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<document id="5D57390E13DF684DF84E555B5256597E" ID-DOI="10.5281/zenodo.179514" ID-GBIF-Dataset="fcf9e9dc-fdc8-4644-8d7f-e7ce840a743a" ID-ISSN="1175-5326" ID-Zenodo-Dep="179514" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459978018765" checkinUser="plazi" docAuthor="Marin, Ivan" docDate="2007" docId="B37F3E3DFF98FF84FF6BEF56B8A33ED5" docLanguage="en" docName="zt01635p021.pdf" docOrigin="Zootaxa 1635" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pontonides tatianae Marin, 2007, n. sp." docType="treatment" docVersion="11" lastPageNumber="15" masterDocId="4F464645FF94FF8AFFFCED7BBB193D5F" masterDocTitle="The coral-associated shrimp genus Pontonides (Caridea, Palaemonidae, Pontoniinae) in Nhatrang Bay, Vietnam, with description of two new species" masterLastPageNumber="21" masterPageNumber="1" pageNumber="13" updateTime="1698231631565" updateUser="plazi">
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<mods:title id="DDD8408DE05DD6E36D133CD20ED81118">The coral-associated shrimp genus Pontonides (Caridea, Palaemonidae, Pontoniinae) in Nhatrang Bay, Vietnam, with description of two new species</mods:title>
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<mods:date id="E0B40EA25991360C957B57CE693BE8A2">2007</mods:date>
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<subSubSection id="73CCDCA0FF98FF86FF6BEF56BBF63F35" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="3B698F2BFF98FF86FF6BEF56BAC93F18" blockId="12.[151,464,557,618]" box="[151,464,557,583]" pageId="12" pageNumber="13">
<heading id="60213847FF98FF86FF6BEF56BAC93F18" bold="true" box="[151,464,557,583]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="09A25339FF98FF86FF6BEF56BAC93F18" bold="true" box="[151,464,557,583]" pageId="12" pageNumber="13">
<taxonomicName id="FCD6F4A8FF98FF86FF6BEF56BA9E3F18" ID-CoL="506490d0-2162-4107-9204-76291924f3fd" box="[151,391,557,583]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tatianae" status="sp. nov.">
<emphasis id="09A25339FF98FF86FF6BEF56BA9E3F18" bold="true" box="[151,391,557,583]" italics="true" pageId="12" pageNumber="13">Pontonides tatianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1291EE42FF98FF86FE72EF56BAC93F18" box="[398,464,557,583]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="3B698F2BFF98FF86FF6BEF2BBBF63F35" blockId="12.[151,464,557,618]" box="[151,239,592,618]" pageId="12" pageNumber="13">
(
<figureCitation id="A3ED93AEFF98FF86FF63EF2BBBFC3F35" box="[159,229,592,618]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="73CCDCA0FF98FF86FF6BEFF6B8673F95" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="3B698F2BFF98FF86FF6BEFF6B8673F95" blockId="12.[151,1437,653,2031]" pageId="12" pageNumber="13">
<emphasis id="09A25339FF98FF86FF6BEFF6BA533FF8" bold="true" box="[151,330,653,679]" pageId="12" pageNumber="13">
<typeStatus id="E46D3189FF98FF86FF6BEFF6BBCB3FF8" box="[151,210,653,679]" pageId="12" pageNumber="13">Type</typeStatus>
material.
</emphasis>
<emphasis id="09A25339FF98FF86FEAFEFF6BADF3FF9" box="[339,454,653,678]" italics="true" pageId="12" pageNumber="13">
<typeStatus id="E46D3189FF98FF86FEAFEFF6BADB3FF9" box="[339,450,653,678]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="43C1CFBBFF98FF86FE31EFF6B92A3FF8" box="[461,563,653,679]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
, Nhatrang Bay: female (pcl.
