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<document ID-DOI="http://doi.org/10.5281/zenodo.6232004" ID-GBIF-Dataset="80959f8f-aa73-499e-93ec-5c07458c7935" ID-GBIF-Taxon="160743180" ID-ISSN="0024-0672" ID-Zenodo-Dep="6232004" checkinTime="1576401033600" checkinUser="plazi" docAuthor="Schilthuizen, Menno &amp; Perreau, Michel" docDate="2008" docId="F97A87E2740BE66B66FAFE85FB96FC30" docLanguage="en" docName="ZM.82.19.189-210.pdf" docOrigin="Zool. Med. Leiden 82 (19)" docStyle="DocumentStyle{}" docTitle="Ptomaphaginus rufus Jeannel 1936" docType="treatment" docVersion="5" lastPageNumber="200" masterDocId="0543FF9A7400E660674CFFA5FFD2FFE5" masterDocTitle="New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)" masterLastPageNumber="210" masterPageNumber="189" pageId="11" pageNumber="200" updateTime="1645572769617" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perreau, Michel</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zool. Med. Leiden</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-01-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>82</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>189</mods:start>
<mods:end>210</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3576287</mods:identifier>
<mods:identifier type="GBIF-Dataset">80959f8f-aa73-499e-93ec-5c07458c7935</mods:identifier>
<mods:identifier type="ISSN">0024-0672</mods:identifier>
<mods:identifier type="Zenodo-Dep">3576287</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6232004" ID-GBIF-Taxon="160743180" ID-Zenodo-Dep="6232004" LSID="urn:lsid:plazi:treatment:F97A87E2740BE66B66FAFE85FB96FC30" httpUri="http://treatment.plazi.org/id/F97A87E2740BE66B66FAFE85FB96FC30" lastPageNumber="200" pageId="11" pageNumber="200">
<subSubSection box="[438,833,288,312]" pageId="11" pageNumber="200" type="nomenclature">
<paragraph blockId="11.[438,833,288,312]" box="[438,833,288,312]" pageId="11" pageNumber="200">
<heading box="[438,833,288,312]" centered="true" fontSize="10" level="3" pageId="11" pageNumber="200" reason="1">
<taxonomicName authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[438,833,288,312]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="200" phylum="Arthropoda" rank="species" species="rufus">
<emphasis bold="true" box="[438,681,288,312]" italics="true" pageId="11" pageNumber="200">Ptomaphaginus rufus</emphasis>
<bibRefCitation author="Jeannel, R." box="[687,833,288,312]" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, N. S" pageId="11" pageNumber="200" pagination="1 - 433" refId="ref10634" refString="Jeannel, R., 1936. Monographie des Catopidae. - Memoires du Museum National d'Histoire Naturelle, N. S. 1: 1 - 433." title="Monographie des Catopidae" type="book chapter" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="200" type="reference_group">
<paragraph blockId="11.[151,1119,349,589]" pageId="11" pageNumber="200">
<treatmentCitationGroup infered="true" pageId="11" pageNumber="200">
<taxonomicName authority="Jeannel, 1936: 56 - 58" authorityName="Jeannel" authorityPageNumber="56" authorityYear="1936" box="[151,524,349,368]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="200" phylum="Arthropoda" rank="species" species="rufus">
<emphasis box="[151,336,349,368]" italics="true" pageId="11" pageNumber="200">Ptomaphaginus rufus</emphasis>
<treatmentCitation author="Jeannel, R." box="[341,524,349,368]" page="56" pageId="11" pageNumber="200" year="1936">
