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<document id="66869709CA1707D8D195652EE6F3883E" ID-CLB-Dataset="4940" ID-DOI="10.11646/zootaxa.4984.1.18" ID-GBIF-Dataset="ee0b9a1a-cbc5-4d39-bb90-1e770cb8160a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4927030" ID-ZooBank="05948731-53D9-4EA5-8ED9-8DA99548AA07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623427669319" checkinUser="plazi" docAuthor="Naumova, Maria, Blagoev, Gergin &amp; Deltshev, Christo" docDate="2021" docId="6667B114FFA65A34FF535F6D3F0C3F34" docLanguage="en" docName="zootaxa.4984.1.18.pdf" docOrigin="Zootaxa 4984 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptopilos memorialis" docType="treatment" docVersion="4" lastPageNumber="235" masterDocId="9A5EC96CFFA15A33FFC459773A77384C" masterDocTitle="Fifty spider species new to the Bulgarian fauna, with a review of some dubious species (Arachnida: Araneae)" masterLastPageNumber="257" masterPageNumber="228" pageNumber="235" updateTime="1699010390252" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9C2CBDF9CE8DDA8261013B12C22C54C8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E91539BD03CEF17134D3231F96A49062">
<mods:title id="A9276BB3ED32C87988DAA340E306B3DB">Fifty spider species new to the Bulgarian fauna, with a review of some dubious species (Arachnida: Araneae)</mods:title>
</mods:titleInfo>
<mods:name id="6EF0F7962C607D84602840E30991879B" type="personal">
<mods:role id="7B27554291C92DCF11141C649063325A">
<mods:roleTerm id="DA9AD26B8269FF0C1A8D6479DCF60231">Author</mods:roleTerm>
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<mods:namePart id="4F53801719A189B9C9A7231C9965D1D4">Naumova, Maria</mods:namePart>
<mods:nameIdentifier id="41C969E2D9F34788AED36018AA0AC7D0" type="ORCID">0000-0003-0060-048X</mods:nameIdentifier>
<mods:affiliation id="061E734780CB70F2A55537902F253A1C">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd., 1000 Sofia, Bulgaria. munny @ abv. bg; https: // orcid. org / 0000 - 0003 - 0060 - 048 X</mods:affiliation>
<mods:nameIdentifier id="78BABD783AA345321B062D320B86D453" type="email">munny@abv.bg</mods:nameIdentifier>
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<mods:name id="54CC867C062210565965B6EACE77BB5F" type="personal">
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<mods:roleTerm id="A9B3FF7E2326D8C9A75949B966C549C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DE451E03C8577724D08561F1A1F49EBC">Blagoev, Gergin</mods:namePart>
<mods:affiliation id="D828EF6774B22D60B58FE8332AFCDB8F">Centre for Biodiversity Genomics, University of Guelph, Guelph, Ontario, Canada.</mods:affiliation>
</mods:name>
<mods:name id="CB5E52DFC83B368D7E22AAC71F0B43ED" type="personal">
<mods:role id="E122007B717C2CCC1BDFB558B98CD2C0">
<mods:roleTerm id="7BBF4B956B4F6A7CDEC0DE04CC1A0C1F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D99A5A00B6FEF0A8924320488845B44">Deltshev, Christo</mods:namePart>
<mods:affiliation id="67423A5393AA5D2359FCE9115258DD63">National Museum of Natural History, Bulgarian Academy of Sciences, 1000 Sofia, Bulgaria.</mods:affiliation>
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<mods:typeOfResource id="CF895C7A98D8D05D26BE88D9C2B3E6A1">text</mods:typeOfResource>