<quantity id="FC2E22CEFF98FF86FC7FEFF6B8F93FF8" box="[899,992,653,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="mm" value="1.8">1.8 mm</quantity>
), Dun I., on black coral
<taxonomicName id="FCD6F4A8FF98FF86FB05EFF6BE853FF9" box="[1273,1436,653,678]" class="Anthozoa" family="Aphanipathidae" genus="Rhipidipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="12" pageNumber="13" phylum="Cnidaria" rank="genus">
<emphasis id="09A25339FF98FF86FB05EFF6BE853FF9" box="[1273,1436,653,678]" italics="true" pageId="12" pageNumber="13">Rhipidipathes</emphasis>
</taxonomicName>
sp., depth
<quantity id="FC2E22CEFF98FF86FEF3EFCBBA573F95" box="[271,334,688,714]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="13" unit="m" value="40.0">40 m</quantity>
, coll. I. Marin,
<date id="4F68A9EBFF98FF86FDFEEFCBB9963F95" box="[514,655,688,714]" pageId="12" pageNumber="13" value="2006-06-02">2 June 2006</date>
(
<collectionCode id="5DC717EEFF98FF86FD63EFCBB9E53F95" box="[671,764,688,714]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="12" pageNumber="13">ZMMU</collectionCode>
Ma 5489).
</paragraph>
</subSubSection>
<subSubSection id="73CCDCA0FF98FF86FF3BEFA9BA213E80" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="3B698F2BFF98FF86FF3BEFA9BA213E80" blockId="12.[151,1437,653,2031]" pageId="12" pageNumber="13">
<emphasis id="09A25339FF98FF86FF3BEFA9BA5E3FB3" bold="true" box="[199,327,722,748]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
Small-sized pontoniine shrimps with subcylindrical body. Carapace smooth, not gibbous. Rostrum long, overreaching distal margin of basal antennular segment. Supraorbital margin well developed, rectangular, with small tooth; orbits well developed. Eyestalk without dorsal tubercle. Fourth and fifth pleura rounded. Second maxilliped without fusiform setae. Third maxilliped with numerous fusiform setae on proximal half of basal segment. First to fifth pereiopods without fusiform setae on ischia. Second pereiopods unequal; major chela covered with tiny tubercles, setose, fingers stout and curved; minor chela sparsely setose; fingers 22.5 times as long as wide, with curved tips. Third pereiopods setose; dactylus simple, curved, acute distally.
</paragraph>
</subSubSection>
<subSubSection id="73CCDCA0FF98FF84FF3BEE9CBE903C63" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="3B698F2BFF98FF86FF3BEE9CBA7339D3" blockId="12.[151,1437,653,2031]" pageId="12" pageNumber="13">
<emphasis id="09A25339FF98FF86FF3BEE9CBA46395E" bold="true" box="[199,351,999,1025]" pageId="12" pageNumber="13">Description.</emphasis>
Female
<typeStatus id="E46D3189FF98FF86FE3AEE93B929395D" box="[454,560,1000,1026]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
, tl.
<quantity id="FC2E22CEFF98FF86FDA0EE93B9A3395D" box="[604,698,1000,1026]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="12" pageNumber="13" unit="mm" value="3.1">3.1 mm</quantity>
, pcl.
<quantity id="FC2E22CEFF98FF86FD06EE93B843395D" box="[762,858,1000,1026]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="mm" value="1.8">1.8 mm</quantity>
. Carapace smooth, not gibbous, with well developed antennal tooth; hepatic tooth absent; anterolateral angle broadly concave. Rostrum slender, elongate, overreaching distal margin of basal antennular segment, oval in cross-section (
<figureCitation id="A3ED93AEFF98FF86FBD6E956BF763918" box="[1066,1135,1069,1095]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
B, C). Supraorbital carina well developed, rectangular, with small teeth, broadly expanding over proximal portion of eyes, reaching midlength of rostrum.