<bibRefCitation author="Jeannel, R." box="[341,524,349,368]" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, N. S" pageId="11" pageNumber="200" pagination="1 - 433" refId="ref10634" refString="Jeannel, R., 1936. Monographie des Catopidae. - Memoires du Museum National d'Histoire Naturelle, N. S. 1: 1 - 433." title="Monographie des Catopidae" type="book chapter" year="1936">Jeannel, 1936: 56-58</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, figs 67-68.—
<typeStatus box="[654,742,349,368]" pageId="11" pageNumber="200">Holotype</typeStatus>
male ‘Type’ [round white label with red rim], ‘Bûkit Timah,
<collectingCountry box="[375,471,376,395]" name="Singapore" pageId="11" pageNumber="200">Singapore</collectingCountry>
, G.E. Bryant,
<date box="[598,652,377,396]" pageId="11" pageNumber="200" value="1909-05-07">7.v.09</date>
’ [white rectangular label], ‘G. Bryant Coll., 1919— 147’ [white rectangular label], ‘TYPE’ [white rectangular label with red print], ‘See slide 7 (Jeannel 1934)’ [white rectangular label], ‘
<taxonomicName authority="Schilthuizen &amp; Perreau, 2008" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[491,696,432,451]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="200" phylum="Arthropoda" rank="species" species="rufus" status="n. sp.">Ptomaphaginus rufus</taxonomicName>
<taxonomicNameLabel box="[700,749,432,451]" pageId="11" pageNumber="200" rank="species">n. sp.</taxonomicNameLabel>
R. Jeannel det.’ [white rectangular label, Jeannel’s hand] in BMNH [examined]. Microscope slide, left-hand side: ‘
<taxonomicName authority="Schilthuizen &amp; Perreau, 2008" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[860,1065,459,478]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="200" phylum="Arthropoda" rank="species" species="rufus" status="n. sp.">Ptomaphaginus rufus</taxonomicName>
<taxonomicNameLabel box="[1070,1119,460,479]" pageId="11" pageNumber="200" rank="species">n. sp.</taxonomicNameLabel>
Jeannel, Type, Singapoore’ [white square label, Jeannel’s hand, the word ‘Type” underlined in red], ‘B.M. 1919-147, org. copul.
<specimenCount box="[442,460,514,534]" pageId="11" pageNumber="200" type="male">♂</specimenCount>
(Slide 7, Jeannel 1934)’ [white square label, the words ‘org. copul. ♂’ in Jeannel’s hand], ‘Type’ [round white label with red rim] in BMNH [examined].—
<treatmentCitation author="Szymczakowski, W." page="99" pageId="11" pageNumber="200" year="1964">
<bibRefCitation author="Szymczakowski, W." journalOrPublisher="Acta Zoologica Cracoviensia" pageId="11" pageNumber="200" pagination="55 - 283" part="9" refId="ref11118" refString="Szymczakowski, W., 1964. Analyse systematique et zoogeographique des Catopidae (Coleoptera) de la region orientale. - Acta Zoologica Cracoviensia 9: 55 - 283." title="Analyse systematique et zoogeographique des Catopidae (Coleoptera) de la region orientale" type="journal article" year="1964">Szymczakowski 1964: 99-101</bibRefCitation>
</treatmentCitation>
,
<figureCitation box="[316,371,570,589]" captionStart="Figs 7-12" captionStartId="17.[151,190,1413,1432]" captionTargetId="figure@18.[493,888,185,983]" captionTargetPageId="18" captionText="Figs 7-12. Fig. 7, Ptomaphaginus burckhardti spec. nov., holotype; aedeagus, dorsal view. Fig. 8, P. burckhardti spec. nov., holotype; aedeagus, lateral view. Fig. 9, P. kurbatovi spec. nov., paratype (Cibodas); aedeagus, dorsal view. Fig. 10, P. kurbatovi spec. nov., paratype (Cibodas); aedeagus, lateral view. Fig. 11, P. sabahensis spec. nov., holotype; aedeagus, dorsal view. Fig. 12, P. sabahensis spec. nov., holotype; aedeagus, lateral view. All scale lines 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3576291" httpUri="https://zenodo.org/record/3576291/files/figure.png" pageId="11" pageNumber="200">fig. 11</figureCitation>
(redescription); Szymczakowski 1970: 287-289,
