217 lines
26 KiB
XML
217 lines
26 KiB
XML
<document id="480A8466573FB8FEBAF9F861E0BEA810" ID-CLB-Dataset="99091" ID-DOI="10.11646/zootaxa.5165.4.6" ID-GBIF-Dataset="d2d13df9-c447-4103-9a4c-d952ee8f3f68" ID-ISSN="1175-5326" ID-Zenodo-Dep="6854006" ID-ZooBank="97EF241E-F5DC-4C40-8B96-580125DCF733" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658127250201" checkinUser="plazi" docAuthor="Pankowski, Maximilian G. & Fanti, Fabrizio" docDate="2022" docId="657D7273FF95FFF32BDEFA4BB864C320" docLanguage="en" docName="zootaxa.5165.4.6.pdf" docOrigin="Zootaxa 5165 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malthodes (Malthodes) markpankowskii M. G. PANKOWSKI & FANTI 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="553" masterDocId="99440A0BFF96FFF62B49FFBCBC2DC411" masterDocTitle="Two new fossil species of soldier beetles (Coleoptera, Cantharidae, Malthininae) from Baltic amber" masterLastPageNumber="556" masterPageNumber="548" pageNumber="551" updateTime="1699360237436" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="EA60319C149528631C803467AE818845" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="438039740CDB68A3ABB22DC0406E970A">
|
||
<mods:title id="925AEF4BEC0ABBB398DBD3D660A1FD0E">Two new fossil species of soldier beetles (Coleoptera, Cantharidae, Malthininae) from Baltic amber</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="F04AD5F672505B18C48E01FE0425ADDD" type="personal">
|
||
<mods:role id="6AE9792D13CB777957FD83464009A1D2">
|
||
<mods:roleTerm id="FB8AA67DFD12861A8E62E8B54CBD6FF6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3FCE14DF4692120844E9D8A3B0EBEE0B">Pankowski, Maximilian G.</mods:namePart>
|
||
<mods:affiliation id="3CA7933A912291CF70A9DB35E1413C5D">16405 Fox Valley Terrace, Rockville, Maryland 20853, USA.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="63A8C77F7E4935F12CAD6D2D1ED20A10" type="personal">
|
||
<mods:role id="88A1612D77263F77AC094F89221AEE06">
|
||
<mods:roleTerm id="73B5635CFCE73156FE7D9FAE2CB267A0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="428CD3A54E491A8D403A4115704B79DC">Fanti, Fabrizio</mods:namePart>
|
||
<mods:affiliation id="D941AC84858F4A1F7FA886BB5586DAAB">Via del Tamburino 69, I- 53040 Piazze (SI), Italy.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="3A2B6B0F72DCF9605E58D7C411AC5F79">text</mods:typeOfResource>
|
||
<mods:relatedItem id="B9B0F11A5EF5DC30347D4013266DA740" type="host">
|
||
<mods:titleInfo id="A9AEBE2092BF121E48907ACEDB26019B">
|
||
<mods:title id="4635B28EF80AA68D7F080A097B4B39BE">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="639AAA4E6887A953C0375C0D29FC3B44">
|
||
<mods:date id="9C233C9E47E4EC7434C1D40566ED7E2E">2022</mods:date>
|
||
<mods:detail id="81CCD8A0D6739EDE6342C8ECF50EC696" type="pubDate">
|
||
<mods:number id="A6EA472E41EE4F682C0E331EAFB50BDA">2022-07-18</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="26AA53D1DA195E559A22CDE3FF274A5D" type="volume">
|
||
<mods:number id="B4B4FFF8F764873582B8E7C209C79A0E">5165</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="21702CC511DEF7E3F70AF98F734C2352" type="issue">
|
||
<mods:number id="AEA9190A3F025CE59E3E617177BB7315">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="86A46A55CD346695278C4C5774103E0B" unit="page">
|
||
<mods:start id="25D2B91CFC078DA82708ED947ACFDA68">548</mods:start>
|
||
<mods:end id="8DB2C16EE0934921D9ABA1A866E996A6">556</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="A5A36AB95D737AB27F877E5DFBA69B0E">journal article</mods:classification>
|
||
<mods:identifier id="44A1D284B63F921935411FC604903326" type="CLB-Dataset">99091</mods:identifier>
|
||
<mods:identifier id="FD2E673AAE8DCB0B783136C614823335" type="DOI">10.11646/zootaxa.5165.4.6</mods:identifier>
|
||
<mods:identifier id="EAFAE95854D0F49BCAD37B2F775DAD82" type="GBIF-Dataset">d2d13df9-c447-4103-9a4c-d952ee8f3f68</mods:identifier>
|
||
<mods:identifier id="6FF0DA9B91A964574163289C260651FB" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="04C87C125477126B18C71A9410ABA26A" type="Zenodo-Dep">6854006</mods:identifier>