</paragraph>
<paragraph id="3B698F2BFF98FF86FF3BE9EEBE673865" blockId="12.[151,1437,653,2031]" pageId="12" pageNumber="13">
Abdominal somites smooth, tergites non-carinate or posteriorly produced; pleura of first to fifth abdominal somites posteroventrally rounded (
<figureCitation id="A3ED93AEFF98FF86FDA5E9C3B987398D" box="[601,670,1208,1234]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
A). Telson slender, about four times as long as wide, not tapering distally, with two pairs of small slender submarginal dorsal spines situated at about 0.5 and 0.8 of telson length, respectively; posterior margin with three pairs of spines: short lateral spines, longer slender intermediate spines, and slender plumose submedian spines, latter about half length of intermediate spines (
<figureCitation id="A3ED93AEFF98FF86FAEEE85BBE4E3865" box="[1298,1367,1312,1338]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
D).
</paragraph>
<paragraph id="3B698F2BFF98FF86FF3BE839B8733803" blockId="12.[151,1437,653,2031]" box="[199,874,1346,1372]" pageId="12" pageNumber="13">Eyes large, well-developed; cornea and eyestalk rounded.</paragraph>
<paragraph id="3B698F2BFF98FF86FF3BE81EB9B238B8" blockId="12.[151,1437,653,2031]" pageId="12" pageNumber="13">
Antennules (
<figureCitation id="A3ED93AEFF98FF86FEA0E81EBA863820" box="[348,415,1381,1407]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
E) with basal segment about twice as long as wide; lateral margin with strong acute distolateral tooth and distinct. Antenna with distolateral margin of basicerite rounded, without distolateral tooth; scaphocerite broad, not exceeding antennular peduncle and rostrum, about 1.5 times as long as maximum width, with well developed distolateral tooth.
</paragraph>
<paragraph id="3B698F2BFF98FF86FF3BE88BBFAB3BA3" blockId="12.[151,1437,653,2031]" pageId="12" pageNumber="13">
Mouthparts without specific features, typical for genus. Mandible (
<figureCitation id="A3ED93AEFF98FF86FC23E88BBF3D3B55" box="[991,1060,1520,1546]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
F) without palp; incisor process slender, tapering distally, with four terminal teeth; molar process well developed. Maxillula normal; palp well developed, bilobed; upper lacinia larger than lower, slightly tapering distally, with strong distal spines; lower lacinia slender, distally rounded, with distal setae. Maxilla with well developed simple rounded palp. First maxilliped (
<figureCitation id="A3ED93AEFF98FF86FEDAEB01BA753BCB" box="[294,364,1658,1684]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
G) with completely fused endites; palp with distal setae; exopod greatly reduced. Second maxilliped with ischium furnished with several simple setae in central portion of mesial margin; epipod oval, without podobranch. Third maxilliped (
<figureCitation id="A3ED93AEFF98FF86FD9AEBBBB9B33B85" box="[614,682,1728,1754]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
H) with broad segments; basal segment with numerous fusiform setae along proximal half of lateral margin; exopod well developed; arthrobranch present.
</paragraph>
<paragraph id="3B698F2BFF98FF86FF3BEA7EBFDE3AB0" blockId="12.[151,1437,653,2031]" pageId="12" pageNumber="13">
First pereiopods slender, unarmed; palm twice as long as wide, subcylindrical; fingers slender, about three times as long as wide, with straight cutting edges and row of stiff setae near tips. Second pereiopods unequal; major chela (
<figureCitation id="A3ED93AEFF98FF86FECEEA31BA613A3B" box="[306,376,1866,1892]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">
Fig.
