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<document id="29012A8CABED2DAF5F594B3AEA0F71D3" ID-CLB-Dataset="27448" ID-DOI="10.11646/zootaxa.4559.2.6" ID-GBIF-Dataset="fe48cda7-f0d3-4a74-88f7-6bb854e9367a" ID-ISSN="1175-5326" ID-Zenodo-Dep="2627008" ID-ZooBank="808CB774-BC1B-428F-AEB6-8266AD3984F1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554321623029" checkinUser="plazi" docAuthor="Zhovnerchuk, Olga &amp; Auger, Philippe" docDate="2019" docId="F67E4822CA73F33810A5EA30D449F907" docLanguage="en" docName="zootaxa.4559.2.6.pdf" docOrigin="Zootaxa 4559 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Georgiobia skuthiae Zhovnerchuk &amp; Auger 2019, n. sp." docType="treatment" docVersion="8" lastPageNumber="332" masterDocId="0A47305ACA77F3331032EC60D62EFFDC" masterDocTitle="Spider mites (Acari: Tetranychidae) from the Black Sea Biosphere Reserve (Ukraine): faunistic survey, reinstatement of the genus Georgiobia Wainstein, 1960 and description of a new species" masterLastPageNumber="338" masterPageNumber="321" pageNumber="325" updateTime="1698717451691" updateUser="ExternalLinkService">
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<mods:title id="D370C06FF2C09BE1430200E94F7276C4">Spider mites (Acari: Tetranychidae) from the Black Sea Biosphere Reserve (Ukraine): faunistic survey, reinstatement of the genus Georgiobia Wainstein, 1960 and description of a new species</mods:title>
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<mods:namePart id="9177A694017B89ED48C192342B04A9ED">Zhovnerchuk, Olga</mods:namePart>
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<mods:namePart id="A8FB58B8F41A76802D5023871F1C3FA5">Auger, Philippe</mods:namePart>
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<mods:date id="0D211CBC494DE4FF145F2D17BB19A524">2019</mods:date>
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<treatment id="F67E4822CA73F33810A5EA30D449F907" ID-DOI="http://doi.org/10.5281/zenodo.5921278" ID-GBIF-Taxon="154534171" ID-Zenodo-Dep="5921278" LSID="urn:lsid:plazi:treatment:F67E4822CA73F33810A5EA30D449F907" httpUri="http://treatment.plazi.org/id/F67E4822CA73F33810A5EA30D449F907" lastPageId="11" lastPageNumber="332" pageId="4" pageNumber="325">
<subSubSection id="36CDAABFCA73F33710A5EA30D7F8F9B6" box="[151,470,1616,1642]" pageId="4" pageNumber="325" type="nomenclature">
<paragraph id="7E68F934CA73F33710A5EA30D7F8F9B6" blockId="4.[151,470,1616,1675]" box="[151,470,1616,1642]" pageId="4" pageNumber="325">
<heading id="25204E58CA73F33710A5EA30D7F8F9B6" bold="true" box="[151,470,1616,1642]" fontSize="11" level="1" pageId="4" pageNumber="325" reason="1">
<emphasis id="4CA32526CA73F33710A5EA30D7F8F9B6" bold="true" box="[151,470,1616,1642]" pageId="4" pageNumber="325">
<taxonomicName id="B9D782B7CA73F33710A5EA30D7A5F9B6" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[151,395,1616,1642]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="325" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA73F33710A5EA30D7A5F9B6" bold="true" box="[151,395,1616,1642]" italics="true" pageId="4" pageNumber="325">Georgiobia skuthiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5790985DCA73F33711A1EA30D7F8F9B6" box="[403,470,1616,1642]" pageId="4" pageNumber="325" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="36CDAABFCA73F33710A5EA12D701F957" box="[151,303,1650,1675]" pageId="4" pageNumber="325" type="description">
<paragraph id="7E68F934CA73F33710A5EA12D701F957" blockId="4.[151,470,1616,1675]" box="[151,303,1650,1675]" pageId="4" pageNumber="325">
(
<figureCitation id="E6ECE5B1CA73F33710ACEA12D70BF957" box="[158,293,1650,1675]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="7.[151,250,1971,1993]" captionStartId-1="8.[151,250,1971,1993]" captionStartId-2="9.[151,250,1975,1997]" captionStartId-3="10.[151,250,1101,1123]" captionStartId-4="11.[151,250,1093,1115]" captionStartId-5="12.[151,250,755,777]" captionTargetBox-0="[179,1415,195,1955]" captionTargetBox-1="[279,1436,238,1924]" captionTargetBox-2="[190,1400,196,1958]" captionTargetBox-3="[150,1436,214,1066]" captionTargetBox-4="[150,1436,195,1071]" captionTargetBox-5="[166,1433,196,713]" captionTargetId-0="figure@7.[176,1415,195,1955]" captionTargetId-1="figure@8.[151,1437,238,1924]" captionTargetId-2="figure@9.[190,1400,193,1958]" captionTargetId-3="figure@10.[150,1436,214,1066]" captionTargetId-4="figure@11.[150,1436,195,1071]" captionTargetId-5="figure@12.[166,1433,191,738]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionText-0="FIGURE 1. Georgiobia skuthiae n. sp., female. A. dorsal aspect, B. detail of medial propodosomal striation, C. dorsal body seta (e 2). Scale bars = 100 µm (A), 10 µm (B), 25 µm (C)." captionText-1="FIGURE 2. Georgiobia skuthiae n. sp., female. A. tarsus I, B. tibia I, C. empodium and claws I, D. empodium and claws II, E. coxisternal setae 1b, 1c (from left to right). Scale bars = 25 µm (AB), 10 µm (CE)." captionText-2="FIGURE 3. Georgiobia skuthiae n. sp., female. A. tarsus and tibia II, B. tarsus and tibia III, C. tarsus and tibia IV, D. palptarsus and palptibia, E. peritremal enlargement, F. ventral striation. Scale bars = 25 µm (AC), 10 µm (DE), 50 µm (F)." captionText-3="FIGURE 4. Georgiobia skuthiae n. sp., Deutonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation, D. peritreme. Scale bars: 100 µm (A), 25 µm (BD)." captionText-4="FIGURE 5. Georgiobia skuthiae n. sp., Protonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation. Scale bars: 50 µm (A), 25 µm (B, C)" captionText-5="FIGURE 6. Schematic representation of the underside of tarsi I and II showing setal insertions in the protonymphs, deutonymphs and adult female of Georgiobia skuthiae n. sp." httpUri-0="https://zenodo.org/record/2627010/files/figure.png" httpUri-1="https://zenodo.org/record/2627012/files/figure.png" httpUri-2="https://zenodo.org/record/2627014/files/figure.png" httpUri-3="https://zenodo.org/record/2627016/files/figure.png" httpUri-4="https://zenodo.org/record/2627018/files/figure.png" httpUri-5="https://zenodo.org/record/2627020/files/figure.png" pageId="4" pageNumber="325">Figures 16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="36CDAABFCA73F33710A5EADAD79FF8BF" pageId="4" pageNumber="325" type="materials_examined">
<paragraph id="7E68F934CA73F33710A5EADAD79FF8BF" blockId="4.[151,1436,1722,2035]" pageId="4" pageNumber="325">
<emphasis id="4CA32526CA73F33710A5EADAD767F90F" bold="true" box="[151,329,1722,1747]" pageId="4" pageNumber="325">Type material.</emphasis>
<typeStatus id="A16C4796CA73F3371166EADAD792F90F" box="[340,444,1722,1747]" pageId="4" pageNumber="325" type="holotype">Holotype</typeStatus>
(female),
<specimenCount id="68D132BDCA73F337120AEADBD48FF90F" box="[568,673,1722,1747]" pageId="4" pageNumber="325" type="female">1 female</specimenCount>
, 1 deutonymph and 1 protonymph
<typeStatus id="A16C4796CA73F3371473EADBD283F90F" box="[1089,1197,1723,1747]" pageId="4" pageNumber="325" type="paratype">paratypes</typeStatus>
from
<taxonomicName id="B9D782B7CA73F33714C8EADBD35FF90E" box="[1274,1393,1723,1746]" class="Magnoliopsida" family="Poaceae" genus="Agropyron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="4" pageNumber="325" phylum="Tracheophyta" rank="genus">
<emphasis id="4CA32526CA73F33714C8EADBD35FF90E" box="[1274,1393,1723,1746]" italics="true" pageId="4" pageNumber="325">Agropyron</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="B9D782B7CA73F33710ACEABFD72EF92B" authorityName="" authorityYear="1895" box="[158,256,1759,1783]" class="Magnoliopsida" family="Poaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="4" pageNumber="325" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
), steppe area of
<collectingCountry id="06C0B9A4CA73F33711D2EABED413F92B" box="[480,573,1758,1783]" name="Ukraine" pageId="4" pageNumber="325">Ukraine</collectingCountry>
,
<collectingRegion id="BC1337D6CA73F3371261EABED49CF92B" box="[595,690,1758,1783]" country="Ukraine" name="Kherson" pageId="4" pageNumber="325">Kherson</collectingRegion>
region, Gola Prystan district, Black Sea biosphere reserve, Solenoozerny (
<geoCoordinate id="1BE39FF3CA73F3371170EB62D7FCF8C7" box="[322,466,1794,1819]" direction="north" orientation="latitude" pageId="4" pageNumber="325" precision="15" value="46.509724">46°3035N</geoCoordinate>
,
<geoCoordinate id="1BE39FF3CA73F33711EDEB62D442F8C7" box="[479,620,1794,1819]" direction="east" orientation="longitude" pageId="4" pageNumber="325" precision="15" value="31.901112">31°5404E</geoCoordinate>
,
<quantity id="B92F54D1CA73F337124BEB63D488F8C6" box="[633,678,1795,1819]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="325" unit="m" value="1.0">1 m</quantity>
a.s.l., flood plain),
<date id="0A69DFF4CA73F33713B4EB62D226F8C7" box="[902,1032,1794,1819]" pageId="4" pageNumber="325" value="2008-06-08">08/06/2008</date>
, coll. S. Pogrebnyak.
