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<document ID-DOI="http://doi.org/10.5281/zenodo.3748525" ID-GBIF-Dataset="6384b520-7e9f-4874-a414-76c2e9b01d74" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="3748525" approvalRequired="1" checkinTime="1586358796837" checkinUser="plazi" docAuthor="Burgin, Connor" docDate="2019" docId="885887A2FFD08A36FF00FCFEF899D778" docLanguage="en" docName="hbmw_9_Rhinolophidae.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docStyle="DocumentStyle{}" docTitle="Rhinolophus malayanus Bonhote in Annandale 1903" docType="treatment" docVersion="22" lastPageNumber="310" masterDocId="7461FFDAFFCF8A29FFCCFFA1FF85D963" masterDocTitle="Rhinolophidae" masterLastPageNumber="332" masterPageNumber="280" pageNumber="310" updateTime="1658949228820" updateUser="felipe">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rhinolophidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burgin, Connor</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Wilson, Don E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Mittermeier, Russell A.</mods:namePart>
</mods:name>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>280</mods:start>
<mods:end>332</mods:end>
</mods:extent>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3748525</mods:identifier>
<mods:identifier type="GBIF-Dataset">6384b520-7e9f-4874-a414-76c2e9b01d74</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3748525</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3808888" ID-GBIF-Taxon="163549955" ID-Zenodo-Dep="3808888" LSID="urn:lsid:plazi:treatment:885887A2FFD08A36FF00FCFEF899D778" httpUri="http://treatment.plazi.org/id/885887A2FFD08A36FF00FCFEF899D778" lastPageNumber="310" pageId="31" pageNumber="310">
<subSubSection box="[204,973,863,924]" pageId="31" pageNumber="310" type="vernacular_names">
<paragraph blockId="31.[204,1655,863,924]" box="[204,973,863,924]" pageId="31" pageNumber="310">
<figureCitation box="[204,273,863,924]" captionStart="On" captionStartId="26.[175,218,4332,4359]" captionTargetBox="[136,3490,134,4724]" captionTargetPageId="25" captionText="On following pages: 52. Eastern Horseshoe Bat (Rhinolophus megaphyllus}; 53. Insular Horseshoe Bat (Rhinolophus keyensis}; 54. Yellow-faced Horseshoe Bat (Rhinolophus virgo}; 55. Madura Horseshoe Bat (Rhinolophus madurensis}; 56. Sulawesi Horseshoe Bat (Rhinolophus celebensis}; 57. Robinson's Horseshoe Bat (Rhinolophus robinsoni); 58. Indochinese Horseshoe Bat (Rhinolophus chasem); 59. Anamban Horseshoe Bat (Rhinolophus nereis}; 60. Bornean Horseshoe Bat (Rhinolophus borneensis); 61. Malayan Horseshoe Bat (Rhinolophus malayanus}; 62. Acuminate Horseshoe Bat (Rhinolophus acuminatus}; 63. Acuminate Horseshoe Bat (Rhinolophus acuminatus}; 64. Little Japanese Horseshoe Bat (Rhinolophus cornutus}; 65. Yaeyama Horseshoe Bat (Rhinolophus perditus}; 66. Mountain Horseshoe Bat (Rhinolophus monticolus}; 67. Shortridge's Horseshoe Bat (Rhinolophus shortridgei}; 68. Formosan Horseshoe Bat (Rhinolophus monoceros}; 69. Least Horseshoe Bat (Rhinolophus pusillus}; 70. Little Nepalese Horseshoe Bat (Rhinolophus subbadius}; 71. Convex Horseshoe Bat (Rhinolophus convexus}; 72. Andaman Horseshoe Bat (Rhinolophus cognatus}; 73. Glossy Horseshoe Bat (Rhinolophus refulgens}; 74. Blyth's Horseshoe Bat (Rhinolophus lepidus};" figureDoi="http://doi.org/10.5281/zenodo.3758605" httpUri="https://zenodo.org/record/3758605/files/figure.png" pageId="31" pageNumber="310">61</figureCitation>
. Malayan Horseshoe
<emphasis bold="true" box="[877,973,863,924]" pageId="31" pageNumber="310">Bat</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[1037,1655,863,924]" pageId="31" pageNumber="310" type="nomenclature">
