/files/figure.png" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">Fig. 13E–F</figureCitation>
). Interocular width equals 3.6–3.8 tranverse eye diameters. Antenna with 10 antennomeres, club subequal in length to antennomeres 2–7 (slightly shorter in female).
<emphasis id="B94E4074B628C37CFCD30CBDA3EBDCC7" bold="true" box="[831,961,187,213]" pageId="11" pageNumber="12">Pronotum:</emphasis>
Surface similar in punctation to frons by except punctures slightly larger. Base with strong marginal bead (Fig. 32B).
<emphasis id="B94E4074B628C37CFC160CD9A465DCEB" bold="true" box="[1018,1103,223,249]" pageId="11" pageNumber="12">Elytra:</emphasis>
Surface with rows of moderately large, shallow punctures.
<emphasis id="B94E4074B628C37CFE1B0D04A259DD0E" bold="true" box="[503,627,258,284]" pageId="11" pageNumber="12">Pygidium:</emphasis>
Surface finely scabrous, dull, setigerous; setae moderately dense, long, tawny in color (
<figureCitation id="130180E3B628C37CFEA50D21A199DD53" box="[329,435,295,321]" captionStart="FIGURE 24" captionStartId="35.[151,250,1840,1864]" captionTargetBox="[152,1423,181,1807]" captionTargetId="figure-15@35.[151,1436,181,1816]" captionTargetPageId="35" captionText="FIGURE 24A–H. Pygidia (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina." figureDoi="http://doi.org/10.5281/zenodo.10100402" httpUri="https://zenodo.org/record/10100402/files/figure.png" pageId="11" pageNumber="12">Fig. 24D</figureCitation>
). In lateral view, surface regularly convex.
<emphasis id="B94E4074B628C37CFC740D21A3F0DD53" bold="true" box="[920,986,295,321]" pageId="11" pageNumber="12">Legs:</emphasis>
Protibia bidentate (
<figureCitation id="130180E3B628C37CFB5B0D21A50ADD53" box="[1207,1312,295,321]" captionStart="FIGURE 18" captionStartId="29.[151,250,1846,1870]" captionTargetBox="[161,1428,191,1813]" captionTargetId="figure-15@29.[151,1436,181,1822]" captionTargetPageId="29" captionText="FIGURE 18A–M. Protibiae (dorsal view). A, Cyclocephala marqueti (male); B, Cyclocephala proxima (male); C, Cyclocephala proxima (female); D, Cyclocephala brevipennis (male); E, Cyclocephala brevipennis (female); F, Cyclocephala fasciolata; G, Cyclocephala santandereana (holotype); H, Cyclocephala dalensi; I, Cyclocephala kaszabi; J, Cyclocephala ligyrina; K, Cyclocephala rufa; L, Cyclocephala arhuacana (holotype); M, Cyclocephala arhuacana (allotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100388" httpUri="https://zenodo.org/record/10100388/files/figure.png" pageId="11" pageNumber="12">Fig. 18D</figureCitation>
). Protarsus enlarged, tarsomeres 2–4 with large ventral lobes, all tarsomeres longitudinally striate on dorsal face, 5
<superScript id="7C4F312EB628C37CFAF90D4CA508DD4A" attach="left" box="[1301,1314,330,344]" fontSize="6" pageId="11" pageNumber="12">th</superScript>
large (
<figureCitation id="130180E3B628C37CFA830D4DA0E3DD9B" captionStart="FIGURE 17" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[178,1404,181,1911]" captionTargetId="figure-29@28.[169,1418,181,1919]" captionTargetPageId="28" captionText="FIGURE 17A–J. Protarsomere 5 (caudal view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100386" httpUri="https://zenodo.org/record/10100386/files/figure.png" pageId="11" pageNumber="12">Fig. 17C</figureCitation>
), median claw moderately curved, without lobes or tooth, but lateral internal surface with longitudinal lines (
<figureCitation id="130180E3B628C37CFF4C0D95A12DDDBC" box="[160,263,403,430]" captionStart="FIGURE 16" captionStartId="27.[151,250,1676,1700]" captionTargetBox="[167,1426,192,1639]" captionTargetId="figure-15@27.[151,1436,181,1652]" captionTargetPageId="27" captionText="FIGURE 16A–J. Adult male large claw (lateral view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100384" httpUri="https://zenodo.org/record/10100384/files/figure.png" pageId="11" pageNumber="12">Fig. 16C</figureCitation>
), apex finely cleft (
<figureCitation id="130180E3B628C37CFE0A0D95A267DDBF" box="[486,589,402,429]" captionStart="FIGURE 15" captionStartId="26.[151,250,1943,1967]" captionTargetBox="[219,1372,189,1907]" captionTargetId="figure-29@26.[206,1381,181,1919]" captionTargetPageId="26" captionText="FIGURE 15A–J. Adult male large claw (caudal view). A, Cyclcephala marqueti; B, Cyclocephala proxima; C, Cyclocephala brevipennis; D, Cyclocephala fasciolata; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina; I, Cyclocephala rufa; J, Cyclocephala arhuacana (holotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100382" httpUri="https://zenodo.org/record/10100382/files/figure.png" pageId="11" pageNumber="12">Fig. 15C</figureCitation>
). Metatarsus longer than metatibia.