<typeStatus id="A16C4796CA73F3371531EB62D345F8C7" box="[1283,1387,1794,1819]" pageId="4" pageNumber="325" type="holotype">Holotype</typeStatus>
and
<typeStatus id="A16C4796CA73F33710A5EB47D72CF8E3" box="[151,258,1831,1855]" pageId="4" pageNumber="325" type="paratype">paratypes</typeStatus>
deposited in the collection I.I. Schmalhausen Institute of Zoology of National Academy of Sciences of
<collectingCountry id="06C0B9A4CA73F33710A5EB2AD6DFF8BF" box="[151,241,1866,1891]" name="Ukraine" pageId="4" pageNumber="325">Ukraine</collectingCountry>
(
<collectingRegion id="BC1337D6CA73F3371132EB2AD714F8BF" box="[256,314,1866,1891]" country="Ukraine" name="Kyiv" pageId="4" pageNumber="325">Kyiv</collectingRegion>
,
<collectingCountry id="06C0B9A4CA73F3371177EB2AD78BF8BF" box="[325,421,1866,1891]" name="Ukraine" pageId="4" pageNumber="325">Ukraine</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="36CDAABFCA73F33610F5EB0ED4CCFF24" lastPageId="5" lastPageNumber="326" pageId="4" pageNumber="325" type="diagnosis">
<paragraph id="7E68F934CA73F33610F5EB0ED4CCFF24" blockId="4.[151,1436,1722,2035]" lastBlockId="5.[151,1437,151,1409]" lastPageId="5" lastPageNumber="326" pageId="4" pageNumber="325">
<emphasis id="4CA32526CA73F33710F5EB0ED785F85B" bold="true" box="[199,427,1902,1927]" pageId="4" pageNumber="325">Diagnosis (female).</emphasis>
Leg I very long, twice as long as body length, about
<quantity id="B92F54D1CA73F3371439EB0ED24AF85A" box="[1035,1124,1902,1927]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="325" unit="mm" value="1.5">1.5 mm</quantity>
, with tarsus almost retrorse distally. Tarsal tectal setae (
<emphasis id="4CA32526CA73F33711EFEBF4D7DFF877" box="[477,497,1940,1963]" italics="true" pageId="4" pageNumber="325">tc</emphasis>
-
<emphasis id="4CA32526CA73F3371236EBF4D436F877" box="[516,536,1940,1963]" italics="true" pageId="4" pageNumber="325">tc</emphasis>
) absent, first primiventral setae (
<emphasis id="4CA32526CA73F3371383EBF4D5E2F877" box="[945,972,1940,1963]" italics="true" pageId="4" pageNumber="325">pv</emphasis>
) absent on tarsus I. Dorsal body setae slender, stout, serrate, inserted on quite strong, granulate, tubercles; inner setae far longer than longitudinal distance to base of setae next behind. Integument with medioprodorsal striations with geometric alveoli; medial striation of hysterosoma transverse, longitudinal to wrinkled between inner and lateral setae. Macrosetae on tibia III and IV as long as or slightly longer than tibia III and IV, respectively. Peritreme ending in a well-developed suboval shaped anastomosis, stylophore emarginate.
</paragraph>
</subSubSection>
<subSubSection id="36CDAABFCA72F33910F5ED64D7A5FA84" lastPageId="10" lastPageNumber="331" pageId="5" pageNumber="326" type="description">
<paragraph id="7E68F934CA72F33610F5ED64D494FE9C" blockId="5.[151,1437,151,1409]" pageId="5" pageNumber="326">
<emphasis id="4CA32526CA72F33610F5ED64D777FEC1" bold="true" box="[199,345,260,285]" pageId="5" pageNumber="326">Description.</emphasis>
Female. Body length 730 (720) including gnathosoma, width 413 (410), gnathosoma 160 (155),
<date id="0A69DFF4CA72F33610A5ED49D685FE98" box="[151,171,297,324]" pageId="5" pageNumber="326">
<emphasis id="4CA32526CA72F33610A5ED49D68DFE9C" box="[151,163,297,320]" italics="true" pageId="5" pageNumber="326">v</emphasis>
<subScript id="E253FB71CA72F3361091ED56D685FE98" attach="left" box="[163,171,310,324]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
</date>
-
<emphasis id="4CA32526CA72F3361086ED47D6ECFE9C" box="[180,194,295,320]" italics="true" pageId="5" pageNumber="326">h</emphasis>
<subScript id="E253FB71CA72F33610F0ED56D6E4FE98" attach="left" box="[194,202,310,324]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
535 (565),
<emphasis id="4CA32526CA72F336117EED49D74DFE9C" box="[332,355,297,320]" italics="true" pageId="5" pageNumber="326">sc</emphasis>
<subScript id="E253FB71CA72F3361156ED56D75FFE98" attach="right" box="[356,369,310,324]" fontSize="6" pageId="5" pageNumber="326">2-</subScript>
<emphasis id="4CA32526CA72F3361143ED49D7A6FE9C" box="[369,392,297,320]" italics="true" pageId="5" pageNumber="326">sc</emphasis>
<subScript id="E253FB71CA72F33611BAED56D7BEFE98" attach="left" box="[392,400,310,324]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
310 (290),
<emphasis id="4CA32526CA72F336123DED49D435FE9C" box="[527,539,297,320]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F336122EED56D40AFE98" attach="right" box="[540,548,310,324]" fontSize="6" pageId="5" pageNumber="326">3</subScript>
-
<emphasis id="4CA32526CA72F336121EED49D416FE9C" box="[556,568,297,320]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F336120BED56D46FFE98" attach="left" box="[569,577,310,324]" fontSize="6" pageId="5" pageNumber="326">3</subScript>
335 (340).