<paragraph blockId="31.[204,1655,863,924]" box="[1037,1655,863,924]" pageId="31" pageNumber="310">
<taxonomicName authorityName="Bonhote in Annandale" authorityYear="1903" box="[1037,1655,863,924]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="malayanus">
<emphasis box="[1037,1655,863,924]" italics="true" pageId="31" pageNumber="310">Rhinolophus malayanus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[202,1527,938,965]" pageId="31" pageNumber="310" type="vernacular_names">
<paragraph blockId="31.[202,1527,938,965]" box="[202,1527,938,965]" pageId="31" pageNumber="310">
<emphasis bold="true" box="[202,306,938,965]" pageId="31" pageNumber="310">French:</emphasis>
Rhinolophe de
<collectingCountry box="[520,630,938,965]" name="Malaysia" pageId="31" pageNumber="310">Malaisie</collectingCountry>
/
<emphasis bold="true" box="[656,781,938,965]" pageId="31" pageNumber="310">German:</emphasis>
Malaya-Hufeisennase
<emphasis bold="true" box="[1087,1230,938,965]" pageId="31" pageNumber="310">/ Spanish:</emphasis>
Herradura de
<collectingCountry box="[1427,1527,938,965]" name="Malaysia" pageId="31" pageNumber="310">Malasia</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="310" type="reference_group">
<paragraph blockId="31.[1023,1822,1036,1605]" pageId="31" pageNumber="310">
Taxonomy.
<taxonomicName authority="Bonhote" authorityName="Bonhote " authorityYear="1903" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="malayanus">
<emphasis box="[1250,1670,1036,1075]" italics="true" pageId="31" pageNumber="310">Rhinolophus malayanus</emphasis>
Bonhote
</taxonomicName>
in Annandale &amp; H.. Robinson, 1903,
</paragraph>
</subSubSection>
<subSubSection box="[1026,1502,1142,1181]" pageId="31" pageNumber="310" type="materials_examined">
<paragraph blockId="31.[1023,1822,1036,1605]" box="[1026,1502,1142,1181]" pageId="31" pageNumber="310">
<materialsCitation ID-GBIF-Occurrence="2597533923" box="[1042,1496,1142,1181]" country="Thailand" location="Biserat, Jalor," pageId="31" pageNumber="310" specimenCount="1" typeStatus="type">
<location LSID="urn:lsid:plazi:treatment:885887A2FFD08A36FF00FCFEF899D778:052E606FFFD08A36FBDEFBD7FA89DDFE" box="[1042,1292,1142,1181]" country="Thailand" name="Biserat, Jalor," pageId="31" pageNumber="310">Biserat,Jalor,</location>
<collectingCountry box="[1319,1496,1142,1181]" name="Thailand" pageId="31" pageNumber="310">Thailand</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="310" type="discussion">
<paragraph blockId="31.[1023,1822,1036,1605]" lastBlockId="31.[203,1825,1619,3611]" pageId="31" pageNumber="310">
Tentatively included in the
<taxonomicName authorityName="J. E. Gray" authorityYear="1834" box="[1614,1822,1195,1234]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="megaphyllus">
<emphasis box="[1614,1822,1195,1234]" italics="true" pageId="31" pageNumber="310">megaphyllus</emphasis>
</taxonomicName>
group but its placement here is uncertain; further genetic work is needed.
<taxonomicName authorityName="Bonhote in Annandale" authorityYear="1903" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="malayanus">
<emphasis italics="true" pageId="31" pageNumber="310">Rhinolophus malayanus</emphasis>
</taxonomicName>
is generally considered to be close to.
<emphasis box="[1373,1488,1407,1446]" italics="true" pageId="31" pageNumber="310">
<taxonomicName authorityName="K. Andersen" authorityYear="1905" box="[1373,1482,1407,1446]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="stheno">stheno</taxonomicName>
,
</emphasis>
but they do not cluster together in genetic studies. Exact placement of.