<date id="4F68A9EBFF98FF86FE96EA31BA613A3B" box="[362,376,1866,1892]" pageId="12" pageNumber="13">8</date>
</figureCitation>
I) with papillose and setose palm; fingers stout, about twice as long as wide, curved, with well developed dentate cutting edges (
<figureCitation id="A3ED93AEFF98FF86FD9FEA16B9B23AD8" box="[611,683,1901,1927]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
L); minor chela (
<figureCitation id="A3ED93AEFF98FF86FC85EA16B8D93AD8" box="[889,960,1901,1927]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
J) with smooth but setose palm; fingers slender about four times as long as wide, with straight cutting edges. Third to fifth pereiopods (
<figureCitation id="A3ED93AEFF98FF86FB0CEAEBBE2D3AF5" box="[1264,1332,1936,1962]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
K) stout, smooth; dactylus simple, curved, distally acute (
<figureCitation id="A3ED93AEFF98FF86FD37EAC9B8163A93" box="[715,783,1970,1996]" captionStart="FIGURE 8" captionStartId="13.[151,255,1905,1929]" captionTargetBox="[158,1436,192,1830]" captionTargetId="figure@13.[151,1436,183,1846]" captionTargetPageId="13" captionText="FIGURE 8. Pontonides tatianae n. sp. Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (B D, I K); 0.7 mm (E, F); 0.5 mm (G, H, L, M)." httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
M). Pleopods not especially modified. Uropods slender, exceeding telson; lateral margin of endopod with small movable spine, without distal tooth.
</paragraph>
<caption id="6FA9DFA3FF99FF87FF6BEA0ABFA53AAF" httpUri="https://zenodo.org/record/179522/files/figure.png" pageId="13" pageNumber="14" targetBox="[158,1436,192,1830]" targetPageId="13">
<paragraph id="3B698F2BFF99FF87FF6BEA0ABFA53AAF" blockId="13.[151,1436,1905,2032]" pageId="13" pageNumber="14">
<emphasis id="09A25339FF99FF87FF6BEA0ABA023AD6" bold="true" box="[151,283,1905,1929]" pageId="13" pageNumber="14">FIGURE 8.</emphasis>
<taxonomicName id="FCD6F4A8FF99FF87FED9EA09BAE73AD6" box="[293,510,1906,1929]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tatianae" status="sp. nov.">
<emphasis id="09A25339FF99FF87FED9EA09BAE73AD6" box="[293,510,1906,1929]" italics="true" pageId="13" pageNumber="14">Pontonides tatianae</emphasis>
</taxonomicName>
<emphasis id="09A25339FF99FF87FDFBEA0AB95F3AD6" bold="true" box="[519,582,1905,1929]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="1291EE42FF99FF87FDFBEA0AB95F3AD6" box="[519,582,1905,1929]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Female holotype (ZMMU Ma 5489). A, habitus. B, front of carapace, dorsal view. C, front of carapace, lateral view; D, telson, E, antennule; F, mandible; G, second maxilliped. H, third maxilliped. I, major second pereiopod. J, minor second pereiopod. K, fingers of major second pereiopod. L, third pereiopod. M, third pereiopod, dactylus. Scale bars: 2 mm (A); 1 mm (BD, IK); 0.7 mm (E, F); 0.5 mm (G, H, L, M).
</paragraph>
</caption>
<paragraph id="3B698F2BFF9AFF84FF3BEDE3BA663DED" blockId="14.[151,1436,152,906]" box="[199,383,152,178]" pageId="14" pageNumber="15">Male unknown.</paragraph>
<paragraph id="3B698F2BFF9AFF84FF3BEDC1BA773C45" blockId="14.[151,1436,152,906]" pageId="14" pageNumber="15">
<emphasis id="09A25339FF9AFF84FF3BEDC1BA7C3D8B" bold="true" box="[199,357,186,212]" pageId="14" pageNumber="15">Colouration.</emphasis>
Body and appendages generally translucent, covered with tiny black chromatophores; longitudinal dark-blue band extending lateroventrally from anterior carapace to sixth abdominal segment (I. Marin, pers. obs.).