|
||
<mods:identifier id="FF47FD1BBE385F8DDC2DF1A2DF7C9C6B" type="ZooBank">97EF241E-F5DC-4C40-8B96-580125DCF733</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="657D7273FF95FFF32BDEFA4BB864C320" ID-DOI="http://doi.org/10.5281/zenodo.6853998" ID-GBIF-Taxon="197817483" ID-Zenodo-Dep="6853998" LSID="urn:lsid:plazi:treatment:657D7273FF95FFF32BDEFA4BB864C320" httpUri="http://treatment.plazi.org/id/657D7273FF95FFF32BDEFA4BB864C320" lastPageId="5" lastPageNumber="553" pageId="3" pageNumber="551">
|
||
<subSubSection id="A5CE90EEFF95FFF52BDEFA4BB859C200" box="[151,1140,1527,1554]" pageId="3" pageNumber="551" type="nomenclature">
|
||
<paragraph id="ED6BC365FF95FFF52BDEFA4BB859C200" blockId="3.[151,1140,1527,1589]" box="[151,1140,1527,1554]" pageId="3" pageNumber="551">
|
||
<heading id="B6237409FF95FFF52BDEFA4BB859C200" bold="true" box="[151,1140,1527,1554]" fontSize="11" level="1" pageId="3" pageNumber="551" reason="1">
|
||
<emphasis id="DFA01F77FF95FFF52BDEFA4BB859C200" bold="true" box="[151,1140,1527,1554]" pageId="3" pageNumber="551">
|
||
<taxonomicName id="2AD4B8E6FF95FFF52BDEFA4BB822C200" authority="M. G. PANKOWSKI & FANTI" authorityName="M. G. PANKOWSKI & FANTI" authorityYear="2022" box="[151,1039,1527,1554]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="551" phylum="Arthropoda" rank="species" species="markpankowskii" status="sp. nov." subGenus="Malthodes">
|
||
<emphasis id="DFA01F77FF95FFF52BDEFA4BBD3BC203" bold="true" box="[151,278,1527,1554]" italics="true" pageId="3" pageNumber="551">Malthodes</emphasis>
|
||
(
|
||
<emphasis id="DFA01F77FF95FFF52A6EFA4BBD8BC203" bold="true" box="[295,422,1527,1554]" italics="true" pageId="3" pageNumber="551">Malthodes</emphasis>
|
||
)
|
||
<emphasis id="DFA01F77FF95FFF52AF1FA4BBEADC203" bold="true" box="[440,640,1527,1554]" italics="true" pageId="3" pageNumber="551">markpankowskii</emphasis>
|
||
M. G. PANKOWSKI & FANTI
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C493A20CFF95FFF52F5FFA4BB859C200" box="[1046,1140,1527,1553]" pageId="3" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5CE90EEFF95FFF22BDEF9A7B85FC62C" lastPageId="4" lastPageNumber="552" pageId="3" pageNumber="551" type="description">
|
||
<paragraph id="ED6BC365FF95FFF52BDEF9A7BD3FC224" blockId="3.[151,1140,1527,1589]" box="[151,274,1563,1589]" pageId="3" pageNumber="551">
|
||
(
|
||
<figureCitation id="75EFDFE0FF95FFF52BD6F9A7BD27C224" box="[159,266,1563,1589]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1870,1894]" captionStartId-1="5.[151,250,1169,1193]" captionTargetBox-0="[151,1436,705,1847]" captionTargetBox-1="[151,1418,181,1144]" captionTargetId-0="figure-238@4.[151,1436,705,1847]" captionTargetId-1="figure-22@5.[151,1436,181,1144]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 3. Malthodes (Malthodes) markpankowskii sp. nov. A: Holotype, habitus, dorsal view, scale bar = 0.5 mm. B: Holotype, habitus, detail of metasternum, scale bar = 0.2 mm. C: Holotype, habitus, ventral view, scale bar = 1.0 mm. D: Holotype, habitus, ventral view (detail), scale bar = 0.5 mm." captionText-1="FIGURE 4. Malthodes (Malthodes) markpankowskii sp. nov. A: Holotype, habitus, detail of last abdominal segments (ventral view), scale bar = 0.1 mm. B: Reconstruction of last abdominal segments, scale bar = 0.1 mm. Illustration by Dr. Carly Tribull." figureDoi-0="http://doi.org/10.5281/zenodo.6854014" figureDoi-1="http://doi.org/10.5281/zenodo.6854018" httpUri-0="https://zenodo.org/record/6854014/files/figure.png" httpUri-1="https://zenodo.org/record/6854018/files/figure.png" pageId="3" pageNumber="551">Figs. 3–4</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="ED6BC365FF95FFF52BDEF9DFBF6CC2B0" blockId="3.[151,1437,1635,2022]" pageId="3" pageNumber="551">
|
||
<emphasis id="DFA01F77FF95FFF52BDEF9DFBD04C26C" bold="true" box="[151,297,1635,1661]" pageId="3" pageNumber="551">Description.</emphasis>
|
||
Adult, winged, male defined on the basis of the last urites strongly modified. Body length: 4.0 mm. Entirely blackish-dark brown without yellow spots on elytra.