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<mods:titleInfo id="9EBAF333A269A27D1F0BF41302EC2E24">
<mods:title id="F843E6A1F63951395D6AC1D7DEF24C47">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="463102DA7309DD1DEBCB7E419A596498">
<mods:date id="2D415390F47591D96B63D8CFA6838D69">2021</mods:date>
<mods:detail id="17580F73868326A751819EDE4ED88082" type="pubDate">
<mods:number id="D7F0419BF49887519F7F038C91B73D56">2021-06-10</mods:number>
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<mods:detail id="B7DBFEA133FCCE3F2CC573CFBE82C7DE" type="volume">
<mods:number id="0CA814EF7047BFE261D71C476AAFC79A">4984</mods:number>
</mods:detail>
<mods:detail id="FF39D5FB314A1010768498CBD52B065C" type="issue">
<mods:number id="4C4CCC84AB5C7A1FA0AB9FBFBC6C52DA">1</mods:number>
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<mods:classification id="DDDB3E54FD23279FB76BAB7A6B7320D7">journal article</mods:classification>
<mods:identifier id="AD417AD17F8A2D261D89B042FAB5752D" type="CLB-Dataset">4940</mods:identifier>
<mods:identifier id="BEC9B7086CA1B17B798A97AE966E0C98" type="DOI">10.11646/zootaxa.4984.1.18</mods:identifier>
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<treatment id="6667B114FFA65A34FF535F6D3F0C3F34" ID-DOI="http://doi.org/10.5281/zenodo.5195560" ID-GBIF-Taxon="184606399" ID-Zenodo-Dep="5195560" LSID="urn:lsid:plazi:treatment:6667B114FFA65A34FF535F6D3F0C3F34" httpUri="http://treatment.plazi.org/id/6667B114FFA65A34FF535F6D3F0C3F34" lastPageNumber="235" pageId="7" pageNumber="235">
<subSubSection id="A6D45389FFA65A34FF535F6D38113E7F" box="[151,614,1561,1588]" pageId="7" pageNumber="235" type="nomenclature">
<paragraph id="EE710002FFA65A34FF535F6D38113E7F" blockId="7.[151,614,1561,1624]" box="[151,614,1561,1588]" pageId="7" pageNumber="235">
<taxonomicName id="29CE7B81FFA65A34FF535F6D38113E7F" ID-CoL="95G4X" authority="(Spassky, 1940)" baseAuthorityName="Spassky" baseAuthorityYear="1940" box="[151,614,1561,1588]" class="Arachnida" family="Gnaphosidae" genus="Leptopilos" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="235" phylum="Arthropoda" rank="species" species="memorialis">
<emphasis id="DCBADC10FFA65A34FF535F6D3BE93E78" box="[151,414,1562,1588]" italics="true" pageId="7" pageNumber="235">Leptopilos memorialis</emphasis>
<emphasis id="DCBADC10FFA65A34FE625F6E38113E7F" bold="true" box="[422,614,1561,1587]" pageId="7" pageNumber="235">(Spassky, 1940)</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A6D45389FFA65A34FF535F493B783E14" box="[151,271,1598,1624]" pageId="7" pageNumber="235" type="description">
<paragraph id="EE710002FFA65A34FF535F493B783E14" blockId="7.[151,614,1561,1624]" box="[151,271,1598,1624]" pageId="7" pageNumber="235">
<figureCitation id="76F51C87FFA65A34FF535F493B783E14" box="[151,271,1598,1624]" captionStart="FIGURES 7" captionStartId="8.[151,264,1240,1264]" captionTargetBox="[271,1316,198,1205]" captionTargetId="figure-17@8.[246,1341,181,1216]" captionTargetPageId="8" captionText="FIGURES 7A–E. Leptopilos memorialis, male (A–C) and female (D–E).A–C Left palp (A prolateral, B ventral, C retrolateral). D Epigyne, macerated, ventral. E Vulva, dorsal. C—conductor, CO—copulatory opening, Em—embolus, FD—fertilization duct, Ho—hood of scape, MA—median apophysis, RTA—retrolateral tibial apophysis, S—spermatheca" figureDoi="http://doi.org/10.5281/zenodo.4927044" httpUri="https://zenodo.org/record/4927044/files/figure.png" pageId="7" pageNumber="235">Figs 7A–E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A6D45389FFA65A34FF535FF238063F40" pageId="7" pageNumber="235" type="materials_examined">
<paragraph id="EE710002FFA65A34FF535FF238063F40" blockId="7.[151,1436,1669,1912]" pageId="7" pageNumber="235">
<emphasis id="DCBADC10FFA65A34FF535FF23B713ED3" bold="true" box="[151,262,1669,1695]" pageId="7" pageNumber="235">Material.</emphasis>
<materialsCitation id="5EA60A5FFFA65A34FECB5FF13E833E88" ID-GBIF-Occurrence="3336412305" collectingDate="2017-06-03" collectorName="M. Naumova" elevation="645" latitude="42.7882" location="Utaynika Bog" longLatPrecision="6" longitude="23.4812" municipality="Western Stara Planina Mts" pageId="7" pageNumber="235" specimenCount="1" specimenCount-female="1" stateProvince="Sofiya-Grad">
<specimenCount id="F8C8CB8BFFA65A34FECB5FF13B4D3EEC" box="[271,314,1670,1696]" pageId="7" pageNumber="235" type="female">1 ♀</specimenCount>
(IBER),
<collectingMunicipality id="0E159A78FFA65A34FE615FF138A23EEC" box="[421,725,1670,1696]" pageId="7" pageNumber="235">Western Stara Planina Mts</collectingMunicipality>
,
<collectingRegion id="2C0ACEE0FFA65A34FD265FF1396A3EEC" box="[738,797,1670,1696]" country="Bulgaria" name="Sofiya-Grad" pageId="7" pageNumber="235">Sofia</collectingRegion>
(Kremikovtsi ward), near
<location id="EB1156D9FFA65A34FB895FF13E9A3EEC" LSID="urn:lsid:plazi:treatment:6667B114FFA65A34FF535F6D3F0C3F34:EB1156D9FFA65A34FB895FF13E9A3EEC" box="[1101,1261,1670,1696]" latitude="42.7882" longLatPrecision="6" longitude="23.4812" municipality="Western Stara Planina Mts" name="Utaynika Bog" pageId="7" pageNumber="235" stateProvince="Sofiya-Grad">Utaynika Bog</location>
,
<geoCoordinate id="8BFA66C5FFA65A34FB3E5FF13F0F3EEC" box="[1274,1400,1669,1696]" degrees="42.7882" direction="north" orientation="latitude" pageId="7" pageNumber="235" precision="5" value="42.7882">N 42.7882</geoCoordinate>
˚,
<geoCoordinate id="8BFA66C5FFA65A34FA4F5FF13A8F3E88" degrees="23.4812" direction="east" orientation="longitude" pageId="7" pageNumber="235" precision="5" value="23.4812">E 23.4812</geoCoordinate>
˚, 1–
<date id="9A7026C2FFA65A34FEE55FDD3BEF3E88" box="[289,408,1706,1732]" pageId="7" pageNumber="235" value="2017-06-03">
<collectingDate id="8A34DF2AFFA65A34FEE55FDD3BEF3E88" box="[289,408,1706,1732]" pageId="7" pageNumber="235" value="2017-06-03">3.06.2017</collectingDate>
</date>
,
<quantity id="2936ADE7FFA65A34FE665FDD3B9D3E88" box="[418,490,1706,1733]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="7" pageNumber="235" unit="m" value="645.0">
<elevation id="65E3E731FFA65A34FE665FDD3B9D3E88" box="[418,490,1706,1733]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="7" pageNumber="235" unit="m" value="645.0">645 m</elevation>
</quantity>
,
<collectorName id="433B65D4FFA65A34FE315FDD38FF3E88" box="[501,648,1706,1732]" pageId="7" pageNumber="235">M. Naumova</collectorName>
leg., under stone in open habitat with scarce vegetation
</materialsCitation>
;
<materialsCitation id="5EA60A5FFFA65A34FB3A5FDD38063F40" ID-GBIF-Occurrence="3336412333" collectingDate="2020-05-30" collectorName="T. Ljubomirov" elevation="301" latitude="42.1255" location="Sinitovo village" longLatPrecision="6" longitude="24.3761" municipality="Besapari" pageId="7" pageNumber="235" specimenCount="2" specimenCount-male="2" stateProvince="Sofiya-Grad">
<specimenCount id="F8C8CB8BFFA65A34FB3A5FDD3F4A3E88" box="[1278,1341,1706,1732]" pageId="7" pageNumber="235" type="male">2 ♂♂</specimenCount>
(IBER),
<collectingMunicipality id="0E159A78FFA65A34FF535FB93A8E3EA4" box="[151,249,1742,1768]" pageId="7" pageNumber="235">Besapari</collectingMunicipality>
hills,
<location id="EB1156D9FFA65A34FEF95FB93B863EA4" LSID="urn:lsid:plazi:treatment:6667B114FFA65A34FF535F6D3F0C3F34:EB1156D9FFA65A34FEF95FB93B863EA4" box="[317,497,1742,1768]" latitude="42.1255" longLatPrecision="6" longitude="24.3761" municipality="Besapari" name="Sinitovo village" pageId="7" pageNumber="235" stateProvince="Sofiya-Grad">Sinitovo village</location>
,
<geoCoordinate id="8BFA66C5FFA65A34FE385FB938003EA4" box="[508,631,1741,1768]" degrees="42.1255" direction="north" orientation="latitude" pageId="7" pageNumber="235" precision="5" value="42.1255">N 42.1255</geoCoordinate>
˚,
<geoCoordinate id="8BFA66C5FFA65A34FD4C5FB939773EA4" box="[648,768,1741,1768]" degrees="24.3761" direction="east" orientation="longitude" pageId="7" pageNumber="235" precision="5" value="24.3761">E 24.3761</geoCoordinate>
˚,
<quantity id="2936ADE7FFA65A34FCD55FB9392E3EA4" box="[785,857,1742,1768]" metricMagnitude="2" metricUnit="m" metricValue="3.01" pageId="7" pageNumber="235" unit="m" value="301.0">
<elevation id="65E3E731FFA65A34FCD55FB9392E3EA4" box="[785,857,1742,1768]" metricMagnitude="2" metricUnit="m" metricValue="3.01" pageId="7" pageNumber="235" unit="m" value="301.0">301 m</elevation>
</quantity>
, 29–
<date id="9A7026C2FFA65A34FC4A5FB93E783EA5" box="[910,1039,1742,1769]" pageId="7" pageNumber="235" value="2020-05-30">
<collectingDate id="8A34DF2AFFA65A34FC4A5FB93E783EA5" box="[910,1039,1742,1769]" pageId="7" pageNumber="235" value="2020-05-30">30.05.2020</collectingDate>
</date>
,
<collectingMethod id="378F7815FFA65A34FBDE5FB93ECE3EA4" box="[1050,1209,1742,1768]" pageId="7" pageNumber="235">Moericke trap</collectingMethod>
,
<collectorName id="433B65D4FFA65A34FB005FB93F113EA4" box="[1220,1382,1742,1768]" pageId="7" pageNumber="235">T. Ljubomirov</collectorName>
leg., open rocky habitat with scarce vegetation.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A6D45389FFA65A34FF035E613F0C3F34" pageId="7" pageNumber="235" type="discussion">
<paragraph id="EE710002FFA65A34FF035E613F0C3F34" blockId="7.[151,1436,1669,1912]" pageId="7" pageNumber="235">
<emphasis id="DCBADC10FFA65A34FF035E613B4C3F7C" bold="true" box="[199,315,1814,1840]" pageId="7" pageNumber="235">Remarks.</emphasis>
Rare species known from
<collectingCountry id="96D94092FFA65A34FDA85E6138C93F7C" box="[620,702,1814,1840]" name="Greece" pageId="7" pageNumber="235">Greece</collectingCountry>
,
<collectingCountry id="96D94092FFA65A34FD0E5E6139503F7C" box="[714,807,1814,1840]" name="Ukraine" pageId="7" pageNumber="235">Ukraine</collectingCountry>
,
<collectingCountry id="96D94092FFA65A34FCF05E6139083F7C" box="[820,895,1814,1840]" name="Russia" pageId="7" pageNumber="235">Russia</collectingCountry>
(Europe to Central Asia),
<collectingCountry id="96D94092FFA65A34FB685E613F453F7C" box="[1196,1330,1814,1840]" name="Kazakhstan" pageId="7" pageNumber="235">Kazakhstan</collectingCountry>
,
<collectingCountry id="96D94092FFA65A34FAF95E613FEB3F7C" box="[1341,1436,1814,1840]" name="Pakistan" pageId="7" pageNumber="235">Pakistan</collectingCountry>
and
<collectingCountry id="96D94092FFA65A34FF015E4D3B453F18" box="[197,306,1850,1876]" name="Mongolia" pageId="7" pageNumber="235">Mongolia</collectingCountry>
(
<bibRefCitation id="8A5F7DF3FFA65A34FE855E4D38033F18" author="World Spider Catalog" box="[321,628,1850,1876]" pageId="7" pageNumber="235" refId="ref14500" refString="World Spider Catalog (2021) World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 28 February 2021) https: // doi. org / 10.24436 / 2" type="book" year="2021">World Spider Catalog 2021</bibRefCitation>
). First record of the genus in
<collectingCountry id="96D94092FFA65A34FC775E4D3E603F18" box="[947,1047,1850,1876]" name="Bulgaria" pageId="7" pageNumber="235">Bulgaria</collectingCountry>
. New photographs of the male palp (
<figureCitation id="76F51C87FFA65A34FF5B5E293B6A3F34" box="[159,285,1886,1912]" captionStart="FIGURES 7" captionStartId="8.[151,264,1240,1264]" captionTargetBox="[271,1316,198,1205]" captionTargetId="figure-17@8.[246,1341,181,1216]" captionTargetPageId="8" captionText="FIGURES 7A–E. Leptopilos memorialis, male (A–C) and female (D–E).A–C Left palp (A prolateral, B ventral, C retrolateral). D Epigyne, macerated, ventral. E Vulva, dorsal. C—conductor, CO—copulatory opening, Em—embolus, FD—fertilization duct, Ho—hood of scape, MA—median apophysis, RTA—retrolateral tibial apophysis, S—spermatheca" figureDoi="http://doi.org/10.5281/zenodo.4927044" httpUri="https://zenodo.org/record/4927044/files/figure.png" pageId="7" pageNumber="235">Figs 7A–C</figureCitation>
) with the specific complex of tegular apophyses and the female genitalia (
<figureCitation id="76F51C87FFA65A34FB985E293EAF3F34" box="[1116,1240,1886,1912]" captionStart="FIGURES 7" captionStartId="8.[151,264,1240,1264]" captionTargetBox="[271,1316,198,1205]" captionTargetId="figure-17@8.[246,1341,181,1216]" captionTargetPageId="8" captionText="FIGURES 7A–E. Leptopilos memorialis, male (A–C) and female (D–E).A–C Left palp (A prolateral, B ventral, C retrolateral). D Epigyne, macerated, ventral. E Vulva, dorsal. C—conductor, CO—copulatory opening, Em—embolus, FD—fertilization duct, Ho—hood of scape, MA—median apophysis, RTA—retrolateral tibial apophysis, S—spermatheca" figureDoi="http://doi.org/10.5281/zenodo.4927044" httpUri="https://zenodo.org/record/4927044/files/figure.png" pageId="7" pageNumber="235">Figs 7D–E</figureCitation>
) are provided.
</paragraph>
</subSubSection>
</treatment>
</document>
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