<emphasis id="B94E4074B628C37CFC0A0D95A415DDBF" bold="true" box="[998,1087,403,429]" pageId="11" pageNumber="12">Venter:</emphasis>
Prosternal process long, stout, apex obliquely flattened into a transverse oval with anterior 2/3 raised as convex “button”.
<emphasis id="B94E4074B628C37CFB600DB0A529DDC2" bold="true" box="[1164,1283,438,464]" pageId="11" pageNumber="12">Genitalia:</emphasis>
Parameres as in
<figureCitation id="130180E3B628C37CFF580DDDA116DDE7" box="[180,316,475,501]" captionStart="FIGURE 27" captionStartId="41.[151,250,1885,1909]" captionTargetBox="[205,1382,190,1855]" captionTargetId="figure-15@41.[193,1395,181,1861]" captionTargetPageId="41" captionText="FIGURE 27A–F. Parameres and phallobases, Cyclocephala fasciolata species group. A, Cyclocephala fasciolata parameres, caudal view; B, Cyclocephala fasciolata phallobase, lateral view; C, Cyclocephala brevipennis parameres, caudal view; D, Cyclocephala brevipennis phallobase, lateral view; E, Cyclocephala santandereana parameres, caudal view; F, Cyclocephala santandereana phallobase, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10100409" httpUri="https://zenodo.org/record/10100409/files/figure.png" pageId="11" pageNumber="12">Fig. 27C–D</figureCitation>
.
</paragraph>
<paragraph id="8B859C66B628C37CFF2B0DF9A187DF07" blockId="11.[151,1437,151,1149]" pageId="11" pageNumber="12">
<emphasis id="B94E4074B628C37CFF2B0DF9A134DE0B" bold="true" box="[199,286,511,537]" pageId="11" pageNumber="12">Female</emphasis>
(
<figureCitation id="130180E3B628C37CFEC10DF9A1AFDE0B" box="[301,389,511,537]" captionStart="FIGURE 2" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[308,1279,181,1919]" captionTargetId="figure-15@13.[308,1279,181,1919]" captionTargetPageId="13" captionText="FIGURE 2A–D. Cyclocephala fasciolata species group. A, Cyclocephala brevipennis male; B, Cyclocephala brevipennis female; C, Cyclocephala fasciolata male; D, Cyclocephala santandereana (holotype). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100348" httpUri="https://zenodo.org/record/10100348/files/figure.png" pageId="11" pageNumber="12">Fig. 2B</figureCitation>
). Length
<quantity id="4CC23183B628C37CFE1C0DF9A2BCDE0B" box="[496,662,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.935" metricValueMax="1.98" metricValueMin="1.89" pageId="11" pageNumber="12" unit="mm" value="19.35" valueMax="19.8" valueMin="18.9">18.9–19.8 mm</quantity>
; width
<quantity id="4CC23183B628C37CFD040DF9A35BDE0B" box="[744,881,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="7.699999999999999" metricValueMax="8.1" metricValueMin="7.3" pageId="11" pageNumber="12" unit="mm" value="7.699999999999999" valueMax="8.1" valueMin="7.3">7.3–8.1 mm</quantity>
. Similar to male, but surface of frons and clypeus with punctation denser. Antennal club slightly shorther than antennomeres 2–7. Pronotum and elytra with punctures stronger and denser. Abdominal ventrite VIII entire. Epipleuron (ventral view) expanded into elongate, laterally expanded flange extending from middle of lateral edge of metacoxa to anterior edge of third sternite; in lateral view, epipleuron beneath flange produced ventrally into strong, posteriorly projecting, acute tooth (
<figureCitation id="130180E3B628C37CFB530E89A500DEBB" box="[1215,1322,655,681]" captionStart="FIGURE 20" captionStartId="31.[151,250,1913,1937]" captionTargetBox="[196,1389,161,1875]" captionTargetId="figure-15@31.[185,1402,151,1890]" captionTargetPageId="31" captionText="FIGURE 20A–G. Female epipleura (ventral view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100394" httpUri="https://zenodo.org/record/10100394/files/figure.png" pageId="11" pageNumber="12">Fig. 20D</figureCitation>
). Pygidial surface shiny, punctures moderately dense and moderate in size, setigerous. In lateral view, surface nearly flat. Protibia tridentate (