|
||
</paragraph>
|
||
<paragraph id="ED6BC365FF95FFF22B8EF917B85FC62C" blockId="3.[151,1437,1635,2022]" lastBlockId="4.[151,1437,151,681]" lastPageId="4" lastPageNumber="552" pageId="3" pageNumber="551">Head exposed, wide, rounded, covered by short setae. Eyes large, very prominent, convex, rounded, inserted in the lateral-upper part of head. Mandibles falciform, elongated. Maxillary palpi 4-segmented, with last palpomere globular and distally pointed. Labial palpi 3-segmented, with last palpomere globular and distally pointed. Antennae filiform, 11-segmented, relatively short as they only reach the apex of elytra and about half of abdomen; antennomere I elongated, slightly club-shaped (enlarged from the middle to the apex); antennomere II about 2.0 times shorter than antennomere I; antennomere III about 1.1 times longer than antennomere II; antennomeres IV–IX subequal in length, longer than antennomere III; antennomere X about 1.4 times shorter than previous ones; antennomere XI elongated, rounded at apex; all antennomeres covered by several long setae. Pronotum strongly transverse (approximately one third wider than long), as wide as head, surface almost flat and barely punctate (shallow punctation) with many short setae, sides straight and strongly bordered, posterior and anterior margin strongly bordered and very slightly enlarged/curved in the middle. Elytra short (revealing four tergites uncovered), wider than pronotum, covered with shallow punctation and several erect setae, parallel-sided, strongly rounded at apexes. Hind wings infuscate, exceeding the elytra and partially covering penultimate tergite but not the last one. Legs slender, pubescent; coxae short, stout; trochanters elongated with rounded apex; femora enlarged, slightly curved; tibiae cylindrical and thin, pro- and mesotibiae shorter than pro- and mesofemora, metatibiae as long as metafemora. Tarsi 5-segmented, pubescent; tarsomere I thin, elongated; tarsomere II shorter than tarsomere I; tarsomere III shorter than second; tarsomere IV strongly bilobed; tarsomere V elongated, slightly curved, slender; claws simple without tooth. Metasternum with rounded posterior margin, covered with many dispersed, short setae. Sternites transverse and pubescent. Penultimate tergite (tg9) wide, subrectangular; last tergite (tg10) narrower than penultimate tergite, elongated, with apical margin rather deeply concave; last sternite (st9) very elongated, as narrow as last tergite, apically almost straight (very slightly emarginate). Aedeagus not visible. Female unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5CE90EEFF92FFF22B8EFDFBB8C4C670" box="[199,1257,583,609]" pageId="4" pageNumber="552" type="etymology">
|
||
<paragraph id="ED6BC365FF92FFF22B8EFDFBB8C4C670" blockId="4.[151,1437,151,681]" box="[199,1257,583,609]" pageId="4" pageNumber="552">
|
||
<emphasis id="DFA01F77FF92FFF22B8EFDFBBD63C670" bold="true" box="[199,334,583,609]" pageId="4" pageNumber="552">Etymology.</emphasis>
|
||
Species named after Mark S. Pankowski, the big-hearted father of the first author.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5CE90EEFF92FFF32B8EFDD7BFCBC1FF" lastPageId="5" lastPageNumber="553" pageId="4" pageNumber="552" type="materials_examined">
|
||
<paragraph id="ED6BC365FF92FFF22B8EFDD7BE25C6B8" blockId="4.[151,1437,151,681]" pageId="4" pageNumber="552">
