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<document ID-DOI="http://doi.org/10.5281/zenodo.3748525" ID-GBIF-Dataset="6384b520-7e9f-4874-a414-76c2e9b01d74" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="3748525" approvalRequired="1" checkinTime="1586358796837" checkinUser="plazi" docAuthor="Burgin, Connor" docDate="2019" docId="885887A2FFE58A03F8ADFC69F798D4E7" docLanguage="en" docName="hbmw_9_Rhinolophidae.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docStyle="DocumentStyle{}" docTitle="Rhinolophus andamanensis Dobson 1872" docType="treatment" docVersion="20" lastPageNumber="321" masterDocId="7461FFDAFFCF8A29FFCCFFA1FF85D963" masterDocTitle="Rhinolophidae" masterLastPageNumber="332" masterPageNumber="280" pageNumber="321" updateTime="1658949228820" updateUser="felipe">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rhinolophidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burgin, Connor</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Wilson, Don E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Mittermeier, Russell A.</mods:namePart>
</mods:name>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>280</mods:start>
<mods:end>332</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3748525</mods:identifier>
<mods:identifier type="GBIF-Dataset">6384b520-7e9f-4874-a414-76c2e9b01d74</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3748525</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3808978" ID-GBIF-Taxon="163549989" ID-Zenodo-Dep="3808978" LSID="urn:lsid:plazi:treatment:885887A2FFE58A03F8ADFC69F798D4E7" httpUri="http://treatment.plazi.org/id/885887A2FFE58A03F8ADFC69F798D4E7" lastPageNumber="321" pageId="42" pageNumber="321">
<subSubSection box="[1889,2716,968,1029]" pageId="42" pageNumber="321" type="vernacular_names">
<paragraph blockId="42.[1889,3496,968,1029]" box="[1889,2716,968,1029]" pageId="42" pageNumber="321">
<figureCitation box="[1889,1959,968,1029]" captionStart="On" captionStartId="38.[159,201,4464,4491]" captionTargetBox="[116,3439,160,4676]" captionTargetPageId="37" captionText="On following pages: 78. New Guinea Broad-eared Horseshoe Bat (Rhinolophus euryotis); 79. Sulawesi Broad-eared Horseshoe Bat (Rhinolophus tatah; 80. Poso Horseshoe Bat (Rhinolophus belligerator); 81. Bornean Woolly Horseshoe Bat (Rhinolophus proconsulis); 82. Arcuate Horseshoe Bat (Rhinolophus arcuatus); 83. Philippine Forest Horseshoe Bat (Rhinolophus inops); 84. Small Rufous Horseshoe Bat (Rhinolophus subrufus); 85. Large Rufous Horseshoe Bat (Rhinolophus rufus); 86. Intermediate Horseshoe Bat (Rhinolophus affinis); 87. Homfray's Horseshoe Bat (Rhinolophus andamanensis); 88. Croslet Horseshoe Bat (Rhinolophus coelophyllus); 89. Shamel's Horseshoe Bat (Rhinolophus shameli); 90. Vietnamese Brown Horseshoe Bat (Rhinolophus microglobosus); 91. Lesser Brown Horseshoe Bat (Rhinolophus stheno)." figureDoi="http://doi.org/10.5281/zenodo.3748533" httpUri="https://zenodo.org/record/3748533/files/figure.png" pageId="42" pageNumber="321">87</figureCitation>
. Homfrays Horseshoe
<emphasis bold="true" box="[2621,2716,968,1029]" pageId="42" pageNumber="321">Bat</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[2781,3496,968,1029]" pageId="42" pageNumber="321" type="nomenclature">
<paragraph blockId="42.[1889,3496,968,1029]" box="[2781,3496,968,1029]" pageId="42" pageNumber="321">
<taxonomicName authorityName="Dobson" authorityYear="1872" box="[2781,3496,968,1029]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="species" species="andamanensis">
<emphasis box="[2781,3496,968,1029]" italics="true" pageId="42" pageNumber="321">Rhinolophus andamanensis</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="321" type="vernacular_names">