<figureCitation id="130180E3B628C37CFEAB0ED1A180DEE3" box="[327,426,727,753]" captionStart="FIGURE 18" captionStartId="29.[151,250,1846,1870]" captionTargetBox="[161,1428,191,1813]" captionTargetId="figure-15@29.[151,1436,181,1822]" captionTargetPageId="29" captionText="FIGURE 18A–M. Protibiae (dorsal view). A, Cyclocephala marqueti (male); B, Cyclocephala proxima (male); C, Cyclocephala proxima (female); D, Cyclocephala brevipennis (male); E, Cyclocephala brevipennis (female); F, Cyclocephala fasciolata; G, Cyclocephala santandereana (holotype); H, Cyclocephala dalensi; I, Cyclocephala kaszabi; J, Cyclocephala ligyrina; K, Cyclocephala rufa; L, Cyclocephala arhuacana (holotype); M, Cyclocephala arhuacana (allotype). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10100388" httpUri="https://zenodo.org/record/10100388/files/figure.png" pageId="11" pageNumber="12">Fig. 18E</figureCitation>
). Protarsus simple. Length of metatarsus shorter than metatibia. Gonocoxite and gonocoxal sternite fused (Fig. 32F).
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB628C37CFF2B0F18A13DD86F" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8B859C66B628C37CFF2B0F18A106DF93" blockId="11.[151,1437,151,1149]" pageId="11" pageNumber="12">
<emphasis id="B94E4074B628C37CFF2B0F18A173DF2A" bold="true" box="[199,345,798,824]" pageId="11" pageNumber="12">Distribution</emphasis>
.
<taxonomicName id="4C3AE7E5B628C37CFE870F19A2B8DF2B" authorityName="Endrodi" authorityYear="1964" box="[363,658,799,825]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="B94E4074B628C37CFE870F19A2B8DF2B" box="[363,658,799,825]" italics="true" pageId="11" pageNumber="12">Cyclocephala brevipennis</emphasis>
</taxonomicName>
is known from
<collectingCountry id="F32DDCF6B628C37CFCBF0F19A39ADF2B" box="[851,944,799,825]" name="Ecuador" pageId="11" pageNumber="12">Ecuador</collectingCountry>
(
<bibRefCitation id="EFABE197B628C37CFC2F0F19A44FDF2B" author="Endrodi, S." box="[963,1125,798,825]" pageId="11" pageNumber="12" refId="ref23141" refString="Endrodi, S. (1985) The Dynastinae of the World. Dr. W. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrödi 1985</bibRefCitation>
;
<bibRefCitation id="EFABE197B628C37CFB980F19A548DF2B" author="Ratcliffe, B. C. &amp; Cave, R. D. &amp; Paucar-Cabrera, A." box="[1140,1378,799,825]" pageId="11" pageNumber="12" pagination="1 - 565" refId="ref24303" refString="Ratcliffe, B. C., Cave, R. D. &amp; Paucar-Cabrera, A. (2020) The dynastine scarab beetles of Ecuador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 32, 1 - 565." type="journal article" year="2020">
Ratcliffe
<emphasis id="B94E4074B628C37CFB0D0F19A53CDF2B" box="[1249,1302,799,825]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
. 2020
</bibRefCitation>
) and
<collectingCountry id="F32DDCF6B628C37CFF7B0F45A12FDF4F" box="[151,261,835,861]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
(
<bibRefCitation id="EFABE197B628C37CFEF80F45A1BFDF4F" author="Neita, J. C." box="[276,405,835,861]" pageId="11" pageNumber="12" pagination="17 - 27" refId="ref23942" refString="Neita, J. C. (2011) Escarabajos (Coleoptera: Scarabaeoidea) del departamento del Choco, Colombia. Revista Biodiversidad Neotropical, 1, 17 - 27. https: // doi. org / 10.18636 / bioneotropical. v 1 i 1.25" type="journal article" year="2011">Neita 2011</bibRefCitation>
). The species was previously reported in
<collectingCountry id="F32DDCF6B628C37CFC8C0F45A3E4DF4F" box="[864,974,835,861]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
but was misidentified as
<taxonomicName id="4C3AE7E5B628C37CFB070F45A553DF4F" authorityName="Bates" authorityYear="1888" box="[1259,1401,835,861]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fasciolata">
<emphasis id="B94E4074B628C37CFB070F45A553DF4F" box="[1259,1401,835,861]" italics="true" pageId="11" pageNumber="12">C. fasciolata</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFABE197B628C37CFF7B0F61A102DF93" author="Neita, J. C." box="[151,296,871,897]" pageId="11" pageNumber="12" pagination="17 - 27" refId="ref23942" refString="Neita, J. C. (2011) Escarabajos (Coleoptera: Scarabaeoidea) del departamento del Choco, Colombia. Revista Biodiversidad Neotropical, 1, 17 - 27. https: // doi. org / 10.18636 / bioneotropical. v 1 i 1.25" type="journal article" year="2011">Neita (2011)</bibRefCitation>
.