<emphasis box="[1357,1529,1513,1552]" italics="true" pageId="31" pageNumber="310">malayanu</emphasis>
s in relation to other
<taxonomicName box="[1142,1361,1566,1605]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="genus">
<emphasis box="[1142,1361,1566,1605]" italics="true" pageId="31" pageNumber="310">Rhinolophus</emphasis>
</taxonomicName>
is still uncertain but the most
<emphasis box="[307,429,1619,1658]" italics="true" pageId="31" pageNumber="310">recent</emphasis>
genetic study placed it as sister to
<emphasis box="[1082,1347,1619,1658]" italics="true" pageId="31" pageNumber="310">
<taxonomicName authorityName="Peters" authorityYear="1871" box="[1082,1341,1619,1658]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="acuminatus">R. acuminatus</taxonomicName>
,
</emphasis>
forming a clade sister to much of the
<emphasis box="[448,1076,1672,1711]" italics="true" pageId="31" pageNumber="310">
<taxonomicName authorityName="J. E. Gray" authorityYear="1834" box="[448,657,1672,1711]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="megaphyllus">megaphyllus</taxonomicName>
,
<taxonomicName authorityName="Temminck" authorityYear="1834" box="[673,811,1672,1711]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="pusillus">pusillus</taxonomicName>
,
<taxonomicName authorityName="Waterhouse" authorityYear="1843" box="[828,1070,1672,1711]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="philippinensis">philippinensis</taxonomicName>
,
</emphasis>
and
<taxonomicName authorityName="Blyth" authorityYear="1844" box="[1177,1317,1672,1711]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="310" phylum="Chordata" rank="species" species="macrotis">
<emphasis box="[1177,1317,1672,1711]" italics="true" pageId="31" pageNumber="310">macrotis</emphasis>
</taxonomicName>
groups. Monotypic.
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="310" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.3750028" ID-Zenodo-Dep="3750028" httpUri="https://zenodo.org/record/3750028/files/figure.png" inLine="true" pageId="31" pageNumber="310" targetBox="[201,994,1042,1600]" targetPageId="31">
<paragraph blockId="31.[203,1825,1619,3611]" pageId="31" pageNumber="310">
<emphasis bold="true" box="[203,441,1725,1764]" pageId="31" pageNumber="310">Distribution.</emphasis>
<collectingCountry box="[453,632,1725,1764]" name="Myanmar" pageId="31" pageNumber="310">Myanmar</collectingCountry>
,
<collectingCountry box="[650,826,1725,1764]" name="Thailand" pageId="31" pageNumber="310">Thailand</collectingCountry>
,
<collectingCountry box="[846,937,1725,1764]" name="Laos" pageId="31" pageNumber="310">Laos</collectingCountry>
,
<collectingCountry box="[952,1119,1725,1764]" name="Vietnam" pageId="31" pageNumber="310">Vietnam</collectingCountry>
,
<collectingCountry box="[1139,1337,1725,1764]" name="Cambodia" pageId="31" pageNumber="310">Cambodia</collectingCountry>
, and N Peninsular
<collectingCountry name="Malaysia" pageId="31" pageNumber="310">Malaysia</collectingCountry>
along with Tarutao and Langkawi Is.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="31" pageNumber="310" type="description">
<paragraph blockId="31.[203,1825,1619,3611]" pageId="31" pageNumber="310">
Descriptive notes. Head-body 36-1-55-
<quantity box="[962,1081,1828,1867]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="310" unit="mm" value="8.0">8 mm</quantity>
, tail 15-26-
<quantity box="[1307,1427,1828,1867]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="31" pageNumber="310" unit="mm" value="9.0">9 mm</quantity>
, ear 12-7-19-
<quantity box="[1694,1810,1828,1867]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="310" unit="mm" value="3.0">3 mm</quantity>
, hindfoot 5-8—9-
<quantity box="[499,611,1881,1920]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="310" unit="mm" value="6.0">6 mm</quantity>
