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<document ID-CLB-Dataset="297097" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="f9373c90-73c5-4ee4-b0f8-4c371f527d91" ID-ISBN="0-8018-8221-4" checkinTime="1670905209323" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="80B561A27E5280B7BB550D6422A9F34B" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Muridae.1189-1531.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Rattus leucopus Gray 1867" docType="treatment" docVersion="4" id="A2267A57FF8A65DA07E1D2E970F9422A" lastPageNumber="1472" masterDocId="A2267A57FF8A65DA07E1D2E970F9422A" masterDocTitle="Order Rodentia - Family Muridae" masterLastPageNumber="1531" masterPageNumber="1189" pageNumber="1472" updateTime="1716781534040" updateUser="ExternalLinkService">
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<mods:title id="E7D0EF010770D864DFC4EF33B6C18325">Order Rodentia - Family Muridae</mods:title>
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<mods:namePart id="15F7220679803D40D008B20AF893B702">Wilson, Don E.</mods:namePart>
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<mods:namePart id="80E53F84ED3865F51C54C20292D9FADE">Reeder, DeeAnn</mods:namePart>
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<mods:title id="E0400F6BBE282DDBCA9A4D27D73C9A2E">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
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<treatment id="80B561A27E5280B7BB550D6422A9F34B" ID-DOI="http://doi.org/10.5281/zenodo.11335498" ID-GBIF-Taxon="231540079" ID-Zenodo-Dep="11335498" LSID="urn:lsid:plazi:treatment:80B561A27E5280B7BB550D6422A9F34B" httpUri="http://treatment.plazi.org/id/80B561A27E5280B7BB550D6422A9F34B" lastPageNumber="1472" pageNumber="1472" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=13001752">
<subSubSection id="63BEA333733B097562890C7D59712D50" pageNumber="1472" type="nomenclature">
<paragraph id="5CED85208B125E7D6794B94D1763F22D" pageNumber="1472">
<taxonomicName id="8C82D2386DF9C067CB19D90A35C07B4E" ID-CoL="6X58L" authority="Gray 1867" authorityName="Gray" authorityYear="1867" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1472" phylum="Chordata" rank="species" species="leucopus">
<emphasis id="642F76CF40D810F18D50E6C76948CFA8" italics="true" pageNumber="1472">Rattus leucopus</emphasis>
Gray 1867
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="85F7B902575CB77F0405449004601A70" pageNumber="1472" type="reference_group">
<paragraph id="BEF97ACADB5EF930A0F297E6AF2737F3" pageNumber="1472">
<treatmentCitation id="FE1FF9AF54EBE315B26568B5A5B803A2" author="Gray" authority="Gray 1867" authorityName="Gray" authorityYear="1867" class="Mammalia" family="Muridae" genus="Rattus" journal="Proc. Zool. Soc. Lond." kingdom="Animalia" order="Rodentia" page="598" pageNumber="1472" phylum="Chordata" rank="species" species="leucopus" title="Rattus leucopus" volume="1867" year="1867">
<taxonomicName id="D36007058D509483FFE7F33CEDE0EE5E" ID-CoL="6X58L" authority="Gray 1867" authorityName="Gray" authorityYear="1867" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1472" phylum="Chordata" rank="species" species="leucopus">
<emphasis id="11E71E7C6C3B0D68FBF829EC44BD46C8" italics="true" pageNumber="1472">Rattus leucopus</emphasis>
Gray 1867
</taxonomicName>
,
<bibCitation id="D418822A4DD7E26B0AF7179C2ACFE006" author="Gray" journal="Proc. Zool. Soc. Lond." pageNumber="1472" pagination="598" title="Rattus leucopus" volume="1867" year="1867">Proc. Zool. Soc. Lond., 1867: 598</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5D54F8C01C15801F45D905BF665563F8" pageNumber="1472" type="materials_examined">
<paragraph id="03FD41D507F588B37AB3A8C6B7644FD9" pageNumber="1472">
<emphasis id="C1632A9CEB395280C3950309318CF958" inLineHeading="true" pageNumber="1472">Type Locality:</emphasis>
<materialsCitation id="980D0714D861502DF37B2A7EBC03687A" ID-GBIF-Occurrence="4867324403" country="Australia" pageNumber="1472" stateProvince="Queensland">
<collectingCountry id="37B0A5233A509CC43E6AF12A4D602D8C" name="Australia">Australia</collectingCountry>
,