</paragraph>
<paragraph id="8B859C66B628C37CFF2B0F8CA3FDDFB7" blockId="11.[151,1437,151,1149]" box="[199,983,906,933]" pageId="11" pageNumber="12">
<emphasis id="B94E4074B628C37CFF2B0F8CA178DFB6" bold="true" box="[199,338,906,932]" pageId="11" pageNumber="12">Life history</emphasis>
. Adults are active during the night and attracted to lights.
</paragraph>
<paragraph id="8B859C66B628C37CFF2B0FA8A13DD86F" blockId="11.[151,1437,151,1149]" pageId="11" pageNumber="12">
<emphasis id="B94E4074B628C37CFF2B0FA8A1A6DFDB" bold="true" box="[199,396,942,969]" pageId="11" pageNumber="12">Locality records</emphasis>
(
<figureCitation id="130180E3B628C37CFE700FA9A1D8DFD8" box="[412,498,943,970]" captionStart="FIGURE 36" captionStartId="50.[151,250,1624,1648]" captionTargetBox="[159,1423,191,1584]" captionTargetId="figure-29@50.[151,1436,181,1600]" captionTargetPageId="50" captionText="FIGURE 36A–C. Cyclocephala marqueti, third instar and pupa. A, Terminal abdominal segment, ventral view; B, stout setar; C, Cyclocephala marqueti, pupa, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10100424" httpUri="https://zenodo.org/record/10100424/files/figure.png" pageId="11" pageNumber="12">Fig. 36</figureCitation>
).
<materialsCitation id="3B52963BB628C37CFDEB0FA9A138D86F" collectingDate="2009-12" collectionCode="IAvH, IAvH-" collectorName="S. Sarria Leg. &amp; J. C. Neita" country="Colombia" elevation="730" latitude="3.4333334" location="Four" longLatPrecision="1307" longitude="-76.8" pageId="11" pageNumber="12" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Valle del Cauca">
<location id="8EE5CABDB628C37CFDEB0FA9A216DFDB" LSID="urn:lsid:plazi:treatment:03932D70B629C37CFF7B097EA13DD86F:8EE5CABDB628C37CFDEB0FA9A216DFDB" box="[519,572,943,969]" country="Colombia" latitude="3.4333334" longLatPrecision="1307" longitude="-76.8" name="Four" pageId="11" pageNumber="12" stateProvince="Valle del Cauca">Four</location>
specimens examined from
<collectionCode id="ED2B04A3B628C37CFC9A0FA9A39DDFDB" box="[886,951,943,969]" pageId="11" pageNumber="12">IAvH</collectionCode>
. “
<collectingCountry id="F32DDCF6B628C37CFC210FA9A46BDFDB" box="[973,1089,943,969]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
,
<collectingRegion id="49FE5284B628C37CFBA10FA9A52ADFDB" box="[1101,1280,943,969]" country="Colombia" name="Valle del Cauca" pageId="11" pageNumber="12">Valle del Cauca</collectingRegion>
/ PNN Farallones de Cali/
<geoCoordinate id="EE0EFAA1B628C37CFED40FD5A1A2DFFF" box="[312,392,979,1005]" degrees="3" direction="north" minutes="26" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="3.4333334">3º26´N</geoCoordinate>
;
<geoCoordinate id="EE0EFAA1B628C37CFE790FD5A1D0DFFF" box="[405,506,979,1005]" degrees="76" direction="west" minutes="48" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="-76.8">76º48´W</geoCoordinate>
.