|
||
<materialsCitation id="5DBCC938FF92FFF22B8EFDD7BE25C6B8" ID-GBIF-Occurrence="3863900302" country="United States of America" county="Smithsonian National Museum of Natural History" location="Male" municipality="Baltic" pageId="4" pageNumber="552" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="DFA01F77FF92FFF22B8EFDD7BD16C694" bold="true" box="[199,315,619,645]" pageId="4" pageNumber="552">
|
||
<typeStatus id="326F7DC7FF92FFF22B8EFDD7BD1BC694" box="[199,310,619,645]" pageId="4" pageNumber="552" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<location id="E80B95BEFF92FFF22A0DFDD7BDADC694" LSID="urn:lsid:plazi:treatment:657D7273FF95FFF32BDEFA4BB864C320:E80B95BEFF92FFF22A0DFDD7BDADC694" box="[324,384,619,645]" country="United States of America" county="Smithsonian National Museum of Natural History" municipality="Baltic" name="Male" pageId="4" pageNumber="552">Male</location>
|
||
, inclusion in
|
||
<collectingMunicipality id="0D0F591FFF92FFF22955FDD7BE72C694" box="[540,607,619,645]" pageId="4" pageNumber="552">Baltic</collectingMunicipality>
|
||
amber, housed at the
|
||
<collectingCounty id="040ABBE9FF92FFF22814FDD7B9B5C694" box="[861,1432,619,645]" pageId="4" pageNumber="552">Smithsonian National Museum of Natural History</collectingCounty>
|
||
,
|
||
<collectingCountry id="95C383F5FF92FFF22BDEFD33BCFFC6B8" box="[151,210,655,681]" name="United States of America" pageId="4" pageNumber="552">USA</collectingCountry>
|
||
, under catalog No. 775571.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="B9AB93EDFF92FFF22BDEF8F2BEBDC3BF" ID-DOI="http://doi.org/10.5281/zenodo.6854014" ID-Zenodo-Dep="6854014" httpUri="https://zenodo.org/record/6854014/files/figure.png" pageId="4" pageNumber="552" startId="4.[151,250,1870,1894]" targetBox="[151,1436,705,1847]" targetPageId="4">
|
||
<paragraph id="ED6BC365FF92FFF22BDEF8F2BEBDC3BF" blockId="4.[151,1436,1870,1966]" pageId="4" pageNumber="552">
|
||
<emphasis id="DFA01F77FF92FFF22BDEF8F2BD3EC377" bold="true" box="[151,275,1870,1894]" pageId="4" pageNumber="552">FIGURE 3.</emphasis>
|
||
<taxonomicName id="2AD4B8E6FF92FFF22A53F8F2BE99C377" authority="Pankowski & Fanti, 2022" authorityName="M. G. PANKOWSKI & FANTI" authorityYear="2022" box="[282,692,1870,1895]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="552" phylum="Arthropoda" rank="species" species="markpankowskii" status="sp. nov." subGenus="Malthodes">
|
||
<emphasis id="DFA01F77FF92FFF22A53F8F2BDA8C377" box="[282,389,1870,1894]" italics="true" pageId="4" pageNumber="552">Malthodes</emphasis>
|
||
(
|
||
<emphasis id="DFA01F77FF92FFF22ADDF8F2BDD2C377" box="[404,511,1870,1894]" italics="true" pageId="4" pageNumber="552">Malthodes</emphasis>
|
||
)
|
||
<emphasis id="DFA01F77FF92FFF22947F8F2BE99C377" box="[526,692,1870,1894]" italics="true" pageId="4" pageNumber="552">markpankowskii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DFA01F77FF92FFF229F3F8F3BF26C376" bold="true" box="[698,779,1871,1895]" pageId="4" pageNumber="552">
|
||
<taxonomicNameLabel id="C493A20CFF92FFF229F3F8F3BF26C376" box="[698,779,1871,1895]" pageId="4" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A: Holotype, habitus, dorsal view, scale bar = 0.5 mm. B: Holotype, habitus, detail of metasternum, scale bar = 0.2 mm. C: Holotype, habitus, ventral view, scale bar = 1.0 mm. D: Holotype, habitus, ventral view (detail), scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9AB93EDFF93FFF32BDEFB2DBCC9C0E0" ID-DOI="http://doi.org/10.5281/zenodo.6854018" ID-Zenodo-Dep="6854018" httpUri="https://zenodo.org/record/6854018/files/figure.png" pageId="5" pageNumber="553" startId="5.[151,250,1169,1193]" targetBox="[151,1418,181,1144]" targetPageId="5">
