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<document ID-DOI="http://doi.org/10.5281/zenodo.3748525" ID-GBIF-Dataset="6384b520-7e9f-4874-a414-76c2e9b01d74" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="3748525" approvalRequired="1" checkinTime="1586358796837" checkinUser="plazi" docAuthor="Burgin, Connor" docDate="2019" docId="885887A2FFDB8A3DFF5CF645F772DBFB" docLanguage="en" docName="hbmw_9_Rhinolophidae.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docStyle="DocumentStyle{}" docTitle="Rhinolophus hilli Aellen 1973" docType="treatment" docVersion="21" lastPageNumber="299" masterDocId="7461FFDAFFCF8A29FFCCFFA1FF85D963" masterDocTitle="Rhinolophidae" masterLastPageNumber="332" masterPageNumber="280" pageNumber="299" updateTime="1658949228820" updateUser="felipe">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rhinolophidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burgin, Connor</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Wilson, Don E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Mittermeier, Russell A.</mods:namePart>
</mods:name>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>280</mods:start>
<mods:end>332</mods:end>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3748525</mods:identifier>
<mods:identifier type="GBIF-Dataset">6384b520-7e9f-4874-a414-76c2e9b01d74</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3748525</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3808942" ID-GBIF-Taxon="163549977" ID-Zenodo-Dep="3808942" LSID="urn:lsid:plazi:treatment:885887A2FFDB8A3DFF5CF645F772DBFB" httpUri="http://treatment.plazi.org/id/885887A2FFDB8A3DFF5CF645F772DBFB" lastPageNumber="299" pageId="20" pageNumber="299">
<subSubSection box="[144,831,2532,2593]" pageId="20" pageNumber="299" type="vernacular_names">
<paragraph blockId="20.[142,1338,2531,2641]" box="[144,831,2532,2593]" pageId="20" pageNumber="299">
<figureCitation box="[144,216,2532,2593]" captionStart="On" captionStartId="14.[164,207,4337,4364]" captionTargetBox="[105,3488,151,4690]" captionTargetPageId="13" captionText="On following pages: 27. African Forest Horseshoe Bat (Rhinolophus silvestris); 28. Upland Horseshoe Bat (Rhinolophus hillorum); 29. Sakeji Horseshoe Bat (Rhinolophus sakejiensis); 30. Bokhara Horseshoe Bat (Rhinolophus bocharicus); 31. Greater Horseshoe Bat (Rhinolophus ferrumequinum); 32. Geoffrey's Horseshoe Bat (Rhinolophus clivosus); 33. Greater Japanese Horseshoe Bat (Rhinolophus nippon); 34. Horacek's Horseshoe Bat (Rhinolophus horaceki); 35. Maclaud's Horseshoe Bat (Rhinolophus maclaudi); 36. Ziama Horseshoe Bat (Rhinolophus ziama); 37. Hill's Horseshoe Bat (Rhinolophus hilli); 38. Kahuzi Horseshoe Bat (Rhinolophus kahuzi); 39. Willard's Horseshoe Bat (Rhinolophus willardi); 40. Ruwenzori Horseshoe Bat (Rhinolophus ruwenzorii); 41. Chinese Horseshoe Bat (Rhinolophus xinanzhongguoensis); 42. Big-eared Horseshoe Bat (Rhinolophus macrotis); 43. Osgood's Horseshoe Bat (Rhinolophus osgoodi); 44. Allen's Horseshoe Bat (Rhinolophus episcopus); 45. Thai Horseshoe Bat (Rhinolophus siamensis); 46, Schnitzler's Horseshoe Bat (Rhinolophus schnitzten); M. King Horseshoe Bat (Rhinolophus rex); 48. Marshalls Horseshoe Bat (Rhinolophus marshalli)." figureDoi="http://doi.org/10.5281/zenodo.3748529" httpUri="https://zenodo.org/record/3748529/files/figure.png" pageId="20" pageNumber="299">37</figureCitation>
. Hills Horseshoe Bat
</paragraph>
</subSubSection>
<subSubSection box="[895,1338,2532,2593]" pageId="20" pageNumber="299" type="nomenclature">
<paragraph blockId="20.[142,1338,2531,2641]" box="[895,1338,2532,2593]" pageId="20" pageNumber="299">
<taxonomicName authorityName="Aellen" authorityYear="1973" box="[895,1338,2532,2593]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="299" phylum="Chordata" rank="species" species="hilli">