</paragraph>
<paragraph id="7E68F934CA72F33610F5ED30D796FDC1" blockId="5.[151,1437,151,1409]" pageId="5" pageNumber="326">
<emphasis id="4CA32526CA72F33610F5ED30D70FFEBB" box="[199,289,336,359]" italics="true" pageId="5" pageNumber="326">Dorsum</emphasis>
. Medial propodosomal plate area with distinct regular polygonal reticulate pattern resembling geometric alveoli filled with fine irregularly punctate striae (
<figureCitation id="E6ECE5B1CA72F3361366ED12D5B7FE57" box="[852,921,370,395]" captionStart="FIGURE 1" captionStartId="7.[151,250,1971,1993]" captionTargetBox="[179,1415,195,1955]" captionTargetId="figure@7.[176,1415,195,1955]" captionTargetPageId="7" captionText="FIGURE 1. Georgiobia skuthiae n. sp., female. A. dorsal aspect, B. detail of medial propodosomal striation, C. dorsal body seta (e 2). Scale bars = 100 µm (A), 10 µm (B), 25 µm (C)." httpUri="https://zenodo.org/record/2627010/files/figure.png" pageId="5" pageNumber="326">Fig. 1</figureCitation>
AB). Lateral striation longitudinal. Rest of idiosoma with medial striation transverse. Striation longitudinal to wrinkled-longitudinal between first three pairs of dorsocentral and lateral setae, areas with more obvious wrinkled striation between dorsocentrals and dorsolaterals of each of the first three rows. Transverse striation between setae
<emphasis id="4CA32526CA72F3361404ED80D26CFE2B" box="[1078,1090,480,503]" italics="true" pageId="5" pageNumber="326">e</emphasis>
<subScript id="E253FB71CA72F3361470ED8DD264FE27" attach="left" box="[1090,1098,493,507]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
and
<emphasis id="4CA32526CA72F336144CEDBFD2A8FE2B" box="[1150,1158,479,503]" italics="true" pageId="5" pageNumber="326">f</emphasis>
<subScript id="E253FB71CA72F33614B4ED8DD2A0FE27" attach="left" box="[1158,1166,493,507]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
, irregular and punctate caudally beyond setae
<emphasis id="4CA32526CA72F33611A7EE66D7B3FDC2" box="[405,413,518,542]" italics="true" pageId="5" pageNumber="326">f</emphasis>
<subScript id="E253FB71CA72F33611AFEE74D79FFDFE" attach="left" box="[413,433,532,546]" fontSize="6" pageId="5" pageNumber="326">1-2</subScript>
.
</paragraph>
<paragraph id="7E68F934CA72F33610F5EE4CD7C7FD65" blockId="5.[151,1437,151,1409]" pageId="5" pageNumber="326">
Prodorsum with three pairs of setae, hysterosoma with ten pairs. Humeral setae (
<emphasis id="4CA32526CA72F3361447EE4DD2AFFD98" box="[1141,1153,557,580]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F33614B3EE5BD2A7FD95" attach="left" box="[1153,1161,571,585]" fontSize="6" pageId="5" pageNumber="326">3</subScript>
) not in lateral position, moved behind first dorsolateral setae
<emphasis id="4CA32526CA72F3361271EE34D461FDB7" box="[579,591,596,619]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F336127DEE01D479FDB3" attach="left" box="[591,599,609,623]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
. Dorsal setae relatively long, linear lanceolate, serrate, inserted on strong tubercles. All dorsal tubercles punctate, finely granulated, cone shaped (
<figureCitation id="E6ECE5B1CA72F33613F4EE1AD209FD4E" box="[966,1063,634,659]" captionStart="FIGURE 1" captionStartId="7.[151,250,1971,1993]" captionTargetBox="[179,1415,195,1955]" captionTargetId="figure@7.[176,1415,195,1955]" captionTargetPageId="7" captionText="FIGURE 1. Georgiobia skuthiae n. sp., female. A. dorsal aspect, B. detail of medial propodosomal striation, C. dorsal body seta (e 2). Scale bars = 100 µm (A), 10 µm (B), 25 µm (C)." httpUri="https://zenodo.org/record/2627010/files/figure.png" pageId="5" pageNumber="326">Fig. 1 C</figureCitation>
); tubercles of members of
<emphasis id="4CA32526CA72F3361564EE1AD34AFD4F" box="[1366,1380,634,659]" italics="true" pageId="5" pageNumber="326">d</emphasis>
<subScript id="E253FB71CA72F3361556EEE8D342FD4A" attach="left" box="[1380,1388,648,662]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
and
<emphasis id="4CA32526CA72F33610A5EEC2D68DFD65" box="[151,163,674,697]" italics="true" pageId="5" pageNumber="326">e</emphasis>
<subScript id="E253FB71CA72F3361091EECFD685FD61" attach="left" box="[163,171,687,701]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
setae obviously contiguous.
</paragraph>
<paragraph id="7E68F934CA72F33610F5EEA7D6C2FC7D" blockId="5.[151,1437,151,1409]" pageId="5" pageNumber="326">
First, second and third pairs of dorsocentral hysterosomal setae surpass insertions of consecutive setae, setae
<emphasis id="4CA32526CA72F33615BAEEA8D3BAFD03" box="[1416,1428,712,735]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F33615A6EEB6D3B2FD38" attach="left" box="[1428,1436,726,740]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
and
<emphasis id="4CA32526CA72F33610F5EE8ED6FBFCDB" box="[199,213,750,775]" italics="true" pageId="5" pageNumber="326">d</emphasis>
<subScript id="E253FB71CA72F33610E7EE9CD6F3FCD6" attach="left" box="[213,221,764,778]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
the longest; setae
<date id="0A69DFF4CA72F336119CEE8FD7EDFCD6" box="[430,451,751,778]" pageId="5" pageNumber="326">
<emphasis id="4CA32526CA72F336119CEE8FD794FCDA" box="[430,442,751,774]" italics="true" pageId="5" pageNumber="326">v</emphasis>
<subScript id="E253FB71CA72F3361189EE9CD7EDFCD6" attach="right" box="[443,451,764,778]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
</date>
,
<emphasis id="4CA32526CA72F33611E3EE8FD7F3FCDA" box="[465,477,751,774]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F33611ECEE9CD7C8FCD6" attach="right" box="[478,486,764,778]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
,
<emphasis id="4CA32526CA72F33611C6EE8FD42EFCDA" box="[500,512,751,774]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F3361233EE9CD427FCD6" attach="left" box="[513,521,764,778]" fontSize="6" pageId="5" pageNumber="326">3</subScript>
and
<emphasis id="4CA32526CA72F3361272EE8ED460FCDB" box="[576,590,750,775]" italics="true" pageId="5" pageNumber="326">h</emphasis>
<subScript id="E253FB71CA72F336127CEE9CD478FCD6" attach="left" box="[590,598,764,778]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
similar in length, the shortest. Setae lengths:
<date id="0A69DFF4CA72F3361468EE8FD2BCFCDB" box="[1114,1170,751,778]" pageId="5" pageNumber="326" value="1992-05-02">
<emphasis id="4CA32526CA72F3361468EE8FD248FCDA" box="[1114,1126,751,774]" italics="true" pageId="5" pageNumber="326">v</emphasis>
<subScript id="E253FB71CA72F3361455EE9CD241FCD6" attach="left" box="[1127,1135,764,778]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
92
</date>
(90);
<emphasis id="4CA32526CA72F33614EAEE8FD2C1FCDA" box="[1240,1263,751,774]" italics="true" pageId="5" pageNumber="326">sc</emphasis>
<subScript id="E253FB71CA72F33614DDEE9CD2D9FCD6" attach="left" box="[1263,1271,764,778]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
123 (121);
<emphasis id="4CA32526CA72F336154FEE8FD3BAFCDA" box="[1405,1428,751,774]" italics="true" pageId="5" pageNumber="326">sc</emphasis>
<subScript id="E253FB71CA72F33615A6EE9CD3B2FCD6" attach="left" box="[1428,1436,764,778]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
105 (117);
<emphasis id="4CA32526CA72F3361120EF76D730FCF1" box="[274,286,790,813]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F336112CEF43D708FCED" attach="left" box="[286,294,803,817]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
168 (135);
<emphasis id="4CA32526CA72F336119AEF76D79AFCF1" box="[424,436,790,813]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F3361186EF43D792FCED" attach="left" box="[436,444,803,817]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
90 (98);
<emphasis id="4CA32526CA72F3361211EF76D401FCF1" box="[547,559,790,813]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F336121DEF43D419FCED" attach="left" box="[559,567,803,817]" fontSize="6" pageId="5" pageNumber="326">3</subScript>
76 (97);
<emphasis id="4CA32526CA72F33612AFEF75D485FCF2" box="[669,683,789,814]" italics="true" pageId="5" pageNumber="326">d</emphasis>