|
||
<paragraph id="ED6BC365FF93FFF32BDEFB2DBCC9C0E0" blockId="5.[151,1436,1169,1265]" pageId="5" pageNumber="553">
|
||
<emphasis id="DFA01F77FF93FFF32BDEFB2DBD38C0B8" bold="true" box="[151,277,1169,1193]" pageId="5" pageNumber="553">FIGURE 4.</emphasis>
|
||
<taxonomicName id="2AD4B8E6FF93FFF32A54FB2DBE96C0B8" authority="Pankowski & Fanti, 2022" authorityName="M. G. PANKOWSKI & FANTI" authorityYear="2022" box="[285,699,1169,1193]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="553" phylum="Arthropoda" rank="species" species="markpankowskii" status="sp. nov." subGenus="Malthodes">
|
||
<emphasis id="DFA01F77FF93FFF32A54FB2DBDA5C0B8" box="[285,392,1169,1193]" italics="true" pageId="5" pageNumber="553">Malthodes</emphasis>
|
||
(
|
||
<emphasis id="DFA01F77FF93FFF32AD0FB2DBE29C0B8" box="[409,516,1169,1193]" italics="true" pageId="5" pageNumber="553">Malthodes</emphasis>
|
||
)
|
||
<emphasis id="DFA01F77FF93FFF3295CFB2DBE96C0B8" box="[533,699,1169,1193]" italics="true" pageId="5" pageNumber="553">markpankowskii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DFA01F77FF93FFF3298AFB2DBF3BC0B8" bold="true" box="[707,790,1169,1193]" pageId="5" pageNumber="553">
|
||
<taxonomicNameLabel id="C493A20CFF93FFF3298AFB2DBF3BC0B8" box="[707,790,1169,1193]" pageId="5" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A: Holotype, habitus, detail of last abdominal segments (ventral view), scale bar = 0.1 mm. B: Reconstruction of last abdominal segments, scale bar = 0.1 mm. Illustration by Dr. Carly Tribull.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="ED6BC365FF93FFF32B8EFAA3B979C128" blockId="5.[151,1437,1311,1842]" box="[199,1364,1311,1338]" pageId="5" pageNumber="553">
|
||
<emphasis id="DFA01F77FF93FFF32B8EFAA3BD4BC128" bold="true" box="[199,358,1311,1337]" pageId="5" pageNumber="553">Type locality.</emphasis>
|
||
Amber mine in the Yantarny settlement, Sambian Peninsula,
|
||
<collectingRegion id="2F100D87FF93FFF32F5FFAA3B8DAC128" box="[1046,1271,1311,1337]" country="Russia" name="Kaliningrad" pageId="5" pageNumber="553">Kaliningrad Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="95C383F5FF93FFF32E4BFAA3B97DC128" box="[1282,1360,1311,1337]" name="Russia" pageId="5" pageNumber="553">Russia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="ED6BC365FF93FFF32B8EFAFFB96DC14C" blockId="5.[151,1437,1311,1842]" box="[199,1344,1347,1374]" pageId="5" pageNumber="553">
|
||
<emphasis id="DFA01F77FF93FFF32B8EFAFFBD41C14C" bold="true" box="[199,364,1347,1373]" pageId="5" pageNumber="553">Type horizon.</emphasis>
|
||
Middle Eocene (Lutetian) (47.8–41.2 Ma) to late Eocene (Priabonian) (37.8–33.9 Ma).
|
||
</paragraph>
|
||
<paragraph id="ED6BC365FF93FFF32B8EFADBB878C190" blockId="5.[151,1437,1311,1842]" box="[199,1109,1383,1410]" pageId="5" pageNumber="553">
|
||
<emphasis id="DFA01F77FF93FFF32B8EFADBBD5FC190" bold="true" box="[199,370,1383,1409]" pageId="5" pageNumber="553">Syninclusions.</emphasis>
|
||
Many stellate hairs, detritus, botanical fragments and air bubbles.