</paragraph>
<paragraph id="3B698F2BFF9AFF84FF3BEC59BE903C63" blockId="14.[151,1436,152,906]" box="[199,1417,290,316]" pageId="14" pageNumber="15">
<emphasis id="09A25339FF9AFF84FF3BEC59BA193C63" bold="true" box="[199,256,290,316]" pageId="14" pageNumber="15">Size.</emphasis>
Non-ovigerous female
<typeStatus id="E46D3189FF9AFF84FDEAEC59B9653C63" box="[534,636,290,316]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(only specimen known), with cl.
<quantity id="FC2E22CEFF9AFF84FBFAEC59BF7A3C63" box="[1030,1123,290,316]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="15" unit="mm" value="2.3">2.3 mm</quantity>
, pcl.
<quantity id="FC2E22CEFF9AFF84FB5CEC59BFE43C63" box="[1184,1277,290,316]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="15" unit="mm" value="1.8">1.8 mm</quantity>
, tl.
<quantity id="FC2E22CEFF9AFF84FAD4EC59BE9C3C63" box="[1320,1413,290,316]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="14" pageNumber="15" unit="mm" value="3.1">3.1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="73CCDCA0FF9AFF84FF3BEC3EB9963C00" box="[199,655,325,351]" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="3B698F2BFF9AFF84FF3BEC3EB9963C00" blockId="14.[151,1436,152,906]" box="[199,655,325,351]" pageId="14" pageNumber="15">
<emphasis id="09A25339FF9AFF84FF3BEC3EBA743C00" bold="true" box="[199,365,325,351]" pageId="14" pageNumber="15">
<typeStatus id="E46D3189FF9AFF84FF3BEC3EBA1D3C00" box="[199,260,325,351]" pageId="14" pageNumber="15">Type</typeStatus>
locality.
</emphasis>
Nhatrang Bay,
<collectingCountry id="43C1CFBBFF9AFF84FDD8EC3EB9923C00" box="[548,651,325,351]" name="Vietnam" pageId="14" pageNumber="15">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="73CCDCA0FF9AFF84FF3BEC1CBA1E3F53" pageId="14" pageNumber="15" type="etymology">
<paragraph id="3B698F2BFF9AFF84FF3BEC1CBA253CFB" blockId="14.[151,1436,152,906]" pageId="14" pageNumber="15">
<emphasis id="09A25339FF9AFF84FF3BEC1CBA553CDE" bold="true" box="[199,332,359,385]" pageId="14" pageNumber="15">Etymology</emphasis>
. The species is named in honour of my mother, Tatiana Marina, and Tatiana Anker, mother of Arthur Anker.
</paragraph>
<paragraph id="3B698F2BFF9AFF84FF3BECD6BA1E3F53" blockId="14.[151,1436,152,906]" pageId="14" pageNumber="15">
<emphasis id="09A25339FF9AFF84FF3BECD6BA293C98" bold="true" box="[199,304,429,455]" pageId="14" pageNumber="15">Ecology.</emphasis>
The species was found on a colony of the bushy black coral,
<taxonomicName id="FCD6F4A8FF9AFF84FBDDECD6BFDD3C99" box="[1057,1220,429,454]" class="Anthozoa" family="Aphanipathidae" genus="Rhipidipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="genus">
<emphasis id="09A25339FF9AFF84FBDDECD6BFDD3C99" box="[1057,1220,429,454]" italics="true" pageId="14" pageNumber="15">Rhipidipathes</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="FCD6F4A8FF9AFF84FAFCECD6BE8E3C98" box="[1280,1431,429,455]" class="Anthozoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="order">Antipatharia</taxonomicName>
,
<taxonomicName id="FCD6F4A8FF9AFF84FF6BECABBA293CB5" box="[151,304,464,490]" class="Anthozoa" family="Antipathidae" kingdom="Animalia" order="Antipatharia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="family">Antipathidae</taxonomicName>
) at a depth of about 40 meters. The shrimps were found at the base of the colony, clinging to the branches.