<subScript id="E253FB71CA72F336129EEF43D49AFCED" attach="left" box="[684,692,803,817]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
160 (150);
<emphasis id="4CA32526CA72F3361304EF75D56AFCF2" box="[822,836,789,814]" italics="true" pageId="5" pageNumber="326">d</emphasis>
<subScript id="E253FB71CA72F3361376EF43D562FCED" attach="left" box="[836,844,803,817]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
130 (122);
<emphasis id="4CA32526CA72F33613FCEF76D5F4FCF1" box="[974,986,790,813]" italics="true" pageId="5" pageNumber="326">e</emphasis>
<subScript id="E253FB71CA72F33613E9EF43D5CDFCED" attach="left" box="[987,995,803,817]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
150 (145);
<emphasis id="4CA32526CA72F3361457EF76D25FFCF1" box="[1125,1137,790,813]" italics="true" pageId="5" pageNumber="326">e</emphasis>
<subScript id="E253FB71CA72F3361443EF43D257FCED" attach="left" box="[1137,1145,803,817]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
130 (130);
<emphasis id="4CA32526CA72F33614CEEF76D32AFCF2" box="[1276,1284,790,814]" italics="true" pageId="5" pageNumber="326">f</emphasis>
<subScript id="E253FB71CA72F3361531EF43D325FCED" attach="left" box="[1283,1291,803,817]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
125 (113);
<emphasis id="4CA32526CA72F33615BEEF76D3BAFCF2" box="[1420,1428,790,814]" italics="true" pageId="5" pageNumber="326">f</emphasis>
<subScript id="E253FB71CA72F33615A6EF43D3B2FCED" attach="left" box="[1428,1436,803,817]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
100 (107);
<emphasis id="4CA32526CA72F3361123EF5CD731FC89" box="[273,287,828,853]" italics="true" pageId="5" pageNumber="326">h</emphasis>
<subScript id="E253FB71CA72F336112DEF2AD709FC84" attach="left" box="[287,295,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
80 (98). Distances between setae:
<emphasis id="4CA32526CA72F3361282EF5DD492FC88" box="[688,700,829,852]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F336128EEF2AD4EAFC84" attach="left" box="[700,708,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
-
<emphasis id="4CA32526CA72F33612FFEF5DD4F7FC88" box="[717,729,829,852]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F33612E8EF2AD4CCFC84" attach="left" box="[730,738,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
55 (50),
<emphasis id="4CA32526CA72F3361378EF5CD576FC89" box="[842,856,828,853]" italics="true" pageId="5" pageNumber="326">d</emphasis>
<subScript id="E253FB71CA72F336136AEF2AD54EFC84" attach="left" box="[856,864,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
-
<emphasis id="4CA32526CA72F336135BEF5CD559FC89" box="[873,887,828,853]" italics="true" pageId="5" pageNumber="326">d</emphasis>
<subScript id="E253FB71CA72F3361345EF2AD551FC84" attach="left" box="[887,895,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
43 (38),
<emphasis id="4CA32526CA72F33613D5EF5DD5DDFC88" box="[999,1011,829,852]" italics="true" pageId="5" pageNumber="326">e</emphasis>
<subScript id="E253FB71CA72F33613C1EF2AD5D5FC84" attach="left" box="[1011,1019,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
-
<emphasis id="4CA32526CA72F3361436EF5DD23EFC88" box="[1028,1040,829,852]" italics="true" pageId="5" pageNumber="326">e</emphasis>
<subScript id="E253FB71CA72F3361423EF2AD237FC84" attach="left" box="[1041,1049,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
48 (45),
<emphasis id="4CA32526CA72F33614B3EF5DD2A3FC88" box="[1153,1165,829,852]" italics="true" pageId="5" pageNumber="326">c</emphasis>
<subScript id="E253FB71CA72F33614BFEF2AD2BBFC84" attach="left" box="[1165,1173,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
-
<emphasis id="4CA32526CA72F33614ACEF5CD282FC89" box="[1182,1196,828,853]" italics="true" pageId="5" pageNumber="326">d</emphasis>
<subScript id="E253FB71CA72F336149EEF2AD29AFC84" attach="left" box="[1196,1204,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
115 (120),
<emphasis id="4CA32526CA72F3361505EF5CD36BFC89" box="[1335,1349,828,853]" italics="true" pageId="5" pageNumber="326">d</emphasis>
<subScript id="E253FB71CA72F3361577EF2AD363FC84" attach="left" box="[1349,1357,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
-
<emphasis id="4CA32526CA72F3361564EF5DD34CFC88" box="[1366,1378,829,852]" italics="true" pageId="5" pageNumber="326">e</emphasis>
<subScript id="E253FB71CA72F3361551EF2AD345FC84" attach="left" box="[1379,1387,842,856]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
105 (140),
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<subScript id="E253FB71CA72F33610DCEF11D6D8FCA3" attach="left" box="[238,246,881,895]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
-
<emphasis id="4CA32526CA72F33610CDEF03D729FCA7" box="[255,263,867,891]" italics="true" pageId="5" pageNumber="326">f</emphasis>
<subScript id="E253FB71CA72F3361135EF11D721FCA3" attach="left" box="[263,271,881,895]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
. 100 (105). Sacral setae
<emphasis id="4CA32526CA72F3361215EF03D401FCA7" box="[551,559,867,891]" italics="true" pageId="5" pageNumber="326">f</emphasis>
<quantity id="B92F54D1CA72F336121DEF11D47AFCA7" box="[559,596,866,895]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="326" unit="in" value="1.0">
<subScript id="E253FB71CA72F336121DEF11D419FCA3" attach="left" box="[559,567,881,895]" fontSize="6" pageId="5" pageNumber="326">1</subScript>
in
</quantity>
normal position, in line with other dorsocentral setae, not contiguous with
<emphasis id="4CA32526CA72F33610A5EFEAD6B1FC7E" box="[151,159,906,930]" italics="true" pageId="5" pageNumber="326">f</emphasis>
<subScript id="E253FB71CA72F33610ACEFF8D688FC7A" attach="left" box="[158,166,920,934]" fontSize="6" pageId="5" pageNumber="326">2</subScript>
setae.
</paragraph>
<paragraph id="7E68F934CA72F33610F5EFD0D5F8FBE9" blockId="5.[151,1437,151,1409]" pageId="5" pageNumber="326">
<emphasis id="4CA32526CA72F33610F5EFD0D778FC15" box="[199,342,944,969]" italics="true" pageId="5" pageNumber="326">Gnathosoma</emphasis>
. Stylophore rounded, slightly emarginate anteriorly, longer than wide 130 (125) long 80 (90) wide. Tibial claw of palpus slightly surpasses tip of palptarsus (
<figureCitation id="E6ECE5B1CA72F3361350EFB4D5EBFC31" box="[866,965,980,1005]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[190,1400,196,1958]" captionTargetId="figure@9.[190,1400,193,1958]" captionTargetPageId="9" captionText="FIGURE 3. Georgiobia skuthiae n. sp., female. A. tarsus and tibia II, B. tarsus and tibia III, C. tarsus and tibia IV, D. palptarsus and palptibia, E. peritremal enlargement, F. ventral striation. Scale bars = 25 µm (AC), 10 µm (DE), 50 µm (F)." httpUri="https://zenodo.org/record/2627014/files/figure.png" pageId="5" pageNumber="326">Fig. 3 D</figureCitation>
). Palptarsus quite short with six setae and one solenidion. Eupathidia
<emphasis id="4CA32526CA72F33611FAEF98D7C9FBCD" box="[456,487,1016,1041]" italics="true" pageId="5" pageNumber="326">ul</emphasis>
ζ,
<emphasis id="4CA32526CA72F3361233EF98D408FBCD" box="[513,550,1016,1041]" italics="true" pageId="5" pageNumber="326">ul</emphasis>
ζ and solenidion subequal in length,
<emphasis id="4CA32526CA72F336138FEF99D5F8FBCC" box="[957,982,1017,1040]" italics="true" pageId="5" pageNumber="326">su</emphasis>
ζ longer about 6. Peritreme anastomosed distally in a suboval enlargement, length 32 (30), width 15 (13) (
<figureCitation id="E6ECE5B1CA72F3361359E87CD5E6FBE9" box="[875,968,1052,1077]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[190,1400,196,1958]" captionTargetId="figure@9.[190,1400,193,1958]" captionTargetPageId="9" captionText="FIGURE 3. Georgiobia skuthiae n. sp., female. A. tarsus and tibia II, B. tarsus and tibia III, C. tarsus and tibia IV, D. palptarsus and palptibia, E. peritremal enlargement, F. ventral striation. Scale bars = 25 µm (AC), 10 µm (DE), 50 µm (F)." httpUri="https://zenodo.org/record/2627014/files/figure.png" pageId="5" pageNumber="326">Fig. 3 E</figureCitation>
).
</paragraph>
<paragraph id="7E68F934CA72F33610F5E824D271FB16" blockId="5.[151,1437,151,1409]" pageId="5" pageNumber="326">
<emphasis id="4CA32526CA72F33610F5E824D722FB87" box="[199,268,1092,1115]" italics="true" pageId="5" pageNumber="326">Venter</emphasis>
. Striation transverse between 1
<superScript id="89A2547CCA72F336125EE85FD458FB91" attach="left" box="[620,630,1087,1101]" fontSize="6" pageId="5" pageNumber="326">st</superScript>
(
<emphasis id="4CA32526CA72F33612B4E823D48CFB87" box="[646,674,1091,1115]" italics="true" pageId="5" pageNumber="326">1a</emphasis>
) and 2
<superScript id="89A2547CCA72F33612DDE85FD4D1FB91" attach="left" box="[751,767,1087,1101]" fontSize="6" pageId="5" pageNumber="326">nd</superScript>
(
<emphasis id="4CA32526CA72F336133DE823D505FB87" box="[783,811,1091,1115]" italics="true" pageId="5" pageNumber="326">3a</emphasis>
) pairs of setae, with irregular pattern medially, obliquelongitudinal laterally between 2
<superScript id="89A2547CCA72F33611C8E806D424FBA8" attach="left" box="[506,522,1126,1140]" fontSize="6" pageId="5" pageNumber="326">nd</superScript>
and 3
<superScript id="89A2547CCA72F336127CE806D475FBA8" attach="left" box="[590,603,1126,1140]" fontSize="6" pageId="5" pageNumber="326">rd</superScript>
(
<emphasis id="4CA32526CA72F3361259E80AD4A6FB5E" box="[619,648,1130,1154]" italics="true" pageId="5" pageNumber="326">4a</emphasis>
) pairs of setae, transverse between members of
<emphasis id="4CA32526CA72F3361490E80AD290FB5E" box="[1186,1214,1130,1154]" italics="true" pageId="5" pageNumber="326">4a</emphasis>
, transverse anterior to aggenital (
<emphasis id="4CA32526CA72F3361102E8EFD762FB7A" box="[304,332,1167,1190]" italics="true" pageId="5" pageNumber="326">ag</emphasis>
) pair of setae, then longitudinal between
<emphasis id="4CA32526CA72F3361300E8EFD560FB7A" box="[818,846,1167,1190]" italics="true" pageId="5" pageNumber="326">ag</emphasis>
setae (
<figureCitation id="E6ECE5B1CA72F3361390E8EDD5D4FB7A" box="[930,1018,1165,1190]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[190,1400,196,1958]" captionTargetId="figure@9.[190,1400,193,1958]" captionTargetPageId="9" captionText="FIGURE 3. Georgiobia skuthiae n. sp., female. A. tarsus and tibia II, B. tarsus and tibia III, C. tarsus and tibia IV, D. palptarsus and palptibia, E. peritremal enlargement, F. ventral striation. Scale bars = 25 µm (AC), 10 µm (DE), 50 µm (F)." httpUri="https://zenodo.org/record/2627014/files/figure.png" pageId="5" pageNumber="326">Fig. 3F</figureCitation>
). Sacculus of spermatheca not seen. Three pairs of pseudanal setae (
<emphasis id="4CA32526CA72F33611C8E8D3D43DFB16" box="[506,531,1203,1226]" italics="true" pageId="5" pageNumber="326">ps</emphasis>
<subScript id="E253FB71CA72F3361221E8A0D406FB12" attach="left" box="[531,552,1216,1230]" fontSize="6" pageId="5" pageNumber="326">1-3</subScript>
) and two pairs of ventrocaudal (
<emphasis id="4CA32526CA72F33613A7E8D2D58DFB17" box="[917,931,1202,1227]" italics="true" pageId="5" pageNumber="326">h</emphasis>
<subScript id="E253FB71CA72F3361391E8A0D599FB12" attach="left" box="[931,951,1216,1230]" fontSize="6" pageId="5" pageNumber="326">2-3</subScript>
) setae present.
</paragraph>
<paragraph id="7E68F934CA72F33610F5E8BAD7C8FA5D" blockId="5.[151,1437,151,1409]" pageId="5" pageNumber="326">
<emphasis id="4CA32526CA72F33610F5E8BAD6D2FB2D" box="[199,252,1242,1265]" italics="true" pageId="5" pageNumber="326">Legs</emphasis>
. Leg I about twice as long as body length, legs II to IV less than half the body length. Leg
<date id="0A69DFF4CA72F3361533E8B9D364FB2D" box="[1281,1354,1240,1265]" pageId="5" pageNumber="326" value="1615-01">I 1615</date>
(1515) long (measured from femur to tarsus), leg II 310 (315), leg III 275 (275), leg IV 335 (345). Length of segments of leg I as follows: trochanter 30 (24), femur 700 (680), genu 55 (55), tibia 640 (575), tarsus 220 (200). Tibia and tarsus I bow-shaped, the latter almost retrorse distally, both with ribbed aspect (
<figureCitation id="E6ECE5B1CA72F3361415E924D245FA81" box="[1063,1131,1348,1373]" captionStart="FIGURE 2" captionStartId="8.[151,250,1971,1993]" captionTargetBox="[279,1436,238,1924]" captionTargetId="figure@8.[151,1437,238,1924]" captionTargetPageId="8" captionText="FIGURE 2. Georgiobia skuthiae n. sp., female. A. tarsus I, B. tibia I, C. empodium and claws I, D. empodium and claws II, E. coxisternal setae 1b, 1c (from left to right). Scale bars = 25 µm (AB), 10 µm (CE)." httpUri="https://zenodo.org/record/2627012/files/figure.png" pageId="5" pageNumber="326">Fig. 2</figureCitation>
AB). Leg setal counts as follows (
<figureCitation id="E6ECE5B1CA72F33610C8E908D769FA5D" box="[250,327,1384,1409]" captionStart="FIGURE 2" captionStartId="8.[151,250,1971,1993]" captionTargetBox="[279,1436,238,1924]" captionTargetId="figure@8.[151,1437,238,1924]" captionTargetPageId="8" captionText="FIGURE 2. Georgiobia skuthiae n. sp., female. A. tarsus I, B. tibia I, C. empodium and claws I, D. empodium and claws II, E. coxisternal setae 1b, 1c (from left to right). Scale bars = 25 µm (AB), 10 µm (CE)." httpUri="https://zenodo.org/record/2627012/files/figure.png" pageId="5" pageNumber="326">Figs. 2</figureCitation>
AB, 3 AC):
</paragraph>
<paragraph id="7E68F934CA72F33610A5E9D1D4CCFA15" blockId="5.[151,738,1457,1589]" box="[151,738,1457,1481]" pageId="5" pageNumber="326">
<date id="0A69DFF4CA72F33610A5E9D1D6FBFA15" box="[151,213,1457,1481]" pageId="5" pageNumber="326">I 2</date>
¯ 1 ¯ 9 ¯ 2 13 + (1) 11 + (1) + 2 duplexes;
</paragraph>
<paragraph id="7E68F934CA72F33610A5E9B5D416FA31" blockId="5.[151,738,1457,1589]" box="[151,568,1492,1517]" pageId="5" pageNumber="326">
<date id="0A69DFF4CA72F33610A5E9B5D6FBFA31" box="[151,213,1493,1517]" pageId="5" pageNumber="326">II 2</date>
¯ 1 6 ¯ 2 7 ¯ 11 + 1 duplex;
</paragraph>
<paragraph id="7E68F934CA72F33610A5E999D7DCF9CD" blockId="5.[151,738,1457,1589]" box="[151,498,1529,1553]" pageId="5" pageNumber="326">
<date id="0A69DFF4CA72F33610A5E999D6FBF9CD" box="[151,213,1529,1553]" pageId="5" pageNumber="326">III 1</date>
¯ 1 2 3 ¯ 5 ¯ 8 + (1);
</paragraph>
<paragraph id="7E68F934CA72F33610A5EA7DD7DEF9E9" blockId="5.[151,738,1457,1589]" box="[151,496,1564,1589]" pageId="5" pageNumber="326">
<date id="0A69DFF4CA72F33610A5EA7DD6FBF9E9" box="[151,213,1565,1589]" pageId="5" pageNumber="326">IV 1</date>
¯ 1 2 3 ¯ 5 8 + (1).
</paragraph>
<paragraph id="7E68F934CA72F33610F5EA05D30BF97D" blockId="5.[151,1437,1636,2022]" pageId="5" pageNumber="326">
True claws and empodia pad-like, the latter with two rows of tenent hairs. Leg I empodial pad and pad-like true claws similar in length (
<figureCitation id="E6ECE5B1CA72F3361197EAE8D42AF97D" box="[421,516,1672,1697]" captionStart="FIGURE 2" captionStartId="8.[151,250,1971,1993]" captionTargetBox="[279,1436,238,1924]" captionTargetId="figure@8.[151,1437,238,1924]" captionTargetPageId="8" captionText="FIGURE 2. Georgiobia skuthiae n. sp., female. A. tarsus I, B. tibia I, C. empodium and claws I, D. empodium and claws II, E. coxisternal setae 1b, 1c (from left to right). Scale bars = 25 µm (AB), 10 µm (CE)." httpUri="https://zenodo.org/record/2627012/files/figure.png" pageId="5" pageNumber="326">Fig. 2 C</figureCitation>
); empodial pads IIIV about twice as long as true claw pads (
<figureCitation id="E6ECE5B1CA72F3361485EAE8D336F97D" box="[1207,1304,1672,1697]" captionStart="FIGURE 2" captionStartId="8.[151,250,1971,1993]" captionTargetBox="[279,1436,238,1924]" captionTargetId="figure@8.[151,1437,238,1924]" captionTargetPageId="8" captionText="FIGURE 2. Georgiobia skuthiae n. sp., female. A. tarsus I, B. tibia I, C. empodium and claws I, D. empodium and claws II, E. coxisternal setae 1b, 1c (from left to right). Scale bars = 25 µm (AB), 10 µm (CE)." httpUri="https://zenodo.org/record/2627012/files/figure.png" pageId="5" pageNumber="326">Fig. 2 D</figureCitation>
).
</paragraph>
<paragraph id="7E68F934CA72F33610F5EACCD52DF81D" blockId="5.[151,1437,1636,2022]" pageId="5" pageNumber="326">
Tarsi without tectal setae (
<emphasis id="4CA32526CA72F33611DEEACED42EF919" box="[492,512,1710,1733]" italics="true" pageId="5" pageNumber="326">tc</emphasis>
-
<emphasis id="4CA32526CA72F3361220EACED408F919" box="[530,550,1710,1733]" italics="true" pageId="5" pageNumber="326">tc</emphasis>
), first primiventral setae (
<emphasis id="4CA32526CA72F3361368EACED55AF919" box="[858,884,1710,1733]" italics="true" pageId="5" pageNumber="326">pv</emphasis>
) absent on tarsus I, present on other tarsi. Tarsus I with two distal duplex setae inserted near each other. Solenidia members of duplex setae subequal, 92 and 91, tactile members also subequal, 6 and 5. On leg II, duplex solenidion 42 (44), tactile setae 8 (8). On tarsi III and IV solenidia 17 (18) and 11, respectively. Long, strong, stout, serrate seta present on tibia III and IV, far less developed on tibia II (
<figureCitation id="E6ECE5B1CA72F3361120EB5CD763F889" box="[274,333,1852,1877]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[190,1400,196,1958]" captionTargetId="figure@9.[190,1400,193,1958]" captionTargetPageId="9" captionText="FIGURE 3. Georgiobia skuthiae n. sp., female. A. tarsus and tibia II, B. tarsus and tibia III, C. tarsus and tibia IV, D. palptarsus and palptibia, E. peritremal enlargement, F. ventral striation. Scale bars = 25 µm (AC), 10 µm (DE), 50 µm (F)." httpUri="https://zenodo.org/record/2627014/files/figure.png" pageId="5" pageNumber="326">Fig 3</figureCitation>
AC). On tibia III, macroseta and leg segment subequal in length, 82 (80) and 82(79), respectively (
<figureCitation id="E6ECE5B1CA72F33610ADEB00D6D0F8A5" box="[159,254,1888,1913]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[190,1400,196,1958]" captionTargetId="figure@9.[190,1400,193,1958]" captionTargetPageId="9" captionText="FIGURE 3. Georgiobia skuthiae n. sp., female. A. tarsus and tibia II, B. tarsus and tibia III, C. tarsus and tibia IV, D. palptarsus and palptibia, E. peritremal enlargement, F. ventral striation. Scale bars = 25 µm (AC), 10 µm (DE), 50 µm (F)." httpUri="https://zenodo.org/record/2627014/files/figure.png" pageId="5" pageNumber="326">Fig. 3 B</figureCitation>
); on tibia IV, macroseta slightly longer than respected segment, 101 (106) and 99 (95), respectively (
<figureCitation id="E6ECE5B1CA72F336155DEB00D6ECF841" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[190,1400,196,1958]" captionTargetId="figure@9.[190,1400,193,1958]" captionTargetPageId="9" captionText="FIGURE 3. Georgiobia skuthiae n. sp., female. A. tarsus and tibia II, B. tarsus and tibia III, C. tarsus and tibia IV, D. palptarsus and palptibia, E. peritremal enlargement, F. ventral striation. Scale bars = 25 µm (AC), 10 µm (DE), 50 µm (F)." httpUri="https://zenodo.org/record/2627014/files/figure.png" pageId="5" pageNumber="326">Fig. 3 C</figureCitation>
). Tarsi III and IV with solenidion not associated with tactile (
<emphasis id="4CA32526CA72F336134BEBE5D5A6F841" box="[889,904,1925,1949]" italics="true" pageId="5" pageNumber="326">ft</emphasis>
); solenidion III, 17 (18) about ¼ the length of tactiles, solenidion
<date id="0A69DFF4CA72F3361142EBC9D781F81D" box="[368,431,1961,1985]" pageId="5" pageNumber="326" value="2011-04">IV 11</date>
about ½ the length of tactiles.