<figureCitation box="[813,901,570,589]" captionStart-0="Figs 7-12" captionStart-1="Figs 13-15. 13" captionStart-2="Figs 16-25. 16" captionStartId-0="17.[151,190,1413,1432]" captionStartId-1="19.[151,189,769,788]" captionStartId-2="20.[151,190,1437,1456]" captionTargetBox-1="[153,1083,201,739]" captionTargetBox-2="[151,1119,192,1408]" captionTargetId-0="figure@18.[493,888,185,983]" captionTargetId-1="figure@19.[459,777,172,750]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionText-0="Figs 7-12. Fig. 7, Ptomaphaginus burckhardti spec. nov., holotype; aedeagus, dorsal view. Fig. 8, P. burckhardti spec. nov., holotype; aedeagus, lateral view. Fig. 9, P. kurbatovi spec. nov., paratype (Cibodas); aedeagus, dorsal view. Fig. 10, P. kurbatovi spec. nov., paratype (Cibodas); aedeagus, lateral view. Fig. 11, P. sabahensis spec. nov., holotype; aedeagus, dorsal view. Fig. 12, P. sabahensis spec. nov., holotype; aedeagus, lateral view. All scale lines 100 µm." captionText-1="Figs 13-15. 13, Ptomaphaginus bryanti Jeannel, aedeagus; reproduced from Jeannel (1936). 14, Ptomaphaginus caroli spec. nov., holotype; aedeagus dorsal view. 15, Ptomaphaginus caroli spec. nov., holotype; aedeagus lateral view." captionText-2="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.3576291" figureDoi-1="http://doi.org/10.5281/zenodo.3576293" figureDoi-2="http://doi.org/10.5281/zenodo.3576295" httpUri-0="https://zenodo.org/record/3576291/files/figure.png" httpUri-1="https://zenodo.org/record/3576293/files/figure.png" httpUri-2="https://zenodo.org/record/3576295/files/figure.png" pageId="11" pageNumber="200">figs 10-20</figureCitation>
(redescription).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="200" type="materials_examined">
<paragraph blockId="11.[151,1120,625,727]" pageId="11" pageNumber="200">
Material.—
<materialsCitation ID-GBIF-Occurrence="2512737955" pageId="11" pageNumber="200">
<collectingCountry box="[264,352,625,644]" name="Malaysia" pageId="11" pageNumber="200">Malaysia</collectingCountry>
: Air Hitam forest reserve,
<quantity box="[608,657,625,644]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="200" unit="km" value="5.0">5 km</quantity>
west of Serdang (
<collectingRegion box="[832,918,625,644]" country="Malaysia" name="Selangor" pageId="11" pageNumber="200">Selangor</collectingRegion>
), in pitfall with beef,
<date box="[151,232,652,671]" pageId="11" pageNumber="200" value="1989-08">viii.1989</date>
, leg. M. Schilthuizen,
<specimenCount box="[439,473,652,672]" pageId="11" pageNumber="200" type="male">1 ♂</specimenCount>
,
<specimenCount box="[484,518,652,672]" pageId="11" pageNumber="200" type="female">1 ♀</specimenCount>
(
<collectionCode box="[529,577,652,671]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="11" pageNumber="200" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2512737962" collectingDate="1989-07-10" collectionCode="BOR" collectorName="M. Schilthuizen" location="Serdang Lama" pageId="11" pageNumber="200" specimenCount="1" specimenCount-male="1" stateProvince="Selangor">
<location LSID="urn:lsid:plazi:treatment:F97A87E2740BE66B66FAFE85FB96FC30:740C602F740BE66B651CFD29FD0AFD45" box="[592,728,652,672]" name="Serdang Lama" pageId="11" pageNumber="200" stateProvince="Selangor">Serdang Lama</location>
(
<collectingRegion box="[740,826,652,671]" country="Malaysia" name="Selangor" pageId="11" pageNumber="200">Selangor</collectingRegion>
), rubber estate,
<date box="[975,1076,652,671]" pageId="11" pageNumber="200" value="1989-07-10">
<collectingDate box="[975,1076,652,671]" pageId="11" pageNumber="200" value="1989-07-10">10.vii.1989</collectingDate>
</date>
, leg.