<paragraph blockId="42.[1888,3308,1047,1127]" box="[1889,3308,1047,1074]" pageId="42" pageNumber="321">
French: Rhinolophe
<emphasis box="[2164,2197,1047,1074]" italics="true" pageId="42" pageNumber="321">de</emphasis>
Homfray
<emphasis box="[2331,2342,1047,1074]" italics="true" pageId="42" pageNumber="321">I</emphasis>
German: Andamanen-Hufeisennase /
<emphasis bold="true" box="[2868,2994,1047,1074]" pageId="42" pageNumber="321">Spanish:</emphasis>
Herradura
<emphasis box="[3147,3181,1047,1074]" italics="true" pageId="42" pageNumber="321">de</emphasis>
Homfray
</paragraph>
<paragraph blockId="42.[1888,3308,1047,1127]" box="[1888,2755,1100,1127]" pageId="42" pageNumber="321">
<emphasis bold="true" box="[1888,2222,1100,1127]" pageId="42" pageNumber="321">Other common names:</emphasis>
<emphasis box="[2231,2360,1100,1127]" italics="true" pageId="42" pageNumber="321">Andaman</emphasis>
Intermediate Horseshoe Bat
</paragraph>
</subSubSection>
<paragraph blockId="42.[2712,3517,1199,1770]" pageId="42" pageNumber="321">
<subSubSection pageId="42" pageNumber="321" type="reference_group">
Taxonomy.
<taxonomicName authority="Dobson, 1872" authorityName="Dobson" authorityYear="1872" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="subSpecies" species="affinis" subSpecies="andamanensis">
<emphasis box="[2954,3517,1199,1238]" italics="true" pageId="42" pageNumber="321">
Rhinolophus affinis
<emphasis box="[2715,2797,1250,1289]" italics="true" pageId="42" pageNumber="321">andamanensis</emphasis>
</emphasis>
Dobson, 1872
</taxonomicName>
,
</subSubSection>
<subSubSection pageId="42" pageNumber="321" type="materials_examined">
<materialsCitation ID-GBIF-Occurrence="2597534029" country="India" location="Andaman Islands" pageId="42" pageNumber="321" specimenCount="1" typeStatus="type">
<location LSID="urn:lsid:plazi:treatment:885887A2FFE58A03F8ADFC69F798D4E7:052E606FFFE58A03F388FB43F226DC6A" box="[3140,3491,1250,1289]" country="India" name="Andaman Islands" pageId="42" pageNumber="321">Andaman Islands</location>
,”
<collectingCountry box="[2713,2816,1306,1345]" name="India" pageId="42" pageNumber="321">India</collectingCountry>
</materialsCitation>
.
</subSubSection>
</paragraph>
<subSubSection pageId="42" pageNumber="321" type="discussion">
<paragraph blockId="42.[2712,3517,1199,1770]" pageId="42" pageNumber="321">
<taxonomicName authorityName="Dobson" authorityYear="1872" box="[2714,3207,1359,1398]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="species" species="andamanensis">
<emphasis box="[2714,3207,1359,1398]" italics="true" pageId="42" pageNumber="321">Rhinolophus andamanensis</emphasis>
</taxonomicName>
is included in the
<taxonomicName authorityName="Temminck" authorityYear="1835" box="[2795,2926,1412,1451]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="species" species="euryotis">
<emphasis box="[2795,2926,1412,1451]" italics="true" pageId="42" pageNumber="321">euryotis</emphasis>
</taxonomicName>
species group and is sister to
<emphasis box="[2713,2881,1465,1504]" italics="true" pageId="42" pageNumber="321">
<taxonomicName authorityName="Horsfield" authorityYear="1823" box="[2713,2876,1465,1504]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="species" species="affinis">R affinis</taxonomicName>
.
</emphasis>
It was
<emphasis box="[3005,3248,1465,1504]" italics="true" pageId="42" pageNumber="321">only recently</emphasis>
recognized as a species distinct from
<taxonomicName authorityName="Horsfield" authorityYear="1823" box="[3145,3309,1518,1558]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="species" species="affinis">
<emphasis box="[3145,3309,1518,1558]" italics="true" pageId="42" pageNumber="321">R affinis</emphasis>
</taxonomicName>
by. Srinivasulu and coworkers in 2019, based on genetic, morphometric, and acoustic data. Monotypic.