<collectingRegion id="920741516E7F0C0B55E92F853988C883" country="Australia" name="Queensland">Queensland</collectingRegion>
, Cape York (as restricted by Thomas
<typeStatus id="51E84A14D140864ADF9A3A8EE7BB1C24">lectotype</typeStatus>
selection; see
<bibRefCitation id="D29AF00E807578E21169BF52C5786CB5" author="Mahoney, J. A. &amp; B. J. Richardson" refId="ref222168" refString="Mahoney, J. A., and B. J. Richardson. 1988. Muridae. Pp. 154 - 192, in Zoological catalogue of Australia. Mammalia (J. L. Bannister, et. al.). Australian Government Publishing Service, Canberra, 5: 1 - 274." year="1988">Mahoney and Richardson, 1988:183</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="ECEE89AC7BBFD14E1FC3CC67BFD93CC9" pageNumber="1472" type="vernacular_names">
<paragraph id="7FDD04422F2FB96C259BFE921E10E16A" pageNumber="1472">
<emphasis id="27D2D0B2745AD0B0C72C94EA3D7D6CE0" inLineHeading="true" pageNumber="1472">Vernacular Names:</emphasis>
<vernacularName id="B7CA7A5663B4BEB7005D81753ECB8AB0" language="eng" pageNumber="1472">
<collectingRegion id="76E33CFA16CF633DF103E1EE3667E626" country="South Africa" name="Western Cape">Cape</collectingRegion>
<collectingRegion id="92AF02EC92D97B111367AFC9B52088BF" country="United Kingdom" name="York">York</collectingRegion>
Rat
</vernacularName>
.
</paragraph>
</subSubSection>
<subSubSection id="795D4527807D95367F6DBBED2B10B631" pageNumber="1472" type="synonym_list">
<paragraph id="7A5BDA5E4C810D79C1D5CD9C3FC8DF27" pageNumber="1472">
<emphasis id="82FD35D0E86740F03220652C512949FA" inLineHeading="true" pageNumber="1472">Synonyms:</emphasis>
<taxonomicName id="2B3806018C610157CF0BF767B03D5E2F" authority="Tate 1951" authorityName="Tate" authorityYear="1951" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1472" phylum="Chordata" rank="species" species="cooktownensis">
<emphasis id="A839773B8215F196158A7A0C3730A312" italics="true" pageNumber="1472">Rattus cooktownensis</emphasis>
<bibRefCitation id="608FF571E9FFCB3A51209CE5B6881760" author="Tate, G. H. H." refId="ref347722" refString="Tate, G. H. H. 1951. Results of the Archbold Expeditions. No. 65. The rodents of Australia and New Guinea. Bulletin of the American Museum of Natural History, 97: 183 - 430." year="1951">Tate 1951</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="5D3D376101DF550EA0DBB1105726E7C8" authority="Troughton 1946" authorityName="Troughton" authorityYear="1946" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1472" phylum="Chordata" rank="species" species="dobodurae">
<emphasis id="5CE34335FC10FCC69BD0C90E2E553637" italics="true" pageNumber="1472">Rattus dobodurae</emphasis>
Troughton 1946
</taxonomicName>
;
<taxonomicName id="A4A1E52C9ABB301B351D81F40B55BD33" authority="Tate 1951" authorityName="Tate" authorityYear="1951" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1472" phylum="Chordata" rank="species" species="mcilwraithi">
<emphasis id="B29A2032C327F757D1A7388741A10AAF" italics="true" pageNumber="1472">Rattus mcilwraithi</emphasis>
<bibRefCitation id="3F51176CAFF085C74D05F1C19F1B09F0" author="Tate, G. H. H." refId="ref347722" refString="Tate, G. H. H. 1951. Results of the Archbold Expeditions. No. 65. The rodents of Australia and New Guinea. Bulletin of the American Museum of Natural History, 97: 183 - 430." year="1951">Tate 1951</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="EFBA7D9FEBB082E588F4518869A94275" authority="(Krefft 1867)" baseAuthorityName="Krefft" baseAuthorityYear="1867" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1472" phylum="Chordata" rank="species" species="personata">
<emphasis id="7E89B5B0040B74BD1592417A9C37F837" italics="true" pageNumber="1472">Rattus personata</emphasis>
(Krefft 1867)
</taxonomicName>
;
<taxonomicName id="ABCD83E3BCD0A52E6EF5E2C060B8F67D" authority="(Jentink 1908)" baseAuthorityName="Jentink" baseAuthorityYear="1908" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1472" phylum="Chordata" rank="species" species="ratticolor">
<emphasis id="B069B28189CCAB85509D27B336B5874B" italics="true" pageNumber="1472">Rattus ratticolor</emphasis>
(Jentink 1908)
</taxonomicName>
;