<collectorName box="[151,300,680,699]" pageId="11" pageNumber="200">M. Schilthuizen</collectorName>
,
<specimenCount box="[308,342,679,699]" pageId="11" pageNumber="200" type="male">1 ♂</specimenCount>
(
<collectionCode box="[352,400,680,699]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="11" pageNumber="200" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2512737978" collectingDate="1989-06-22" collectingDateMax="1989-06-27" collectingDateMin="1989-06-22" collectionCode="BOR" collectorName="M. Schilthuizen" location="Taman Negara" pageId="11" pageNumber="200" specimenCount="1" specimenCount-male="1" stateProvince="Pahang">
<location LSID="urn:lsid:plazi:treatment:F97A87E2740BE66B66FAFE85FB96FC30:740C602F740BE66B66D2FD0DFDFBFD5E" box="[414,553,680,699]" name="Taman Negara" pageId="11" pageNumber="200" stateProvince="Pahang">Taman Negara</location>
(
<collectingRegion box="[563,637,680,699]" country="Malaysia" name="Pahang" pageId="11" pageNumber="200">Pahang</collectingRegion>
), primary forest,
<date box="[797,919,680,699]" pageId="11" pageNumber="200" value="1989-06-22" valueMax="1989-06-27" valueMin="1989-06-22">
<collectingDate box="[797,919,680,699]" pageId="11" pageNumber="200" value="1989-06-22" valueMax="1989-06-27" valueMin="1989-06-22">22-27.vi.1989</collectingDate>
</date>
, leg.
<collectorName box="[966,1115,680,699]" pageId="11" pageNumber="200">M. Schilthuizen</collectorName>
,
<specimenCount box="[151,185,707,727]" pageId="11" pageNumber="200" type="male">1 ♂</specimenCount>
(
<collectionCode box="[197,245,708,727]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="11" pageNumber="200" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="200" type="discussion">
<paragraph blockId="11.[151,1119,767,981]" pageId="11" pageNumber="200">
Remarks.— This species has been recorded from several sites in
<collectingCountry box="[903,1016,767,791]" name="Singapore" pageId="11" pageNumber="200">Singapore</collectingCountry>
(
<bibRefCitation author="Jeannel, R." journalOrPublisher="Memoires du Museum National d'Histoire Naturelle, N. S" pageId="11" pageNumber="200" pagination="1 - 433" refId="ref10634" refString="Jeannel, R., 1936. Monographie des Catopidae. - Memoires du Museum National d'Histoire Naturelle, N. S. 1: 1 - 433." title="Monographie des Catopidae" type="book chapter" year="1936">Jeannel, 1936</bibRefCitation>
; Szymczakowski, 1970), which is also the provenance of the
<typeStatus box="[903,1005,799,823]" pageId="11" pageNumber="200">holotype</typeStatus>
. A female
<typeStatus box="[151,251,831,855]" pageId="11" pageNumber="200">paratype</typeStatus>
derives from the west coast of Sumatra, but
<bibRefCitation author="Szymczakowski, W." box="[761,1016,830,854]" journalOrPublisher="Acta Zoologica Cracoviensia" pageId="11" pageNumber="200" pagination="55 - 283" part="9" refId="ref11118" refString="Szymczakowski, W., 1964. Analyse systematique et zoogeographique des Catopidae (Coleoptera) de la region orientale. - Acta Zoologica Cracoviensia 9: 55 - 283." title="Analyse systematique et zoogeographique des Catopidae (Coleoptera) de la region orientale" type="journal article" year="1964">Szymczakowski (1964)</bibRefCitation>
suspects this specimen may actually belong to the related
<taxonomicName authority="Szymczakowski. Our" authorityName="Szymczakowski. Our" authorityYear="1964" box="[702,1048,862,886]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="200" phylum="Arthropoda" rank="species" species="tarsalis">
<emphasis box="[702,803,862,886]" italics="true" pageId="11" pageNumber="200">P. tarsalis</emphasis>
Szymczakowski. Our
</taxonomicName>
material provides the first localities outside of
<collectingCountry box="[617,733,894,918]" name="Singapore" pageId="11" pageNumber="200">Singapore</collectingCountry>
: in the Malaysian states of Pahang and Selangor, which suggests the species is more widely distributed across the Malay Peninsula. The new specimens closely match Szymczakowski’s (1970) redescription.
</paragraph>
</subSubSection>
</treatment>
</document>
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