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="321" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.3750084" ID-Zenodo-Dep="3750084" httpUri="https://zenodo.org/record/3750084/files/figure.png" inLine="true" pageId="42" pageNumber="321" targetBox="[1884,2682,1204,1763]" targetPageId="42">
<paragraph blockId="42.[2712,3517,1199,1770]" lastBlockId="42.[1879,3513,1783,3460]" pageId="42" pageNumber="321">
<emphasis bold="true" box="[2712,2950,1731,1770]" pageId="42" pageNumber="321">Distribution.</emphasis>
Andaman Is (North Andaman, Interview, Middle Andaman, Baratang, South Andaman, and Little Andaman).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="42" pageNumber="321" type="description">
<paragraph blockId="42.[1879,3513,1783,3460]" pageId="42" pageNumber="321">
Descriptive
<emphasis bold="true" box="[2110,2221,1835,1874]" pageId="42" pageNumber="321">notes.</emphasis>
Head-body 58-
<quantity box="[2519,2626,1835,1874]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="321" unit="mm" value="2.0">2 mm</quantity>
(average), tail 17-
<quantity box="[2964,3142,1835,1874]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.1" metricValueMin="0.9" pageId="42" pageNumber="321" unit="mm" value="20.0" valueMax="31.0" valueMin="9.0">9—31 mm</quantity>
, ear 18-1-23-
<quantity box="[3391,3503,1835,1874]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="42" pageNumber="321" unit="mm" value="9.0">9 mm</quantity>
, hindfoot 9-7—14-
<quantity box="[2209,2325,1887,1926]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="42" pageNumber="321" unit="mm" value="7.0">7 mm</quantity>
, forearm 46-7—56-
<quantity box="[2681,2796,1887,1926]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="42" pageNumber="321" unit="mm" value="6.0">6 mm</quantity>
. Dorsal pelage is brown to bright orange, with ventral pelage very similar. Ears are small. Noseleaf has parallel-sided lancet with pointed tip; connecting process is rounded and sparsely haired; sella is slightly concave and pandurate in shape; horseshoe is
<emphasis box="[2762,2932,2049,2088]" italics="true" pageId="42" pageNumber="321">relatively</emphasis>
broad and covers muzzle, with well-defined median emargination, and distinct lateral leaflets. Lower lip has three mental grooves. Baculum is
<quantity box="[2420,2572,2155,2194]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="42" pageNumber="321" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2—3 mm</quantity>
long, has deeply notched ventral margin of basal cone but is less notched on dorsal margin; it has smaller notches on lateral sides; shaft is roughly circular in cross section and bends downward toward tip. Skull is robust; sagittal
<emphasis box="[1970,2060,2314,2353]" italics="true" pageId="42" pageNumber="321">crest</emphasis>
is well developed; interorbital region is broad; anterior median swellings are comparatively less inflated; posterior swellings are well defined; zygomatic arches are robust; canines are robust. P2 is small and within tooth row; P3 is small and extruded from tooth row, allowing P2 and P4 to touch.
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="321" type="biology_ecology">
<paragraph blockId="42.[1879,3513,1783,3460]" box="[1885,2720,2528,2567]" pageId="42" pageNumber="321">Habitat. Forested habitats across the islands.</paragraph>
<paragraph blockId="42.[1879,3513,1783,3460]" box="[1885,2560,2581,2620]" pageId="42" pageNumber="321">
<emphasis bold="true" box="[1885,2067,2581,2620]" pageId="42" pageNumber="321">Food and</emphasis>
Feeding. No information.
</paragraph>
<paragraph blockId="42.[1879,3513,1783,3460]" box="[1885,2390,2634,2673]" pageId="42" pageNumber="321">
<emphasis bold="true" box="[1885,2068,2634,2673]" pageId="42" pageNumber="321">Breeding.</emphasis>
No information.
</paragraph>
<paragraph blockId="42.[1879,3513,1783,3460]" pageId="42" pageNumber="321">
<emphasis bold="true" box="[1883,2203,2687,2726]" pageId="42" pageNumber="321">Activity patterns.</emphasis>
Homfrays Horseshoe Bat roosts in limestone caves, forest caves, and sometimes holes and tree hollows. Calls have a FM/CF/FM shape with a peak F recorded at 56-4-58-5 kHz (mean 57-6 kHz).
</paragraph>
<paragraph blockId="42.[1879,3513,1783,3460]" pageId="42" pageNumber="321">
<emphasis bold="true" box="[1885,2832,2846,2885]" pageId="42" pageNumber="321">Movements, Home range and Social organization.</emphasis>
Homfrays Horseshoe Bat roosts in large colonies and has been observed roosting with Andaman Horseshoe Bats (.
<emphasis italics="true" pageId="42" pageNumber="321">
<taxonomicName authorityName="K Andersen" authorityYear="1906" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="species" species="cognatus">cognatus</taxonomicName>
),
</emphasis>
Diadem Leaf-nosed Bats (Hipposideros
<emphasis box="[2781,2954,2951,2990]" italics="true" pageId="42" pageNumber="321">diadema),</emphasis>
Andersens Leaf-nosed Bats (
<emphasis box="[1904,2115,3003,3042]" italics="true" pageId="42" pageNumber="321">H. gentilis),</emphasis>
Grand Leaf-nosed Bats (
<emphasis box="[2602,2821,3003,3042]" italics="true" pageId="42" pageNumber="321">H. grandis),</emphasis>
and Horsfields
<taxonomicName box="[3144,3267,3003,3042]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="genus">Myotis</taxonomicName>
(
<emphasis italics="true" pageId="42" pageNumber="321">
<taxonomicName baseAuthorityName="Temminck" baseAuthorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="species" species="horsfieldii">Myotis horsfieldii</taxonomicName>
).
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="321" type="conservation">
<paragraph blockId="42.[1879,3513,1783,3460]" pageId="42" pageNumber="321">
<emphasis bold="true" box="[1885,2372,3110,3149]" pageId="42" pageNumber="321">Status and Conservation.</emphasis>
<collectingRegion box="[2392,2466,3110,3149]" country="Cameroon" name="North" pageId="42" pageNumber="321">Not</collectingRegion>
assessed as a separate species on
<emphasis box="[3148,3213,3110,3149]" italics="true" pageId="42" pageNumber="321">The</emphasis>
IUCN
<emphasis box="[3349,3507,3110,3149]" italics="true" pageId="42" pageNumber="321">ed List,</emphasis>
where it is included under the Intermediate Horseshoe Bat (
<emphasis box="[3057,3235,3162,3202]" italics="true" pageId="42" pageNumber="321">
<taxonomicName authorityName="Horsfield" authorityYear="1823" box="[3057,3224,3162,3202]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="42" pageNumber="321" phylum="Chordata" rank="species" species="affinis">R. affinis</taxonomicName>
)
</emphasis>
as Least Concern. Homfrays Horseshoe Bat has a restricted range, but appears to be
<emphasis box="[3339,3510,3216,3255]" italics="true" pageId="42" pageNumber="321">relatively</emphasis>
common throughout the Andaman Islands. Further studies into the species ecology and potential threats are needed.
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="321" type="reference_group">
<paragraph blockId="42.[1879,3513,1783,3460]" pageId="42" pageNumber="321">
<emphasis bold="true" box="[1885,2093,3380,3407]" pageId="42" pageNumber="321">Bibliography.</emphasis>
Srinivasulu, Srinivasulu, Srinivasulu,
<emphasis box="[2658,2725,3380,3407]" italics="true" pageId="42" pageNumber="321">Gopi</emphasis>
et al. (2017), Srinivasulu, Srinivasulu, Srinivasulu &amp; Jones (2019).
</paragraph>
</subSubSection>
</treatment>
</document>