|
||
</paragraph>
|
||
<paragraph id="ED6BC365FF93FFF32B8EFA37BFCBC1FF" blockId="5.[151,1437,1311,1842]" pageId="5" pageNumber="553">
|
||
<emphasis id="DFA01F77FF93FFF32B8EFA37BDF9C1B4" bold="true" box="[199,468,1419,1445]" pageId="5" pageNumber="553">Systematic placement.</emphasis>
|
||
This new, extinct species clearly belongs to the subfamily
|
||
<taxonomicName id="2AD4B8E6FF93FFF32FCBFA37B924C1B4" authorityName="Kiesenwetter" authorityYear="1852" box="[1154,1289,1419,1445]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="553" phylum="Arthropoda" rank="subFamily" subFamily="Malthininae">Malthininae</taxonomicName>
|
||
based on its last maxillary palpomere that is globular and pointed distally. Its rounded head behind eyes, filiform antennae and strongly modified
|
||
<emphasis id="DFA01F77FF93FFF32A2FFA6FBDF6C1FC" box="[358,475,1491,1517]" italics="true" pageId="5" pageNumber="553">terminalia</emphasis>
|
||
place the new species in the genus
|
||
<taxonomicName id="2AD4B8E6FF93FFF32820FA6FBFF2C1FC" authorityName="Kiesenwetter" authorityYear="1852" box="[873,991,1491,1517]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="553" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFA01F77FF93FFF32820FA6FBFF2C1FC" box="[873,991,1491,1517]" italics="true" pageId="5" pageNumber="553">Malthodes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5CE90EEFF93FFF32B8EFA4BB9ACC2F8" pageId="5" pageNumber="553" type="diagnosis">
|
||
<paragraph id="ED6BC365FF93FFF32B8EFA4BB9ACC2F8" blockId="5.[151,1437,1311,1842]" pageId="5" pageNumber="553">
|
||
<emphasis id="DFA01F77FF93FFF32B8EFA4BBDFDC200" bold="true" box="[199,464,1527,1553]" pageId="5" pageNumber="553">Differential diagnosis.</emphasis>
|
||
No sister group of
|
||
<taxonomicName id="2AD4B8E6FF93FFF329FDFA4BBFC6C200" authority="Pankowski & Fanti, 2022" authorityName="M. G. PANKOWSKI & FANTI" authorityYear="2022" box="[692,1003,1527,1553]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="553" phylum="Arthropoda" rank="species" species="markpankowskii" status="sp. nov.">
|
||
<emphasis id="DFA01F77FF93FFF329FDFA4BBFC6C200" box="[692,1003,1527,1553]" italics="true" pageId="5" pageNumber="553">Malthodes markpankowskii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DFA01F77FF93FFF328BDFA4BB87DC200" bold="true" box="[1012,1104,1527,1553]" pageId="5" pageNumber="553">
|
||
<taxonomicNameLabel id="C493A20CFF93FFF328BDFA4BB87DC200" box="[1012,1104,1527,1553]" pageId="5" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has been found living in the Baltic region or Central Europe, and no other fossil species of
|
||
<taxonomicName id="2AD4B8E6FF93FFF32805F9A7BFEFC224" authorityName="Kiesenwetter" authorityYear="1852" box="[844,962,1563,1589]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="553" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DFA01F77FF93FFF32805F9A7BFEFC224" box="[844,962,1563,1589]" italics="true" pageId="5" pageNumber="553">Malthodes</emphasis>
|
||
</taxonomicName>
|
||
has this combination of characters (in particular the shape of the
|
||
<emphasis id="DFA01F77FF93FFF32AD3F983BE22C248" box="[410,527,1599,1625]" italics="true" pageId="5" pageNumber="553">terminalia</emphasis>
|
||
) shown by
|
||
<taxonomicName id="2AD4B8E6FF93FFF329D8F983BFE8C248" authority="Pankowski & Fanti, 2022" authorityName="M. G. PANKOWSKI & FANTI" authorityYear="2022" box="[657,965,1599,1625]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="553" phylum="Arthropoda" rank="species" species="markpankowskii" status="sp. nov.">
|
||
<emphasis id="DFA01F77FF93FFF329D8F983BFE8C248" box="[657,965,1599,1625]" italics="true" pageId="5" pageNumber="553">Malthodes markpankowskii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DFA01F77FF93FFF32882F983B809C248" bold="true" box="[971,1060,1599,1625]" pageId="5" pageNumber="553">