, forearm 38-
<quantity box="[856,1038,1881,1920]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="4.4" metricValueMin="0.2" pageId="31" pageNumber="310" unit="mm" value="23.0" valueMax="44.0" valueMin="2.0">2—44 mm</quantity>
; weight 4-5—9-
<quantity box="[1313,1374,1881,1920]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="31" pageNumber="310" unit="g" value="7.0">7 g</quantity>
. Dorsal pelage is either brown (brown morph) or bright cinnamon (orange morph), whereas ventral pelage is either buff or whitish. Ears are of medium size. Noseleaf has triangular lancet with concave or nearly parallel lateral margins and elongated tip; connecting process is rounded; sella is
<emphasis box="[542,712,2096,2135]" italics="true" pageId="31" pageNumber="310">relatively</emphasis>
wide with broadly rounded top, and lateral margins are almost parallel; horseshoe is
<emphasis box="[741,911,2149,2188]" italics="true" pageId="31" pageNumber="310">relatively</emphasis>
broad (6-
<quantity box="[1099,1250,2149,2188]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="31" pageNumber="310" unit="mm" value="7.5" valueMax="9.0" valueMin="6.0">6-9 mm</quantity>
) and nearly completely covers muzzle, has moderately developed lateral leaflets, and a shallow
<emphasis box="[1441,1508,2201,2240]" italics="true" pageId="31" pageNumber="310">but</emphasis>
present median emargination. Lower lip has three mental grooves. Skull is moderately robust and zygomatic width is only slightly larger than mastoid width; anterior median swellings are well developed and wider than they are long; posterior compartments are well inflated; sagittal crest is rather weak; frontal depression is
<emphasis box="[1304,1473,2413,2452]" italics="true" pageId="31" pageNumber="310">relatively</emphasis>
elongate but shallow; supraorbital crests are more or less distinct P2 is of medium size, with distinct cusp, and is within tooth row, separating C1 and P4;
<emphasis box="[1203,1228,2519,2558]" italics="true" pageId="31" pageNumber="310">P</emphasis>
3 is small and slightly to almost completely extrudes labially from tooth row, which allows P2 and P4 to touch. Chromosomal
<emphasis box="[327,570,2627,2666]" italics="true" pageId="31" pageNumber="310">complement</emphasis>
has 2n = 62 and FNa = 60.
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="310" type="biology_ecology">
<paragraph blockId="31.[203,1825,1619,3611]" pageId="31" pageNumber="310">
<emphasis bold="true" box="[203,356,2679,2718]" pageId="31" pageNumber="310">Habitat</emphasis>
Mainly recorded from areas with limestone karst and sandstone mountains for roosting. Malayan Horseshoe Bats have been recorded in lowland mixed deciduous forest, hill evergreen forest, highly disturbed dry dipterocarp forest, and agricultural areas, indicating that the species is
<emphasis box="[953,1106,2838,2877]" italics="true" pageId="31" pageNumber="310">resilient</emphasis>
in response to disturbance. It is found at elevations from sea level up to c.
<quantity box="[857,1004,2891,2930]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="31" pageNumber="310" unit="m" value="1400.0">1400 m</quantity>
.
</paragraph>
<paragraph blockId="31.[203,1825,1619,3611]" box="[205,876,2944,2983]" pageId="31" pageNumber="310">
<emphasis bold="true" box="[205,387,2944,2983]" pageId="31" pageNumber="310">Food and</emphasis>
Feeding. No information.
</paragraph>
<paragraph blockId="31.[203,1825,1619,3611]" box="[205,1585,2997,3036]" pageId="31" pageNumber="310">
Breeding.
<emphasis box="[401,577,2997,3036]" italics="true" pageId="31" pageNumber="310">Pregnant</emphasis>
females were captured in late February in
<collectingCountry box="[1383,1580,2997,3036]" name="Cambodia" pageId="31" pageNumber="310">Cambodia</collectingCountry>
.