</paragraph>
</subSubSection>
<subSubSection id="73CCDCA0FF9AFF84FF3BEF6EB8A33ED5" pageId="14" pageNumber="15" type="distribution">
<paragraph id="3B698F2BFF9AFF84FF3BEF6EBFD13F70" blockId="14.[151,1436,152,906]" box="[199,1224,533,559]" pageId="14" pageNumber="15">
<emphasis id="09A25339FF9AFF84FF3BEF6EBA7E3F70" bold="true" box="[199,359,533,559]" pageId="14" pageNumber="15">Distribution.</emphasis>
Western Pacific: presently known only from the
<typeStatus id="E46D3189FF9AFF84FC5AEF6EB8C03F70" box="[934,985,533,559]" pageId="14" pageNumber="15">type</typeStatus>
locality in
<collectingCountry id="43C1CFBBFF9AFF84FBA1EF6EBFDD3F70" box="[1117,1220,533,559]" name="Vietnam" pageId="14" pageNumber="15">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="3B698F2BFF9AFF84FF3BEF4CB8A33ED5" blockId="14.[151,1436,152,906]" pageId="14" pageNumber="15">
<emphasis id="09A25339FF9AFF84FF3BEF4CBAD93F0E" bold="true" box="[199,448,567,593]" pageId="14" pageNumber="15">Systematic position.</emphasis>
<taxonomicName id="FCD6F4A8FF9AFF84FE3BEF43B9AB3F0E" box="[455,690,568,593]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tatianae" status="sp. nov.">
<emphasis id="09A25339FF9AFF84FE3BEF43B9AB3F0E" box="[455,690,568,593]" italics="true" pageId="14" pageNumber="15">Pontonides tatianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1291EE42FF9AFF84FD46EF4CB9E53F0E" box="[698,764,567,593]" pageId="14" pageNumber="15" rank="species">
<emphasis id="09A25339FF9AFF84FD46EF4CB9E53F0E" bold="true" box="[698,764,567,593]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
shares with
<emphasis id="09A25339FF9AFF84FC73EF43BF283F0E" box="[911,1073,568,593]" italics="true" pageId="14" pageNumber="15">P. a s p e r u l a t u s</emphasis>
and
<emphasis id="09A25339FF9AFF84FB91EF43BFF93F0E" box="[1133,1248,568,593]" italics="true" pageId="14" pageNumber="15">P. u n c i g e r</emphasis>
the presence of a long rostrum (reaching to the distal margin of the proximal segment of the antennular peduncle), the supraorbital carina bearing a sharp tooth, the major chela of the second pereiopods being covered with small tubercles, and the marginal position of the dorsal spines of the telson. This unique feature combination suggests that
<taxonomicName id="FCD6F4A8FF9AFF84FEF0EFB9BA813F84" box="[268,408,706,731]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tataianae" status="sp. nov.">
<emphasis id="09A25339FF9AFF84FEF0EFB9BA813F84" box="[268,408,706,731]" italics="true" pageId="14" pageNumber="15">P. tataianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1291EE42FF9AFF84FE63EFB9BAF83F83" box="[415,481,706,732]" pageId="14" pageNumber="15" rank="species">
<emphasis id="09A25339FF9AFF84FE63EFB9BAF83F83" bold="true" box="[415,481,706,732]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="FCD6F4A8FF9AFF84FE0CEFB9B98C3F84" box="[496,661,706,731]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="asperulatus">
<emphasis id="09A25339FF9AFF84FE0CEFB9B98C3F84" box="[496,661,706,731]" italics="true" pageId="14" pageNumber="15">P. asperulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="FCD6F4A8FF9AFF84FD32EFB9B85C3F84" box="[718,837,706,731]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="09A25339FF9AFF84FD32EFB9B85C3F84" box="[718,837,706,731]" italics="true" pageId="14" pageNumber="15">P. unciger</emphasis>
</taxonomicName>
form a morphologically well-defined group within
<taxonomicName id="FCD6F4A8FF9AFF84FF6BEF9EBA023FA1" box="[151,283,741,766]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="09A25339FF9AFF84FF6BEF9EBA023FA1" box="[151,283,741,766]" italics="true" pageId="14" pageNumber="15">Pontonides</emphasis>
</taxonomicName>
(see also
<bibRefCitation id="5F47F2DAFF9AFF84FE6EEF9EB9393FA0" author="Bruce" box="[402,544,741,767]" pageId="14" pageNumber="20" refString="Bruce, A. J. (2005) Pontoniine shrimps from Papua New Guinea, with description of two new genera, Cainonia and Colemonia (Crustacea: Decapoda: Palaemonidae). Memoirs of the Queensland Museum, 51, 333 - 383." type="journal article" year="2005">Bruce 2005</bibRefCitation>
).