|
||
<taxonomicNameLabel id="C493A20CFF93FFF32882F983B809C248" box="[971,1060,1599,1625]" pageId="5" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
One fossil species that is vaguely similar is
|
||
<taxonomicName id="2AD4B8E6FF93FFF32A4EF9DFBE56C26C" authority="Fanti, 2019" authorityName="Fanti" authorityYear="2019" box="[263,635,1635,1661]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="553" phylum="Arthropoda" rank="species" species="neumanni">
|
||
<emphasis id="DFA01F77FF93FFF32A4EF9DFBDDEC26C" box="[263,499,1635,1661]" italics="true" pageId="5" pageNumber="553">Malthodes neumanni</emphasis>
|
||
<bibRefCitation id="8945BE94FF93FFF32AB2F9DFBE56C26C" author="Fanti, F." box="[507,635,1635,1661]" pageId="5" pageNumber="553" pagination="89 - 96" refId="ref3775" refString="Fanti, F. (2019) First fossil soldier beetles (Coleoptera Cantharidae) from Bitterfeld amber, Germany. Zitteliana, 93, 89 - 96." type="journal article" year="2019">Fanti, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
from Bitterfeld amber. It differs by the shorter last sternite (st9) with a more concave anterior margin (
|
||
<bibRefCitation id="8945BE94FF93FFF32ABBF93BBE5DC2B0" author="Fanti, F." box="[498,624,1671,1697]" pageId="5" pageNumber="553" pagination="89 - 96" refId="ref3775" refString="Fanti, F. (2019) First fossil soldier beetles (Coleoptera Cantharidae) from Bitterfeld amber, Germany. Zitteliana, 93, 89 - 96." type="journal article" year="2019">Fanti 2019</bibRefCitation>
|
||
), and by its shorter last tergite (tg10). The other slightly similar species is
|
||
<taxonomicName id="2AD4B8E6FF93FFF32BDEF917BDC4C2D4" authority="Fanti, 2018" authorityName="Fanti" authorityYear="2018" box="[151,489,1707,1733]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="553" phylum="Arthropoda" rank="species" species="meriae">
|
||
<emphasis id="DFA01F77FF93FFF32BDEF917BD4EC2D4" box="[151,355,1707,1733]" italics="true" pageId="5" pageNumber="553">Malthodes meriae</emphasis>
|
||
Fanti, 2018
|
||
</taxonomicName>
|
||
from Baltic amber. Compared with the new species described here,
|
||
<taxonomicName id="2AD4B8E6FF93FFF32FA0F910B971C2D4" authorityName="Fanti" authorityYear="2018" box="[1257,1372,1708,1733]" class="Insecta" family="Cantharidae" genus="Malthodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="553" phylum="Arthropoda" rank="species" species="meriae">
|
||
<emphasis id="DFA01F77FF93FFF32FA0F910B971C2D4" box="[1257,1372,1708,1733]" italics="true" pageId="5" pageNumber="553">M. meriae</emphasis>
|
||
</taxonomicName>
|
||
has a much smaller and shorter last tergite, as well as a last sternite with a more concave anterior margin (Fanti 2018).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5CE90EEFF93FFF32B8EF94FB864C320" pageId="5" pageNumber="553" type="discussion">
|
||
<paragraph id="ED6BC365FF93FFF32B8EF94FB864C320" blockId="5.[151,1437,1311,1842]" pageId="5" pageNumber="553">
|
||
<emphasis id="DFA01F77FF93FFF32B8EF94FBD16C31C" bold="true" box="[199,315,1779,1805]" pageId="5" pageNumber="553">Remarks.</emphasis>
|
||
The yellow rectangular amber piece measures approximately 26x13x
|
||
<quantity id="2A2C6E80FF93FFF32F1BF94FB8B8C31F" box="[1106,1173,1779,1806]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="553" unit="mm" value="4.0">4 mm</quantity>
|
||
and weighs
|
||
<quantity id="2A2C6E80FF93FFF32E6CF94FB9B4C31F" box="[1317,1433,1779,1806]" metricMagnitude="-3" metricUnit="kg" metricValue="1.2" pageId="5" pageNumber="553" unit="g" value="1.2">1.2 grams</quantity>
|
||
. The inclusion is complete and clearly visible except for small areas around the head.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |