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<document id="BB9E6706882A688DD17E9D9FEFD134C7" ID-CLB-Dataset="132131" ID-DOI="10.5962/bhl.part.150202" ID-GBIF-Dataset="bbe4d5b4-7c73-424d-b871-74880c01bb45" ID-ISSN="0035-418X" ID-Zenodo-Dep="5828663" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641578883276" checkinUser="carolina" docAuthor="Arredondo, Nathalia J. &amp; Gil de Pertierra, Alicia A." docDate="2012" docId="F96A8786D541FFF1FF2EFC3AD21EBB70" docLanguage="en" docName="RevSuisseZool.119.4.425-439.pdf.imf" docOrigin="Revue suisse de Zoologie 119 (4)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Neoechinorhynchus Stiles &amp; Hassall 1905" docType="key" docVersion="9" lastPageNumber="437" masterDocId="0553FFFED54AFFFDFFB9FF83D63DB86B" masterDocTitle="A new species of Neoechinorhynchus (Eoacanthocephala: Neoechinorhynchidae) from Pachyurus bonariensis (Perciformes: Sciaenidae) from the Paraná River basin in Argentina, with comments on two other species of the genus" masterLastPageNumber="439" masterPageNumber="425" pageNumber="436" updateTime="1699296794393" updateUser="ExternalLinkService">
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<mods:title id="FFB19E927D66C91AB08A21AA28A09910">A new species of Neoechinorhynchus (Eoacanthocephala: Neoechinorhynchidae) from Pachyurus bonariensis (Perciformes: Sciaenidae) from the Paraná River basin in Argentina, with comments on two other species of the genus</mods:title>
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<treatment id="F96A8786D541FFF1FF2EFC3AD21EBB70" ID-DOI="http://doi.org/10.5281/zenodo.7037164" ID-GBIF-Taxon="199862959" ID-Zenodo-Dep="7037164" LSID="urn:lsid:plazi:treatment:F96A8786D541FFF1FF2EFC3AD21EBB70" httpUri="http://treatment.plazi.org/id/F96A8786D541FFF1FF2EFC3AD21EBB70" lastPageId="12" lastPageNumber="437" pageId="11" pageNumber="436">
<subSubSection id="39D9651BD541FFF6FF2EFC3AD52DBBBB" box="[151,784,952,977]" pageId="11" pageNumber="436" type="nomenclature">
<paragraph id="717C3690D541FFF6FF2EFC3AD52DBBBB" blockId="11.[151,784,952,977]" box="[151,784,952,977]" pageId="11" pageNumber="436">
<heading id="2A3481FCD541FFF6FF2EFC3AD52DBBBB" box="[151,784,952,977]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="436" reason="2">
Key to the South American species of
<taxonomicName id="B6C34D13D541FFF6FD8CFC3BD535BBBB" ID-CoL="63DTQ" authorityName="Stiles &amp; Hassall" authorityYear="1905" box="[565,776,952,976]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="11" pageNumber="436" phylum="Acanthocephala" rank="genus">
<emphasis id="43B7EA82D541FFF6FD8CFC3BD535BBBB" box="[565,776,952,976]" italics="true" pageId="11" pageNumber="436">Neoechinorhynchus</emphasis>
</taxonomicName>
:
</heading>
</paragraph>
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<subSubSection id="39D9651BD541FFF1FF2EFC63D21EBB70" lastPageId="12" lastPageNumber="437" pageId="11" pageNumber="436" type="key">
<keyStep id="CA372E35D541FFF6FF2EFC63D20CBC13" pageId="11" pageNumber="436">
<paragraph id="717C3690D541FFF6FF2EFC63D213BC53" blockId="11.[151,1073,992,1656]" pageId="11" pageNumber="436">
<keyLead id="CA3295A5D541FFF6FF2EFC63D213BC53" pageId="11" pageNumber="436">
1a. Eggs with concentric membranes, without polar prolongations of the fertilization membrane; subgenus
<taxonomicName id="B6C34D13D541FFF6FDEAFB83D725BC53" ID-CoL="63DTQ" authority="Stiles &amp; Hassall, 1905" authorityName="Stiles &amp; Hassall" authorityYear="1905" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="11" pageNumber="436" phylum="Acanthocephala" rank="genus">
<emphasis id="43B7EA82D541FFF6FDEAFB83D51BBC73" box="[595,806,1024,1048]" italics="true" pageId="11" pageNumber="436">Neoechinorhynchus</emphasis>
Stiles &amp; Hassall, 1905
</taxonomicName>
....................................................... 2
</keyLead>
</paragraph>
<paragraph id="717C3690D541FFF6FF2EFBC3D20CBC13" blockId="11.[151,1073,992,1656]" pageId="11" pageNumber="436">
<keyLead id="CA3295A5D541FFF6FF2EFBC3D20CBC13" pageId="11" pageNumber="436">
1b. Eggs with polar prolongations of the fertilization membrane; subgenus
<taxonomicName id="B6C34D13D541FFF6FF5AFBE3D425BC13" ID-CoL="7P5JK" authority="Van Cleave, 1928" authorityName="Van Cleave" authorityYear="1928" box="[227,536,1120,1144]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="11" pageNumber="436" phylum="Acanthocephala" rank="subGenus" subGenus="Hebesoma">
<emphasis id="43B7EA82D541FFF6FF5AFBE3D769BC13" box="[227,340,1120,1144]" italics="true" pageId="11" pageNumber="436">Hebesoma</emphasis>
Van Cleave, 1928
</taxonomicName>
..................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CA372E35D541FFF6FF2EFB03D20CBC93" pageId="11" pageNumber="436">
<paragraph id="717C3690D541FFF6FF2EFB03D20CBCD3" blockId="11.[151,1073,992,1656]" pageId="11" pageNumber="436">
<keyLead id="CA3295A5D541FFF6FF2EFB03D20CBCD3" pageId="11" pageNumber="436">2a. Trunk ovoid, elliptical or fusiform, swollen equatorially; proboscis without apical organ............................................ 3</keyLead>
</paragraph>
<paragraph id="717C3690D541FFF6FF2EFB43D20CBC93" blockId="11.[151,1073,992,1656]" pageId="11" pageNumber="436">
<keyLead id="CA3295A5D541FFF6FF2EFB43D20CBC93" pageId="11" pageNumber="436">2b. Trunk elongated, swollen anteriorly; proboscis with or without apical organ....................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="CA372E35D541FFF6FF2EFA83D20CBD13" pageId="11" pageNumber="436">
<paragraph id="717C3690D541FFF6FF2EFA83D20CBD53" blockId="11.[151,1073,992,1656]" pageId="11" pageNumber="436">
<keyLead id="CA3295A5D541FFF6FF2EFA83D20CBD53" pageId="11" pageNumber="436">3a. Lemnisci much longer than proboscis receptacle (twice or more); male reproductive system occupies about 50% of trunk..................... 4</keyLead>
</paragraph>
<paragraph id="717C3690D541FFF6FF2EFAC3D20CBD13" blockId="11.[151,1073,992,1656]" pageId="11" pageNumber="436">
<keyLead id="CA3295A5D541FFF6FF2EFAC3D20CBD13" pageId="11" pageNumber="436">3b. Lemnisci longer than proboscis receptacle; male reproductive system occupies more than 60% of trunk.................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="CA372E35D541FFF6FF2EFA03D20CBE53" pageId="11" pageNumber="436">
<paragraph id="717C3690D541FFF6FF2EFA03D20CBDB3" blockId="11.[151,1073,992,1656]" pageId="11" pageNumber="436">
<keyLead id="CA3295A5D541FFF6FF2EFA03D20CBDB3" pageId="11" pageNumber="436">
4a. Sexual dimorphisms present; neck twice longer than the proboscis; lemnisci subequal, overlapping anterior testes; eggs drop-shaped...............................
<taxonomicName id="B6C34D13D541FFF6FD82FA42D20CBDB3" ID-CoL="7VZHD" authority="Brasil-Sato &amp; Pavanelli, 1998" authorityName="Brasil-Sato &amp; Pavanelli" authorityYear="1998" box="[571,1073,1472,1496]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="11" pageNumber="425" phylum="Acanthocephala" rank="species" species="pimelodi" subGenus="Neoechinorhynchus">
<emphasis id="43B7EA82D541FFF6FD82FA42D470BDB3" box="[571,589,1473,1496]" italics="true" pageId="11" pageNumber="436">N</emphasis>
. (
<emphasis id="43B7EA82D541FFF6FDDAFA42D448BDB3" box="[611,629,1473,1496]" italics="true" pageId="11" pageNumber="436">N</emphasis>
.)
<emphasis id="43B7EA82D541FFF6FD35FA43D4D4BDB3" box="[652,745,1472,1496]" italics="true" pageId="11" pageNumber="436">pimelodi</emphasis>
Brasil-Sato &amp; Pavanelli, 1998
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="717C3690D541FFF6FF2EFA63D20CBE53" blockId="11.[151,1073,992,1656]" pageId="11" pageNumber="436">
<keyLead id="CA3295A5D541FFF6FF2EFA63D20CBE53" pageId="11" pageNumber="436">
4b. Sexual dimorphisms absent; neck short; lenmisci unequal, reaching level of testes, but not overlapping them; eggs elongate.........................................
<emphasis id="43B7EA82D541FFF6FD97F9A2D47DBE53" box="[558,576,1569,1592]" italics="true" pageId="11" pageNumber="436">N</emphasis>
.
<taxonomicName id="B6C34D13D541FFF6FDF7F9A3D20CBE53" ID-CoL="7VZHH" authority="Nickol &amp; Thatcher, 1971" authorityName="Nickol &amp; Thatcher" authorityYear="1971" box="[590,1073,1568,1592]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="11" pageNumber="436" phylum="Acanthocephala" rank="species" species="prochilodorum" subGenus="Neoechinorhynchus">
(
<emphasis id="43B7EA82D541FFF6FDEEF9A2D454BE53" box="[599,617,1569,1592]" italics="true" pageId="11" pageNumber="436">N</emphasis>
.)
<emphasis id="43B7EA82D541FFF6FDC6F9A3D523BE53" box="[639,798,1568,1592]" italics="true" pageId="11" pageNumber="436">prochilodorum</emphasis>
Nickol &amp; Thatcher, 1971
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CA372E35D541FFF1FF2EF9C3D21FB970" lastPageId="12" lastPageNumber="437" pageId="11" pageNumber="436">
<paragraph id="717C3690D541FFF6FF2EF9C3D20CBE13" blockId="11.[151,1073,992,1656]" pageId="11" pageNumber="436">
<keyLead id="CA3295A5D541FFF6FF2EF9C3D20CBE13" pageId="11" pageNumber="436">
5a. Neck short; lemnisci slightly longer than proboscis receptacle; cement gland almost same size than testes.
<emphasis id="43B7EA82D541FFF6FDE7F9E2D44DBE13" box="[606,624,1633,1656]" italics="true" pageId="11" pageNumber="436">N</emphasis>
.
<taxonomicName id="B6C34D13D541FFF6FDC5F9E3D20CBE13" ID-CoL="7VZHC" authority="Machado Filho, 1959" authorityName="Machado Filho" authorityYear="1959" box="[636,1073,1632,1656]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="11" pageNumber="436" phylum="Acanthocephala" rank="species" species="paraguayensis" subGenus="Neoechinorhynchus">
(
<emphasis id="43B7EA82D541FFF6FD3CF9E2D4AABE13" box="[645,663,1633,1656]" italics="true" pageId="11" pageNumber="436">N</emphasis>
.)
<emphasis id="43B7EA82D541FFF6FD15F9E2D575BE13" box="[684,840,1633,1656]" italics="true" pageId="11" pageNumber="436">paraguayensis</emphasis>
Machado Filho, 1959
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="717C3690D546FFF1FF31FF60D21FB970" blockId="12.[136,1059,227,796]" pageId="12" pageNumber="437">
<keyLead id="CA3295A5D546FFF1FF31FF60D21FB970" pageId="12" pageNumber="437">
5b. Neck long; lemnisci much longer than proboscis receptacle; cement gland bigger than testes................
<emphasis id="43B7EA82D546FFF1FD10FE87D486B970" box="[681,699,260,283]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.
<taxonomicName id="B6C34D13D546FFF1FD71FE80D21FB970" ID-CoL="7VZHM" authority="Thatcher, 1981" authorityName="Thatcher" authorityYear="1981" box="[712,1058,259,283]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="12" pageNumber="437" phylum="Acanthocephala" rank="species" species="pterodoridis" subGenus="Neoechinorhynchus">
(
<emphasis id="43B7EA82D546FFF1FD68FE87D4DEB970" box="[721,739,260,283]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.)
<emphasis id="43B7EA82D546FFF1FD40FE80D541B970" box="[761,892,259,283]" italics="true" pageId="12" pageNumber="437">pterodoridis</emphasis>
Thatcher, 1981
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CA372E35D546FFF1FF31FEA0D21FB9F0" pageId="12" pageNumber="437">
<paragraph id="717C3690D546FFF1FF31FEA0D21FB930" blockId="12.[136,1059,227,796]" pageId="12" pageNumber="437">
<keyLead id="CA3295A5D546FFF1FF31FEA0D21FB930" pageId="12" pageNumber="437">
6a. Proboscis with apical organ not observed; tegument with 1–2 dorsally and 1–3 ventral giant nuclei...............
<emphasis id="43B7EA82D546FFF1FD6BFEC7D4D9B930" box="[722,740,324,347]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.
<taxonomicName id="B6C34D13D546FFF1FD48FEC0D21FB930" ID-CoL="7VZJ7" authority="Vizcaino, 1992" authorityName="Vizcaino" authorityYear="1992" box="[753,1058,323,347]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="12" pageNumber="437" phylum="Acanthocephala" rank="species" species="villoldoi" subGenus="Neoechinorhynchus">
(
<emphasis id="43B7EA82D546FFF1FD43FEC7D531B930" box="[762,780,324,347]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.)
<emphasis id="43B7EA82D546FFF1FC9BFEC0D546B930" box="[802,891,323,347]" italics="true" pageId="12" pageNumber="437">villoldoi</emphasis>
<bibRefCitation id="15524B61D546FFF1FC38FEC0D21FB930" author="VIZCAINO, S. I." box="[897,1058,323,347]" journalOrPublisher="Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Serie Zoologia" pageId="12" pageNumber="437" pagination="179 - 184" part="63" refId="ref8396" refString="VIZCAINO, S. I. 1992. Especie nueva del genero Neoechinorhynchus (Acanthocephala-Neoechinorhynchidae) parasita del peces de Argentina. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Serie Zoologia 63: 179 - 184." title="Especie nueva del genero Neoechinorhynchus (Acanthocephala-Neoechinorhynchidae) parasita del peces de Argentina" type="journal article" year="1992">Vizcaíno, 1992</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="717C3690D546FFF1FF31FEE0D21FB9F0" blockId="12.[136,1059,227,796]" pageId="12" pageNumber="437">
<keyLead id="CA3295A5D546FFF1FF31FEE0D21FB9F0" pageId="12" pageNumber="437">6b. Probosis with apical organ; tegument with 5 dorsally and 1–2 ventral giant nuclei.................................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="CA372E35D546FFF1FF31FE27D21FB990" pageId="12" pageNumber="437">
<paragraph id="717C3690D546FFF1FF31FE27D21FB9B0" blockId="12.[136,1059,227,796]" pageId="12" pageNumber="437">
<keyLead id="CA3295A5D546FFF1FF31FE27D21FB9B0" pageId="12" pageNumber="437">
7a. Dorsal tegument with 4 prominent pre-equatorial giant nuclei and the fifth post-equatorial..........
<emphasis id="43B7EA82D546FFF1FD82FE47D470B9B0" box="[571,589,452,475]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.
<taxonomicName id="B6C34D13D546FFF1FDE3FE40D21FB9B0" ID-CoL="7VZH3" authority="Machado Filho, 1954" authorityName="Machado Filho" authorityYear="1954" box="[602,1058,451,475]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="12" pageNumber="437" phylum="Acanthocephala" rank="species" species="macronucleatus" subGenus="Neoechinorhynchus">
(
<emphasis id="43B7EA82D546FFF1FDDAFE47D448B9B0" box="[611,629,452,475]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.)
<emphasis id="43B7EA82D546FFF1FD35FE40D508B9B0" box="[652,821,451,475]" italics="true" pageId="12" pageNumber="437">macronucleatus</emphasis>
<bibRefCitation id="15524B61D546FFF1FC85FE40D21FB9B0" author="Machado Filho" box="[828,1058,451,475]" firstAuthor="Machado Filho" pageId="12" pageNumber="437" pagination="55 - 57" refId="ref7895" refString="MACHADO FILHO, D. A. 1954. Uma nova especie do genero Neoechinorhynchus (Hamann) (Neoechinorhynchidae, Acanthocephala). Revista Brasileira de Biologia 14: 55 - 57." type="journal article" year="1954">Machado Filho, 1954</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="717C3690D546FFF1FF31FE67D21FB990" blockId="12.[136,1059,227,796]" box="[136,1058,483,508]" pageId="12" pageNumber="437">
<keyLead id="CA3295A5D546FFF1FF31FE67D21FB990" box="[136,1058,483,508]" pageId="12" pageNumber="437">7b. Dorsal tegument with giant nuclei not clustered in pre-equatorial region.... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="CA372E35D546FFF1FF31FD80D21FBAD0" pageId="12" pageNumber="437">
<paragraph id="717C3690D546FFF1FF31FD80D21FBA30" blockId="12.[136,1059,227,796]" pageId="12" pageNumber="437">
<keyLead id="CA3295A5D546FFF1FF31FD80D21FBA30" pageId="12" pageNumber="437">
8a. Apical organ about half length than proboscis; elongated cement gland, separated for a distance of testes. Coiled vagina associated to paravaginal muscles........................
<emphasis id="43B7EA82D546FFF1FD6BFDC7D4D9BA30" box="[722,740,580,603]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.
<taxonomicName id="B6C34D13D546FFF1FD4BFDC0D21FBA30" ID-CoL="7VZG9" authority="Golvan, 1956" authorityName="Golvan" authorityYear="1956" box="[754,1058,579,603]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="12" pageNumber="437" phylum="Acanthocephala" rank="species" species="buttnerae" subGenus="Neoechinorhynchus">
(
<emphasis id="43B7EA82D546FFF1FD42FDC7D530BA30" box="[763,781,580,603]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.)
<emphasis id="43B7EA82D546FFF1FC9AFDC0D5B4BA30" box="[803,905,579,603]" italics="true" pageId="12" pageNumber="437">buttnerae</emphasis>
Golvan, 1956
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="717C3690D546FFF1FF31FDE0D21FBAD0" blockId="12.[136,1059,227,796]" pageId="12" pageNumber="437">
<keyLead id="CA3295A5D546FFF1FF31FDE0D21FBAD0" pageId="12" pageNumber="437">
8b. Apical organ large, almost same length than proboscis; elongated cement gland overlaps testes. Proboscis with two lateral hooks larger than other in first circle...........................
<emphasis id="43B7EA82D546FFF1FD6DFD27D4DBBAD0" box="[724,742,676,699]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.
<taxonomicName id="B6C34D13D546FFF1FD4AFD20D21FBAD0" ID-CoL="7VZGK" authority="Noronha, 1973" authorityName="Noronha" authorityYear="1973" box="[755,1058,675,699]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="12" pageNumber="437" phylum="Acanthocephala" rank="species" species="curemai" subGenus="Neoechinorhynchus">
(
<emphasis id="43B7EA82D546FFF1FD45FD27D533BAD0" box="[764,782,676,699]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.)
<emphasis id="43B7EA82D546FFF1FC9CFD27D546BAD0" box="[805,891,676,699]" italics="true" pageId="12" pageNumber="437">curemai</emphasis>
Noronha, 1973
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="717C3690D546FFF1FF31FD40D21EBB70" blockId="12.[136,1059,227,796]" pageId="12" pageNumber="437">
<keyLead id="CA3295A5D546FFF1FF31FD40D21EBB70" pageId="12" pageNumber="437">
<keyStep id="CA372E35D546FFF1FF31FD40D21EBB70" pageId="12" pageNumber="437">
9a. Proboscis with prominent apical organ; neck relatively long, trunk elongated, swollen anteriorly; male reproductive system 79%, female reproductive system 32%.........................
<emphasis id="43B7EA82D546FFF1FCB2FC87D520BB70" box="[779,797,772,795]" italics="true" pageId="12" pageNumber="437">N</emphasis>
.
<taxonomicName id="B6C34D13D546FFF1FC92FC80D5DFBB70" authorityName="Arredondo &amp; Gil de Pertierra" authorityYear="2012" box="[811,994,771,795]" class="Eoacanthocephala" family="Neoechinorhynchidae" genus="Neoechinorhynchus" kingdom="Animalia" order="Neoechinorhynchida" pageId="12" pageNumber="437" phylum="Acanthocephala" rank="species" species="colastinense" status="sp. nov." subGenus="Hebesoma">
(
<emphasis id="43B7EA82D546FFF1FC8DFC87D57ABB70" box="[820,839,772,795]" italics="true" pageId="12" pageNumber="437">H</emphasis>
.)
<emphasis id="43B7EA82D546FFF1FCE7FC80D5DFBB70" box="[862,994,771,795]" italics="true" pageId="12" pageNumber="437">colastinense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="588457F9D546FFF1FC50FC87D21EBB70" box="[1001,1059,772,795]" pageId="12" pageNumber="437" rank="species">sp. n.</taxonomicNameLabel>
</keyStep>
</keyLead>
</paragraph>
</subSubSection>
</treatment>
</document>
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