<taxonomicName id="C19910DBA06ABA068E96A2690BCD8C77" authority="(Peters and Doria 1881)" baseAuthorityName="Peters and Doria" baseAuthorityYear="1881" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1472" phylum="Chordata" rank="species" species="ringens">
<emphasis id="B3CD2A137A3C6BE119A0A410DCB5B0C1" italics="true" pageNumber="1472">Rattus ringens</emphasis>
(Peters and Doria 1881)
</taxonomicName>
;
<taxonomicName id="58E990CFB0E97036346E3FA33F040AFD" authority="(Alston 1879)" baseAuthorityName="Alston" baseAuthorityYear="1879" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageNumber="1472" phylum="Chordata" rank="species" species="terra-reginae">
<emphasis id="F4AE0DAD313815496C04F5FFDB76F9BB" italics="true" pageNumber="1472">Rattus terra-reginae</emphasis>
(Alston 1879)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="ED679E32FEA09CA09B9C9B9F7F76C9D9" pageNumber="1472" type="distribution">
<paragraph id="E039A697BBD27835DB0B9131EAA8A8E2" pageNumber="1472">
<emphasis id="B98421B54B38A2395C6686B5943B12AA" inLineHeading="true" pageNumber="1472">Distribution:</emphasis>
<collectingCountry id="E28CFA591889F9BF005FBD49DE681581" name="Australia">Australia</collectingCountry>
,
<collectingRegion id="36C12F7EF32CBABF3B267C04F80F3D87" country="Australia" name="Queensland">Queensland</collectingRegion>
: one population ranges from the tip of Cape York south down E side of the peninsula to vicinity of Coen, another from region of Cooktown south along the coast to Tully; all records are east of the Great Dividing Range (
<bibRefCitation id="04F7C8AE3F7C51EAC0D966706E3D0AAA" author="Moore, L. A. &amp; L. Leung" refId="ref246295" refString="Moore, L. A., and L. Leung. 1995. Cape York rat, Rattus leucopus. Pp. 653 - 655, in Mammals of Australia (R. Strahan, ed.). Smithsonian Institution Press, Washington, D. C., 756 pp." year="1995">Moore and Leung, 1995</bibRefCitation>
;
<bibRefCitation id="4A4301BAD903E8FF1600D16107053F28" author="Taylor, J. M. &amp; B. E. Horner" refId="ref348251" refString="Taylor, J. M., and B. E. Horner. 1973. Results of the Archbold Expeditions. No. 98. Systematics of native Australian Rattus (Rodentia, Muridae). Bulletin of the American Museum of Natural History, 150: 1 - 130." year="1973">Taylor and Horner, 1973</bibRefCitation>
;
<bibRefCitation id="5A1C18A9B6AB64B4AA619BE350344813" author="Watts, C. H. S. &amp; H. J. Aslin" refId="ref384328" refString="Watts, C. H. S., and H. J. Aslin. 1981. The rodents of Australia. Angus and Robertson, Sydney, 321 pp." year="1981">Watts and Aslin, 1981</bibRefCitation>
). New
<collectingCountry id="C4ED56ADFF55CDD78108AA5313DA72EB" name="Guinea-Bissau">Guinea</collectingCountry>
: widespread in lowlands south of Central Cordillera, in N and S lowland regions fringing the Owen Stanley Range in E
<collectingCountry id="9F30D9D4018275A8075274157E883122" name="Papua New Guinea">Papua New Guinea</collectingCountry>
(Flannery, 1995
<emphasis id="FAE7363DB45ED0E36AA3DC7F205E1206" italics="true" pageNumber="1472">a</emphasis>
;
<bibRefCitation id="E452FF2E64BE6B5360240A55AF5E1F0A" author="Taylor, J. M. &amp; J. H. Calaby &amp; H. M. Van Deusen" refId="ref348303" refString="Taylor, J. M., J. H. Calaby, and H. M. Van Deusen. 1982. A revision of the genus Rattus (Rodentia, Muridae) in the New Guinean region. Bulletin of the American Museum of Natural History, 173: 177 - 336." year="1982">Taylor et al., 1982:232</bibRefCitation>
); also Wokam Isl in the Aru Isls (Flannery, 1995
<emphasis id="8B8994EEEB992A9B027D815B6B97F479" italics="true" pageNumber="1472">b</emphasis>
). Altitudinal range, sea level to
<quantity id="576134E58687B04393A71C3D08D22396" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</quantity>
(Flannery, 1995
<emphasis id="E4D42146806A483476AFB706011E3502" italics="true" pageNumber="1472">a</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="6690EC1496C4B3C1858290E84084FA38" pageNumber="1472" type="conservation">
<paragraph id="65594A9D3190A7A351E4B875FD1E2CFD" pageNumber="1472">
<emphasis id="16CAC0307F17CEEF067061AD7004BE74" inLineHeading="true" pageNumber="1472">Conservation:</emphasis>
<collectionCode id="8BA788415C1F16D419507407E6029AD8">IUCN</collectionCode>
Lower Risk (lc).
</paragraph>
</subSubSection>
<subSubSection id="D6F2836A8A332D448520618EEB9B2D00" pageNumber="1472" type="discussion">
<paragraph id="62E1A3766D6021A06A9A0778DDEEB975" pageNumber="1472">
<emphasis id="8C5F350BE720ACE4D846B1DE555E06D6" inLineHeading="true" pageNumber="1472">Discussion:</emphasis>
<taxonomicName id="E36F126AA5DA9CDD25F64108972BE759" authorityName="Gray" authorityYear="1867" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="leucopus">
<emphasis id="9B207A0EF946C208ECA815BB69611028" italics="true" pageNumber="1472">Rattus leucopus</emphasis>
</taxonomicName>
species group. This species and
<taxonomicName id="EC7D72E4B1EB17682E371A13AC790C6B" authorityName="Gould" authorityYear="1857" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="sordidus">
<emphasis id="080BA9F20F0E6B338DA2D7A59F5D794D" italics="true" pageNumber="1472">R. sordidus</emphasis>
</taxonomicName>
are the only two native
<taxonomicName id="5890858ABEF7CD10AACAEA628897E180" baseAuthorityName="Musser and Newcomb" baseAuthorityYear="1983" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="7F5DDC570026FC302C9F6478A1D740EA" italics="true" pageNumber="1472">Rattus</emphasis>
</taxonomicName>
occurring on both New
<collectingCountry id="607B74BEFBEBB80425D06DBC94371E46" name="Guinea-Bissau">Guinea</collectingCountry>
and the NE coastal region of
<collectingCountry id="A9F2681442176478EE9B7BF283516600" name="Australia">Australia</collectingCountry>
(
<bibRefCitation id="AC25B3E534A4F7EB02B1849AD6D0CF30" author="Taylor, J. M. &amp; J. H. Calaby &amp; H. M. Van Deusen" refId="ref348303" refString="Taylor, J. M., J. H. Calaby, and H. M. Van Deusen. 1982. A revision of the genus Rattus (Rodentia, Muridae) in the New Guinean region. Bulletin of the American Museum of Natural History, 173: 177 - 336." year="1982">Taylor et al., 1982</bibRefCitation>
). Morphologically related to other species of
<taxonomicName id="27908B9700F06AB0C853EA1BFCDF37BF" baseAuthorityName="Musser and Newcomb" baseAuthorityYear="1983" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
<emphasis id="1EBE9A822907600778CEEEF86682C2C2" italics="true" pageNumber="1472">Rattus</emphasis>
</taxonomicName>
native to New
<collectingCountry id="ADB2D5218C79CB07621EC87216BDEBCF" name="Guinea-Bissau">Guinea</collectingCountry>
(
<bibRefCitation id="D9C420EEFE9689FC221C698019425DB8" author="Taylor, J. M. &amp; J. H. Calaby &amp; H. M. Van Deusen" refId="ref348303" refString="Taylor, J. M., J. H. Calaby, and H. M. Van Deusen. 1982. A revision of the genus Rattus (Rodentia, Muridae) in the New Guinean region. Bulletin of the American Museum of Natural History, 173: 177 - 336." year="1982">Taylor et al., 1982</bibRefCitation>
). Morphological data interpreted by
<bibRefCitation id="42811FF537B614010EF8BB90844FC950" author="Taylor, J. M. &amp; B. E. Horner" refId="ref348251" refString="Taylor, J. M., and B. E. Horner. 1973. Results of the Archbold Expeditions. No. 98. Systematics of native Australian Rattus (Rodentia, Muridae). Bulletin of the American Museum of Natural History, 150: 1 - 130." year="1973">Taylor and Horner (1973)</bibRefCitation>
to indicate close affiliation between
<taxonomicName id="DA7E4EFF59BFAC8DEB086F9A486DA0C9" authorityName="Gray" authorityYear="1867" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="leucopus">
<emphasis id="B4A6C79ABB45CC9D87E416D72DE4CB1C" italics="true" pageNumber="1472">R. leucopus</emphasis>
</taxonomicName>
and
<taxonomicName id="7D058E7E636B8A8A7F36CEF6B5A01F77" authorityName="Waterhouse" authorityYear="1839" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="fuscipes">
<emphasis id="5E89E2DD5AB1CCB1DA18050F9D9C3505" italics="true" pageNumber="1472">R. fuscipes</emphasis>
</taxonomicName>
from coastal Queensland; allozymic data discordant with this view (
<bibRefCitation id="EB1DB2AF2A4CF3A0B74403C6A67DEADA" author="Baverstock, P. R. &amp; M. Gelder &amp; A. Jahnke" refId="ref24788" refString="Baverstock, P. R., M. Gelder, and A. Jahnke. 1983 a. Chromosome evolution in Australian Rattus- - G-banding and hybrid meiosis. Genetica, 60: 93 - 103." year="1983">
Baverstock et al., 1983
<emphasis id="43B7222C16749D93C248B870948E2E96" italics="true" pageNumber="1472">a</emphasis>
</bibRefCitation>
, 1986). Australian segment reviewed by
<bibRefCitation id="E14CFC75F09D492600B7D7680562EEDB" author="Watts, C. H. S. &amp; H. J. Aslin" refId="ref384328" refString="Watts, C. H. S., and H. J. Aslin. 1981. The rodents of Australia. Angus and Robertson, Sydney, 321 pp." year="1981">Watts and Aslin (1981)</bibRefCitation>
,
<bibRefCitation id="59BAB4E6BCF24169D0C80D9D96A8F02D" author="Mahoney, J. A. &amp; B. J. Richardson" refId="ref222168" refString="Mahoney, J. A., and B. J. Richardson. 1988. Muridae. Pp. 154 - 192, in Zoological catalogue of Australia. Mammalia (J. L. Bannister, et. al.). Australian Government Publishing Service, Canberra, 5: 1 - 274." year="1988">Mahoney and Richardson (1988)</bibRefCitation>
, and
<bibRefCitation id="982A1EB9EBB58A26CF34F87964EEF5DC" author="Moore, L. A. &amp; L. Leung" refId="ref246295" refString="Moore, L. A., and L. Leung. 1995. Cape York rat, Rattus leucopus. Pp. 653 - 655, in Mammals of Australia (R. Strahan, ed.). Smithsonian Institution Press, Washington, D. C., 756 pp." year="1995">Moore and Leung (1995)</bibRefCitation>
; New
<collectingCountry id="58E0F30F84F50A056B2DB59BEEA51761" name="Guinea-Bissau">Guinea</collectingCountry>
segment reviewed by Flannery (1995
<emphasis id="52F986A2CF8153692777D4CF3E69C78E" italics="true" pageNumber="1472">a</emphasis>
).
<bibRefCitation id="0FBB05CE636DCE145DA4B9887C60127C" author="Leung, L. K. - P." refId="ref210651" refString="Leung, L. K. - P. 1999 b. Ecology of Australian tropical rainforest mammals. III. The Cape York rat, Rattus leucopus (Muridae: Rodentia). Wildlife Research, 1999, 26: 317 - 328." year="1999">
Leung (1999
<emphasis id="D92FD811280A13E9491D864C5A56DFE0" italics="true" pageNumber="1472">b</emphasis>
)
</bibRefCitation>
provided the first detailed ecological study of
<taxonomicName id="5715BD6658DA15AAD782FF8E6B661F8D" authorityName="Gray" authorityYear="1867" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="leucopus">
<emphasis id="7F9867A629C9FCCA2BF5F1829ECEE509" italics="true" pageNumber="1472">R. leucopus</emphasis>
</taxonomicName>
in
<collectingCountry id="90630DAF80871451D72F1FDE692D9EEC" name="Australia">Australia</collectingCountry>
.
<bibRefCitation id="2852C7344E791F152263D49DDA8893BC" author="Leary, T. &amp; L. Seri" refId="ref208130" refString="Leary, T., and L. Seri. 1997. An annotated checklist of mammals recorded in the Kikori River Basin, Papua New Guinea. Science in New Guinea, 23 (2): 79 - 100." year="1997">Leary and Seri (1997)</bibRefCitation>
discussed specimens taken in the Kikori River Basin of S Papua.
</paragraph>
</subSubSection>
</treatment>
</document>