</paragraph>
<paragraph id="7E68F934CA72F33610F5EBADD3BDF839" blockId="5.[151,1437,1636,2022]" box="[199,1427,1997,2022]" pageId="5" pageNumber="326">
Proximal coxisternal seta
<emphasis id="4CA32526CA72F33611DBEBADD42AF83A" box="[489,516,1997,2022]" italics="true" pageId="5" pageNumber="326">1b</emphasis>
slender 56 (50), distal coxisternal seta
<emphasis id="4CA32526CA72F3361389EBADD5F8F839" box="[955,982,1997,2021]" italics="true" pageId="5" pageNumber="326">1c</emphasis>
shorter 19 (21), serrate, stout (
<figureCitation id="E6ECE5B1CA72F336151DEBADD3A9F839" box="[1327,1415,1997,2022]" captionStart="FIGURE 2" captionStartId="8.[151,250,1971,1993]" captionTargetBox="[279,1436,238,1924]" captionTargetId="figure@8.[151,1437,238,1924]" captionTargetPageId="8" captionText="FIGURE 2. Georgiobia skuthiae n. sp., female. A. tarsus I, B. tibia I, C. empodium and claws I, D. empodium and claws II, E. coxisternal setae 1b, 1c (from left to right). Scale bars = 25 µm (AB), 10 µm (CE)." httpUri="https://zenodo.org/record/2627012/files/figure.png" pageId="5" pageNumber="326">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="7E68F934CA71F33510F5ECF7D777FF0B" blockId="6.[151,1436,151,856]" pageId="6" pageNumber="327">
Deutonymph (n=1), (
<figureCitation id="E6ECE5B1CA71F3351186ECF7D7D7FF6C" box="[436,505,151,176]" captionStart="FIGURE 4" captionStartId="10.[151,250,1101,1123]" captionTargetBox="[150,1436,214,1066]" captionTargetId="figure@10.[150,1436,214,1066]" captionTargetPageId="10" captionText="FIGURE 4. Georgiobia skuthiae n. sp., Deutonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation, D. peritreme. Scale bars: 100 µm (A), 25 µm (BD)." httpUri="https://zenodo.org/record/2627016/files/figure.png" pageId="6" pageNumber="327">Fig. 4</figureCitation>
). Body length 500 including gnathosoma, width 290, gnathosoma
<date id="0A69DFF4CA71F33514EFECF8D335FF68" box="[1245,1307,152,180]" pageId="6" pageNumber="327" value="1980-05-02">
80,
<emphasis id="4CA32526CA71F3351535ECF9D33DFF6C" box="[1287,1299,153,176]" italics="true" pageId="6" pageNumber="327">v</emphasis>
<subScript id="E253FB71CA71F3351521ECC6D335FF68" attach="left" box="[1299,1307,166,180]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
</date>
-
<emphasis id="4CA32526CA71F3351516ECF7D31CFF6C" box="[1316,1330,151,176]" italics="true" pageId="6" pageNumber="327">h</emphasis>
<subScript id="E253FB71CA71F3351500ECC6D314FF68" attach="left" box="[1330,1338,166,180]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
400,
<emphasis id="4CA32526CA71F335154AECF9D3A1FF6C" box="[1400,1423,153,176]" italics="true" pageId="6" pageNumber="327">sc</emphasis>
<subScript id="E253FB71CA71F33515BDECC6D3B2FF68" attach="left" box="[1423,1436,166,180]" fontSize="6" pageId="6" pageNumber="327">2-</subScript>
<emphasis id="4CA32526CA71F33510A5ECA0D680FF0B" box="[151,174,192,215]" italics="true" pageId="6" pageNumber="327">sc</emphasis>
<subScript id="E253FB71CA71F335109CECADD698FF07" attach="left" box="[174,182,205,219]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
215,
<emphasis id="4CA32526CA71F33510C0ECA0D6D0FF0B" box="[242,254,192,215]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33510CCECADD728FF07" attach="left" box="[254,262,205,219]" fontSize="6" pageId="6" pageNumber="327">3</subScript>
-
<emphasis id="4CA32526CA71F335113DECA0D735FF0B" box="[271,283,192,215]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F335112EECADD70AFF07" attach="left" box="[284,292,205,219]" fontSize="6" pageId="6" pageNumber="327">3</subScript>
232.
</paragraph>
<paragraph id="7E68F934CA71F33510F5EC86D738FDA6" blockId="6.[151,1436,151,856]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F33510F5EC86D70FFF21" box="[199,289,230,253]" italics="true" pageId="6" pageNumber="327">Dorsum</emphasis>
. Medial prodorsal striation resembling that of female, with geometric alveoli filled with fine irregular striae becoming punctate close to setae
<date id="0A69DFF4CA71F335126FED6AD45FFEFA" box="[605,625,266,294]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F335126FED6AD447FEFD" box="[605,617,266,289]" italics="true" pageId="6" pageNumber="327">v</emphasis>
<subScript id="E253FB71CA71F335125BED78D45FFEFA" attach="left" box="[617,625,280,294]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
</date>
, lateral striation longitudinal (
<figureCitation id="E6ECE5B1CA71F33513FFED69D201FEFE" box="[973,1071,265,290]" captionStart="FIGURE 4" captionStartId="10.[151,250,1101,1123]" captionTargetBox="[150,1436,214,1066]" captionTargetId="figure@10.[150,1436,214,1066]" captionTargetPageId="10" captionText="FIGURE 4. Georgiobia skuthiae n. sp., Deutonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation, D. peritreme. Scale bars: 100 µm (A), 25 µm (BD)." httpUri="https://zenodo.org/record/2627016/files/figure.png" pageId="6" pageNumber="327">Fig. 4 B</figureCitation>
). Overall hysterosomal striation pattern transverse between members of setae
<emphasis id="4CA32526CA71F3351287ED51D4EFFE94" box="[693,705,305,328]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33512F3ED5ED4E7FE90" attach="left" box="[705,713,318,332]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
to
<emphasis id="4CA32526CA71F33512CAED51D52EFE95" box="[760,768,305,329]" italics="true" pageId="6" pageNumber="327">f</emphasis>
<subScript id="E253FB71CA71F3351332ED5ED526FE90" attach="left" box="[768,776,318,332]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
, longitudinal laterally. Humeral setae (
<emphasis id="4CA32526CA71F33514EBED51D2CBFE94" box="[1241,1253,305,328]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33514D7ED5ED2C3FE90" attach="left" box="[1253,1261,318,332]" fontSize="6" pageId="6" pageNumber="327">3</subScript>
) not in lateral position, moved posterior to first dorsolateral setae
<emphasis id="4CA32526CA71F33512EAED38D4CAFEB3" box="[728,740,344,367]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33512D6ED05D4C2FEAF" attach="left" box="[740,748,357,371]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
. Dorsal setae as in female, inserted on strong tubercles (
<figureCitation id="E6ECE5B1CA71F3351556ED37D68BFE4A" captionStart="FIGURE 4" captionStartId="10.[151,250,1101,1123]" captionTargetBox="[150,1436,214,1066]" captionTargetId="figure@10.[150,1436,214,1066]" captionTargetPageId="10" captionText="FIGURE 4. Georgiobia skuthiae n. sp., Deutonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation, D. peritreme. Scale bars: 100 µm (A), 25 µm (BD)." httpUri="https://zenodo.org/record/2627016/files/figure.png" pageId="6" pageNumber="327">Figs. 4</figureCitation>
AC). Dorsal tubercles cone shaped with irregular striation punctate but not finely granulated (
<figureCitation id="E6ECE5B1CA71F3351511ED1DD3A0FE4A" box="[1315,1422,381,406]" captionStart="FIGURE 4" captionStartId="10.[151,250,1101,1123]" captionTargetBox="[150,1436,214,1066]" captionTargetId="figure@10.[150,1436,214,1066]" captionTargetPageId="10" captionText="FIGURE 4. Georgiobia skuthiae n. sp., Deutonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation, D. peritreme. Scale bars: 100 µm (A), 25 µm (BD)." httpUri="https://zenodo.org/record/2627016/files/figure.png" pageId="6" pageNumber="327">Fig. 4 C</figureCitation>
); tubercles of members of setae
<emphasis id="4CA32526CA71F33511C5EDC2D42BFE67" box="[503,517,418,443]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F3351237EDD0D423FE62" attach="left" box="[517,525,432,446]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
,
<emphasis id="4CA32526CA71F335122FEDC3D407FE66" box="[541,553,419,442]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F335121BEDD0D41FFE62" attach="left" box="[553,561,432,446]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
and of
<emphasis id="4CA32526CA71F33512BEEDC2D4BAFE66" box="[652,660,418,442]" italics="true" pageId="6" pageNumber="327">f</emphasis>
<subScript id="E253FB71CA71F33512A6EDD0D4B2FE62" attach="left" box="[660,668,432,446]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
obviously contiguous. First, second and third pairs of dorsocentral hysterosomals much longer than distances between consecutive setae, setae
<emphasis id="4CA32526CA71F33513C0EDAAD5D0FE3D" box="[1010,1022,458,481]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33513CCEDB7D228FE39" attach="left" box="[1022,1030,471,485]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
and
<emphasis id="4CA32526CA71F335140CEDA8D262FE3D" box="[1086,1100,456,481]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F335147EEDB7D27AFE39" attach="left" box="[1100,1108,471,485]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
setae the longest while setae
<emphasis id="4CA32526CA71F33510A5ED8FD68BFDD4" box="[151,165,495,520]" italics="true" pageId="6" pageNumber="327">h</emphasis>