<quantity id="4CC23183B628C37CFDEA0FD5A264DFFF" box="[518,590,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="11" pageNumber="12" unit="m" value="730.0">
<elevation id="00177B55B628C37CFDEA0FD5A264DFFF" box="[518,590,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="11" pageNumber="12" unit="m" value="730.0">730 m</elevation>
</quantity>
/ Malaise. 8/01/00-10/10/00 /
<collectorName id="26CFF9B0B628C37CFC460FD5A462DFFF" box="[938,1096,979,1005]" pageId="11" pageNumber="12">S. Sarria Leg.</collectorName>
M. 1104” [
<specimenCount id="9D3C57EFB628C37CFB270FD4A4F3DFFF" box="[1227,1241,978,1005]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
-IAvH-E-13143] and “
<collectingCountry id="F32DDCF6B628C37CFF3E0FF1A16CD803" box="[210,326,1015,1041]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
,
<collectingRegion id="49FE5284B628C37CFEBF0FF1A22CD803" box="[339,518,1015,1041]" country="Colombia" name="Valle del Cauca" pageId="11" pageNumber="12">Valle del Cauca</collectingRegion>
/ PNN Farallones de Cali /
<geoCoordinate id="EE0EFAA1B628C37CFCA40FF1A3B6D803" box="[840,924,1015,1041]" degrees="3" direction="north" minutes="26" orientation="latitude" pageId="11" pageNumber="12" precision="925" value="3.4333334">3º26´N</geoCoordinate>
;
<geoCoordinate id="EE0EFAA1B628C37CFC450FF1A424D803" box="[937,1038,1015,1041]" degrees="76" direction="west" minutes="48" orientation="longitude" pageId="11" pageNumber="12" precision="925" value="-76.8">76º48´W</geoCoordinate>
.
<quantity id="4CC23183B628C37CFBF70FF1A44ED803" box="[1051,1124,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="11" pageNumber="12" unit="m" value="730.0">
<elevation id="00177B55B628C37CFBF70FF1A44ED803" box="[1051,1124,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="11" pageNumber="12" unit="m" value="730.0">730 m</elevation>
</quantity>
/ Malaise. 7/18/00-8/1/00 /
<collectorName id="26CFF9B0B628C37CFF7B081DA11CD827" box="[151,310,1051,1077]" pageId="11" pageNumber="12">S. Sarria Leg.</collectorName>
M. 1101” [
<specimenCount id="9D3C57EFB628C37CFE57081CA1E3D827" box="[443,457,1050,1077]" pageId="11" pageNumber="12" type="female">♀</specimenCount>
IAvH-E-27167]. “
<collectingCountry id="F32DDCF6B628C37CFD71081DA33BD827" box="[669,785,1051,1077]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
, Chocó, Quibdó / Corregimiento de Tutunendo, / Vereda Cara de Perro. / 5º45´07.9´N; 76º45´16.8´W. 340 / m alt.
<date id="FF84BAA6B628C37CFCCD0839A3BED84B" box="[801,916,1087,1113]" pageId="11" pageNumber="12" value="2009-12">
<collectingDate id="EFC0434EB628C37CFCCD0839A3BED84B" box="[801,916,1087,1113]" pageId="11" pageNumber="12" value="2009-12">Dic. 2009</collectingDate>
</date>
.
<collectorName id="26CFF9B0B628C37CFC730839A432D84B" box="[927,1048,1087,1113]" pageId="11" pageNumber="12">J. C. Neita</collectorName>
M.” [
<specimenCount id="9D3C57EFB628C37CFBB60839A451D84B" box="[1114,1147,1087,1113]" pageId="11" pageNumber="12" type="male">2♂</specimenCount>
-
<collectionCode id="ED2B04A3B628C37CFB6F0839A4FAD84B" box="[1155,1232,1087,1113]" pageId="11" pageNumber="12">IAvH-</collectionCode>
E-216416,
<collectionCode id="ED2B04A3B628C37CFAB90839A5B6D84B" box="[1365,1436,1087,1113]" pageId="11" pageNumber="12">IAvH-</collectionCode>
E-216417]
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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