</paragraph>
<paragraph blockId="31.[203,1825,1619,3611]" pageId="31" pageNumber="310">
<emphasis bold="true" box="[203,519,3049,3088]" pageId="31" pageNumber="310">Activity patterns.</emphasis>
The Malayan Horseshoe Bat is nocturnal, roosting in limestone caves during the day. Torpid individuals have been captured in day roosts during July, in western
<collectingCountry box="[363,542,3152,3191]" name="Thailand" pageId="31" pageNumber="310">Thailand</collectingCountry>
. Call shape is FM/CF/FM with peak F recorded at 72—80 kHz in northern
<collectingCountry box="[391,567,3205,3244]" name="Thailand" pageId="31" pageNumber="310">Thailand</collectingCountry>
,
<quantity box="[589,753,3205,3244]" metricMagnitude="0" metricUnit="m" metricValue="2.1336" metricValueMax="2.1844" metricValueMin="2.0827999999999998" pageId="31" pageNumber="310" unit="in" value="84.0" valueMax="86.0" valueMin="82.0">82—86 in</quantity>
central
<collectingCountry box="[910,1086,3205,3244]" name="Thailand" pageId="31" pageNumber="310">Thailand</collectingCountry>
, 83—91 kHz in southern
<collectingCountry box="[1557,1733,3205,3244]" name="Thailand" pageId="31" pageNumber="310">Thailand</collectingCountry>
, 75— 82 kHz in
<collectingCountry box="[400,599,3257,3296]" name="Cambodia" pageId="31" pageNumber="310">Cambodia</collectingCountry>
, 76-79 kHz in
<collectingCountry box="[879,1059,3257,3296]" name="Myanmar" pageId="31" pageNumber="310">Myanmar</collectingCountry>
, and 77—80 kHz in
<collectingCountry box="[1422,1513,3257,3296]" name="Laos" pageId="31" pageNumber="310">Laos</collectingCountry>
.
</paragraph>
<paragraph blockId="31.[203,1825,1619,3611]" pageId="31" pageNumber="310">
<emphasis bold="true" box="[205,909,3313,3352]" pageId="31" pageNumber="310">Movements, Home range and Social</emphasis>
organization. Malayan Horseshoe Bats roost in colonies, with up to 3000 individuals recorded in one cave in western
<collectingCountry box="[1520,1697,3366,3405]" name="Thailand" pageId="31" pageNumber="310">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="310" type="conservation">
<paragraph blockId="31.[203,1825,1619,3611]" pageId="31" pageNumber="310">
<emphasis bold="true" box="[206,680,3419,3458]" pageId="31" pageNumber="310">Status and Conservation.</emphasis>
Classified as Least Concern on
<emphasis box="[1291,1629,3419,3458]" italics="true" pageId="31" pageNumber="310">The IUCN Red List.</emphasis>
There do not seem to be any major threats currently affecting this widespread species.
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="310" type="reference_group">
<paragraph blockId="31.[203,1825,1619,3611]" pageId="31" pageNumber="310">
<emphasis bold="true" box="[207,413,3531,3558]" pageId="31" pageNumber="310">Bibliography.</emphasis>
Bates, Bumrungsri &amp; Csorba (2008b), Csorba eta/. (2003), Francis (2008a), Harada, Minezawa
<emphasis italics="true" pageId="31" pageNumber="310">et al.</emphasis>
(1982),
<emphasis box="[359,476,3584,3611]" italics="true" pageId="31" pageNumber="310">Kruskop</emphasis>
(2013a),
<emphasis box="[608,725,3584,3611]" italics="true" pageId="31" pageNumber="310">Soisook</emphasis>
(2008),
<emphasis box="[845,962,3584,3611]" italics="true" pageId="31" pageNumber="310">Soisook</emphasis>
et
<emphasis box="[1008,1040,3584,3611]" italics="true" pageId="31" pageNumber="310">al.</emphasis>
(2008), Stoffberg
<emphasis box="[1308,1376,3584,3611]" italics="true" pageId="31" pageNumber="310">et al.</emphasis>
(2010), Zhang Lin et
<emphasis box="[1683,1714,3584,3611]" italics="true" pageId="31" pageNumber="310">al.</emphasis>
(2018).
</paragraph>
</subSubSection>
</treatment>
</document>