<emphasis box="[895,1338,2532,2593]" italics="true" pageId="20" pageNumber="299">Rhinolophus hilli</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[143,1293,2610,2637]" pageId="20" pageNumber="299" type="vernacular_names">
<paragraph blockId="20.[142,1338,2531,2641]" box="[143,1293,2610,2637]" pageId="20" pageNumber="299">
French: Rhinolophe de
<emphasis box="[466,506,2610,2637]" italics="true" pageId="20" pageNumber="299">Hill</emphasis>
/
<emphasis bold="true" box="[533,658,2610,2637]" pageId="20" pageNumber="299">German:</emphasis>
Hill-Hufeisennase / Spanish: Herradura de
<emphasis box="[1253,1293,2610,2637]" italics="true" pageId="20" pageNumber="299">Hill</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[973,1769,2708,2747]" pageId="20" pageNumber="299" type="reference_group">
<paragraph blockId="20.[973,1775,2708,3280]" box="[973,1769,2708,2747]" pageId="20" pageNumber="299">
Taxonomy.
<taxonomicName authority="Aellen, 1973" authorityName="Aellen" authorityYear="1973" box="[1203,1762,2708,2747]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="299" phylum="Chordata" rank="species" species="hilli">
<emphasis box="[1203,1507,2708,2747]" italics="true" pageId="20" pageNumber="299">Rhinolophus hilli</emphasis>
Aellen, 1973
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="299" type="materials_examined">
<paragraph blockId="20.[973,1775,2708,3280]" pageId="20" pageNumber="299">
<materialsCitation ID-GBIF-Occurrence="3864669315" country="Rwanda" elevation="2300" location="Uwinka, Prefecture]. Cyangugu" pageId="20" pageNumber="299" specimenCount="1" typeStatus="type">
<location LSID="urn:lsid:plazi:treatment:885887A2FFDB8A3DFF5CF645F772DBFB:052E606FFFDB8A3DFC13F56BF9D8D392" box="[991,1629,2762,2802]" country="Rwanda" name="Uwinka, Prefecture]. Cyangugu" pageId="20" pageNumber="299">Uwinka, Prefecture]. Cyangugu</location>
,
<collectingCountry name="Rwanda" pageId="20" pageNumber="299">Rwanda</collectingCountry>
, alt.
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<elevation box="[1113,1259,2815,2854]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="299" unit="m" value="2300.0">2300 m</elevation>
</quantity>
</materialsCitation>
.”
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="299" type="discussion">
<paragraph blockId="20.[973,1775,2708,3280]" pageId="20" pageNumber="299">
<taxonomicName authorityName="Aellen" authorityYear="1973" box="[975,1272,2869,2908]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="299" phylum="Chordata" rank="species" species="hilli">
<emphasis box="[975,1272,2869,2908]" italics="true" pageId="20" pageNumber="299">Rhinolophus hilli</emphasis>
</taxonomicName>
is in the
<taxonomicName authorityName="Pousargues" authorityYear="1897" box="[1471,1627,2869,2908]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="299" phylum="Chordata" rank="species" species="maclaudi">
<emphasis box="[1471,1627,2869,2908]" italics="true" pageId="20" pageNumber="299">maclaudi</emphasis>
</taxonomicName>
species group and was considered a subspecies of
<taxonomicName authorityName="Pousargues" authorityYear="1897" box="[975,1186,2975,3014]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="299" phylum="Chordata" rank="species" species="maclaudi">
<emphasis box="[975,1186,2975,3014]" italics="true" pageId="20" pageNumber="299">R maclaudi</emphasis>
</taxonomicName>
and
<emphasis box="[1281,1770,2975,3015]" italics="true" pageId="20" pageNumber="299">
subsequently
<taxonomicName authorityName="Hill" authorityYear="1942" box="[1534,1764,2975,3015]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="20" pageNumber="299" phylum="Chordata" rank="species" species="ruwenzorii">R ruwenzorii</taxonomicName>
.