<taxonomicName id="FCD6F4A8FF9AFF84FDC8EF9EB8063FA1" box="[564,799,741,766]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tatianae" status="sp. nov.">
<emphasis id="09A25339FF9AFF84FDC8EF9EB8063FA1" box="[564,799,741,766]" italics="true" pageId="14" pageNumber="15">Pontonides tatianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1291EE42FF9AFF84FCD4EF9EB8723FA0" box="[808,875,741,767]" pageId="14" pageNumber="15" rank="species">
<emphasis id="09A25339FF9AFF84FCD4EF9EB8723FA0" bold="true" box="[808,875,741,767]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="FCD6F4A8FF9AFF84FBF4EF9EBFB63FA1" box="[1032,1199,741,766]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="asperulatus">
<emphasis id="09A25339FF9AFF84FBF4EF9EBFB63FA1" box="[1032,1199,741,766]" italics="true" pageId="14" pageNumber="15">P. asperulatus</emphasis>
</taxonomicName>
by the smaller size, and the rounded posteroventral margins of the abdominal pleura. Furthermore the two species have different colour patterns and are associated with different hosts. The rounded pleura of the fourth and fifth abdominal segments and the absence of fusiform setae on the second maxilliped enable to separate
<taxonomicName id="FCD6F4A8FF9AFF84FB64EE36BE0D3E39" box="[1176,1300,845,870]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tatianae" status="sp. nov.">
<emphasis id="09A25339FF9AFF84FB64EE36BE0D3E39" box="[1176,1300,845,870]" italics="true" pageId="14" pageNumber="15">P. tatianae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1291EE42FF9AFF84FAE7EE36BE443E38" box="[1307,1373,845,871]" pageId="14" pageNumber="15" rank="species">
<emphasis id="09A25339FF9AFF84FAE7EE36BE443E38" bold="true" box="[1307,1373,845,871]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
from
<taxonomicName id="FCD6F4A8FF9AFF84FF6BEE0BBA153ED6" box="[151,268,880,905]" class="Malacostraca" family="Palaemonidae" genus="Pontonides" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="unciger">
<emphasis id="09A25339FF9AFF84FF6BEE0BBA153ED6" box="[151,268,880,905]" italics="true" pageId="14" pageNumber="15">P. unciger</emphasis>
</taxonomicName>
, which also associates with
<taxonomicName id="FCD6F4A8FF9AFF84FDA4EE0BB9E23ED6" box="[600,763,880,905]" class="Anthozoa" family="Aphanipathidae" genus="Rhipidipathes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antipatharia" pageId="14" pageNumber="15" phylum="Cnidaria" rank="genus">
<emphasis id="09A25339FF9AFF84FDA4EE0BB9E23ED6" box="[600,763,880,905]" italics="true" pageId="14" pageNumber="15">Rhipidipathes</emphasis>
</taxonomicName>
sp. (see below).
</paragraph>
</subSubSection>
</treatment>
</document>