</emphasis>
Recent morphological studies support its recognition as a distinct species.
<emphasis box="[1562,1762,3082,3121]" italics="true" pageId="20" pageNumber="299">Monotypic</emphasis>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3749980" ID-Zenodo-Dep="3749980" httpUri="https://zenodo.org/record/3749980/files/figure.png" inLine="true" pageId="20" pageNumber="299" targetBox="[142,942,2711,3272]" targetPageId="20">
<paragraph blockId="20.[973,1775,2708,3280]" pageId="20" pageNumber="299">
Distribution. SW
<collectingCountry box="[1313,1465,3135,3174]" name="Rwanda" pageId="20" pageNumber="299">Rwanda</collectingCountry>
in Uwinka and Ruta Bansugera in Nyungwe Forest National Park.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="20" pageNumber="299" type="description">
<paragraph blockId="20.[144,1775,3292,4664]" pageId="20" pageNumber="299">
Descriptive
<emphasis bold="true" box="[389,503,3292,3331]" pageId="20" pageNumber="299">notes.</emphasis>
<emphasis box="[530,754,3292,3331]" italics="true" pageId="20" pageNumber="299">Head—body</emphasis>
62-
<quantity box="[837,962,3292,3331]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="20" pageNumber="299" unit="mm" value="7.0">7 mm</quantity>
, tail 29'
<quantity box="[1138,1265,3292,3331]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="299" unit="mm" value="3.0">3 mm</quantity>
, ear 28-
<quantity box="[1444,1568,3292,3331]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="20" pageNumber="299" unit="mm" value="7.0">7 mm</quantity>
, hindfoot 12-
<quantity box="[205,325,3347,3386]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="299" unit="mm" value="2.0">2 mm</quantity>
, forearm 54-
<quantity box="[585,700,3347,3386]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="299" unit="mm" value="2.0">2 mm</quantity>
and 54-
<quantity box="[870,987,3347,3386]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="299" unit="mm" value="3.0">3 mm</quantity>
; weight 16-
<quantity box="[1217,1279,3347,3386]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="20" pageNumber="299" unit="g" value="5.0">5 g</quantity>
(all measurements from one specimen
<emphasis box="[443,573,3401,3440]" italics="true" pageId="20" pageNumber="299">except</emphasis>
forearm which refers to two). Dorsal and ventral pelage is dark brown. There is no known orange morph. Males lack axillary tufts. Ears are of medium length (
<emphasis box="[482,503,3505,3544]" italics="true" pageId="20" pageNumber="299">c</emphasis>
.53%
<emphasis box="[616,660,3505,3544]" italics="true" pageId="20" pageNumber="299">of</emphasis>
forearm length), with nine internal folds, and very large antitragus. Noseleaf has subtriangular lancet that is conspicuously taller than sella; connecting process is largely reduced, low, and concave, leaving large depression between sella and lancet; sella is naked, upright, and about parallel to lancet, having concave sides and distinctly broadened and rounded spoon-shaped top; narial lobes at base of sella are
<emphasis box="[623,704,3773,3812]" italics="true" pageId="20" pageNumber="299">very</emphasis>
enlarged, forming nearly circular cup; nostrils are bordered by semicircular raised rims that are parallel to inner cup; and horseshoe is of medium width at c.
<quantity box="[524,668,3879,3918]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="299" unit="mm" value="12.0">12 mm</quantity>
, covers muzzle, has lateral leaflets, and has conspicuous median emargination. Lower lip has conspicuous medial groove and two indistinct lateral grooves.
<emphasis box="[447,567,3986,4025]" italics="true" pageId="20" pageNumber="299">Wings</emphasis>
and uropatagium are dark gray. Skull is large and slender, with weak zygomatic arches (zygomatic width is slightly less than mastoid width); nasal swellings are high domed, with roughly circular chambers when viewed dorsally; frontal depression is very deep; sagittal crest is moderately developed anteriorly and
<emphasis box="[144,275,4197,4236]" italics="true" pageId="20" pageNumber="299">weakly</emphasis>
developed posteriorly; and interpterygoid groove is
<emphasis box="[1286,1367,4197,4236]" italics="true" pageId="20" pageNumber="299">very</emphasis>
deep. P2 is small and completely displaced labially; C1 and P4 nearly touch; P3 is small and slightly to completely displaced labially; and P2 and P4 do
<emphasis box="[980,1047,4305,4344]" italics="true" pageId="20" pageNumber="299">not</emphasis>
touch.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="299" type="biology_ecology">
<paragraph blockId="20.[144,1775,3292,4664]" box="[145,1445,4356,4395]" pageId="20" pageNumber="299">
<emphasis bold="true" box="[145,299,4356,4395]" pageId="20" pageNumber="299">Habitat.</emphasis>
Montane tropical moist forest at elevations of
<quantity box="[1187,1439,4356,4395]" metricMagnitude="3" metricUnit="m" metricValue="2.131" metricValueMax="2.5119999999999996" metricValueMin="1.75" pageId="20" pageNumber="299" unit="m" value="2131.0" valueMax="2512.0" valueMin="1750.0">1750-2512 m</quantity>
.
</paragraph>
<paragraph blockId="20.[144,1775,3292,4664]" box="[145,824,4412,4451]" pageId="20" pageNumber="299">
<emphasis bold="true" box="[145,503,4412,4451]" pageId="20" pageNumber="299">Food and Feeding.</emphasis>
No information.
</paragraph>
<paragraph blockId="20.[144,1775,3292,4664]" pageId="20" pageNumber="299">
Breeding.
<typeStatus box="[350,525,4465,4504]" pageId="20" pageNumber="299">Holotype</typeStatus>
<emphasis box="[542,585,4465,4504]" italics="true" pageId="20" pageNumber="299">of</emphasis>
Hills Horseshoe Bat was a
<emphasis box="[1126,1302,4465,4504]" italics="true" pageId="20" pageNumber="299">pregnant</emphasis>
female collected in late August.
</paragraph>
<paragraph blockId="20.[144,1775,3292,4664]" box="[144,1266,4571,4610]" pageId="20" pageNumber="299">
<emphasis bold="true" box="[144,466,4571,4610]" pageId="20" pageNumber="299">Activity patterns.</emphasis>
Hills Horseshoe Bats likely roost in caves.
</paragraph>
<paragraph blockId="20.[144,1775,3292,4664]" box="[146,1415,4625,4664]" pageId="20" pageNumber="299">
<emphasis bold="true" box="[146,1092,4625,4664]" pageId="20" pageNumber="299">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="299" type="conservation">
<paragraph blockId="20.[1868,3500,363,667]" pageId="20" pageNumber="299">
<emphasis bold="true" box="[1868,2354,364,403]" pageId="20" pageNumber="299">Status and Conservation.</emphasis>
Classified as Critically Endangered on
<emphasis box="[3141,3496,364,403]" italics="true" pageId="20" pageNumber="299">The IUCN ed List.</emphasis>
Hills Horseshoe Bat is known from only two specimens collected in a region seriously affected by civil war. It is probably threatened by general habitat loss, roost destruction, and possibly use as bushmeat. It is found in Nyungwe Forest National Park.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="299" type="reference_group">
<paragraph blockId="20.[1868,3500,363,667]" pageId="20" pageNumber="299">
<emphasis bold="true" box="[1869,2076,582,609]" pageId="20" pageNumber="299">Bibliography.</emphasis>
ACR (2018), Csorba et al. (2003), Fahr (2010, 2013e), ahr et al. (2002), Kerbis Peterhans et al. (2013), Smith &amp; Hood (1980).
</paragraph>
</subSubSection>
</treatment>
</document>