<subScript id="E253FB71CA71F3351097ED9ED683FDD0" attach="left" box="[165,173,510,524]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
the shortest (
<figureCitation id="E6ECE5B1CA71F3351176ED8FD78BFDD4" box="[324,421,495,520]" captionStart="FIGURE 4" captionStartId="10.[151,250,1101,1123]" captionTargetBox="[150,1436,214,1066]" captionTargetId="figure@10.[150,1436,214,1066]" captionTargetPageId="10" captionText="FIGURE 4. Georgiobia skuthiae n. sp., Deutonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation, D. peritreme. Scale bars: 100 µm (A), 25 µm (BD)." httpUri="https://zenodo.org/record/2627016/files/figure.png" pageId="6" pageNumber="327">Fig. 4 A</figureCitation>
). Setae lengths:
<date id="0A69DFF4CA71F335126FED90D4B9FDD4" box="[605,663,496,524]" pageId="6" pageNumber="327" value="1982-05-02">
<emphasis id="4CA32526CA71F335126FED90D447FDDB" box="[605,617,496,519]" italics="true" pageId="6" pageNumber="327">v</emphasis>
<subScript id="E253FB71CA71F335125BED9ED45FFDD0" attach="left" box="[617,625,510,524]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
82
</date>
;
<emphasis id="4CA32526CA71F3351291ED90D494FDDB" box="[675,698,496,519]" italics="true" pageId="6" pageNumber="327">sc</emphasis>
<subScript id="E253FB71CA71F3351289ED9ED4EDFDD0" attach="left" box="[699,707,510,524]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
110;
<emphasis id="4CA32526CA71F3351330ED90D537FDDB" box="[770,793,496,519]" italics="true" pageId="6" pageNumber="327">sc</emphasis>
<subScript id="E253FB71CA71F335132BED9ED50FFDD0" attach="left" box="[793,801,510,524]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
114;
<emphasis id="4CA32526CA71F3351352ED90D542FDDB" box="[864,876,496,519]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F335135EED9ED55AFDD0" attach="left" box="[876,884,510,524]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
154;
<emphasis id="4CA32526CA71F3351386ED90D5EEFDDB" box="[948,960,496,519]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33513F2ED9ED5E6FDD0" attach="left" box="[960,968,510,524]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
104;
<emphasis id="4CA32526CA71F335143AED90D23AFDDB" box="[1032,1044,496,519]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F3351427ED9ED233FDD0" attach="left" box="[1045,1053,510,524]" fontSize="6" pageId="6" pageNumber="327">3</subScript>
96;
<emphasis id="4CA32526CA71F335147DED8FD273FDD4" box="[1103,1117,495,520]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F335146EED9ED24AFDD0" attach="left" box="[1116,1124,510,524]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
146;
<emphasis id="4CA32526CA71F3351496ED8FD29CFDD4" box="[1188,1202,495,520]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F3351481ED9ED295FDD0" attach="left" box="[1203,1211,510,524]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
122;
<emphasis id="4CA32526CA71F33514C8ED90D328FDDB" box="[1274,1286,496,519]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F3351535ED9ED321FDD0" attach="left" box="[1287,1295,510,524]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
134;
<emphasis id="4CA32526CA71F335157DED90D375FDDB" box="[1359,1371,496,519]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F3351569ED9ED34DFDD0" attach="left" box="[1371,1379,510,524]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
108;
<emphasis id="4CA32526CA71F33510A5EE76D6B1FDF2" box="[151,159,534,558]" italics="true" pageId="6" pageNumber="327">f</emphasis>
<subScript id="E253FB71CA71F33510ACEE44D688FDEE" attach="left" box="[158,166,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
108;
<emphasis id="4CA32526CA71F33510D7EE76D6C3FDF2" box="[229,237,534,558]" italics="true" pageId="6" pageNumber="327">f</emphasis>
<subScript id="E253FB71CA71F33510DFEE44D6DBFDEE" attach="left" box="[237,245,548,562]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
94;
<emphasis id="4CA32526CA71F3351111EE76D71FFDF3" box="[291,305,534,559]" italics="true" pageId="6" pageNumber="327">h</emphasis>
<subScript id="E253FB71CA71F3351103EE44D717FDEE" attach="left" box="[305,313,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
66. Distances between setae:
<emphasis id="4CA32526CA71F33512BAEE77D4BAFDF2" box="[648,660,535,558]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33512A6EE44D4B2FDEE" attach="left" box="[660,668,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
-
<emphasis id="4CA32526CA71F3351297EE76D49DFDF3" box="[677,691,534,559]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F3351281EE44D495FDEE" attach="left" box="[691,699,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
60,
<emphasis id="4CA32526CA71F33512DEEE76D4D4FDF3" box="[748,762,534,559]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F33512C8EE44D52CFDEE" attach="left" box="[762,770,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
-
<emphasis id="4CA32526CA71F3351339EE77D539FDF2" box="[779,791,535,558]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F3351325EE44D531FDEE" attach="left" box="[791,799,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
100,
<emphasis id="4CA32526CA71F335136FEE77D547FDF2" box="[861,873,535,558]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F3351358EE44D55CFDEE" attach="right" box="[874,882,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
-
<emphasis id="4CA32526CA71F3351349EE76D5ADFDF2" box="[891,899,534,558]" italics="true" pageId="6" pageNumber="327">f</emphasis>
<subScript id="E253FB71CA71F33513B1EE44D5A5FDEE" attach="left" box="[899,907,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
80 (distances between members of
<emphasis id="4CA32526CA71F3351528EE77D308FDF2" box="[1306,1318,535,558]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F3351514EE44D300FDEE" attach="left" box="[1318,1326,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
,
<emphasis id="4CA32526CA71F335150EEE76D364FDF3" box="[1340,1354,534,559]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F3351578EE44D37CFDEE" attach="left" box="[1354,1362,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
and
<emphasis id="4CA32526CA71F33515BAEE77D3BAFDF2" box="[1416,1428,535,558]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F33515A6EE44D3B2FDEE" attach="left" box="[1428,1436,548,562]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
setae were not assessed because of the mounting effect and setal insertions on strong tubercles that caused large variations).
</paragraph>
<paragraph id="7E68F934CA71F33510F5EEE5D5B7FD42" blockId="6.[151,1436,151,856]" box="[199,921,645,670]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F33510F5EEE5D778FD42" box="[199,342,645,670]" italics="true" pageId="6" pageNumber="327">Gnathosoma</emphasis>
. Peritreme anastomosed distally, 16 long (
<figureCitation id="E6ECE5B1CA71F3351300EEE5D5A3FD42" box="[818,909,645,670]" captionStart="FIGURE 4" captionStartId="10.[151,250,1101,1123]" captionTargetBox="[150,1436,214,1066]" captionTargetId="figure@10.[150,1436,214,1066]" captionTargetPageId="10" captionText="FIGURE 4. Georgiobia skuthiae n. sp., Deutonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation, D. peritreme. Scale bars: 100 µm (A), 25 µm (BD)." httpUri="https://zenodo.org/record/2627016/files/figure.png" pageId="6" pageNumber="327">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="7E68F934CA71F33510F5EECDD20AFD36" blockId="6.[151,1436,151,856]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F33510F5EECDD722FD18" box="[199,268,685,708]" italics="true" pageId="6" pageNumber="327">Venter</emphasis>
. Ventral striation transverse between 1
<superScript id="89A2547CCA71F33512F5EEC8D4FFFD6A" attach="left" box="[711,721,680,694]" fontSize="6" pageId="6" pageNumber="327">st</superScript>
(
<emphasis id="4CA32526CA71F33512D6EECCD52EFD18" box="[740,768,684,708]" italics="true" pageId="6" pageNumber="327">1a</emphasis>
) and 2
<superScript id="89A2547CCA71F3351362EEC8D54EFD6A" attach="left" box="[848,864,680,694]" fontSize="6" pageId="6" pageNumber="327">nd</superScript>
(
<emphasis id="4CA32526CA71F3351343EECCD5A3FD18" box="[881,909,684,708]" italics="true" pageId="6" pageNumber="327">3a</emphasis>
) pairs of setae, with irregular pattern between 2
<superScript id="89A2547CCA71F3351097EEAFD69BFD01" attach="left" box="[165,181,719,733]" fontSize="6" pageId="6" pageNumber="327">nd</superScript>
and 3
<superScript id="89A2547CCA71F33510CBEEAFD728FD01" attach="left" box="[249,262,719,733]" fontSize="6" pageId="6" pageNumber="327">rd</superScript>
(
<emphasis id="4CA32526CA71F3351124EEB3D71CFD37" box="[278,306,723,747]" italics="true" pageId="6" pageNumber="327">4a</emphasis>
) pairs of setae and transverse from setae
<emphasis id="4CA32526CA71F3351333EEB3D533FD37" box="[769,797,723,747]" italics="true" pageId="6" pageNumber="327">4a</emphasis>
to aggenital (
<emphasis id="4CA32526CA71F335138AEEB4D5FAFD37" box="[952,980,724,747]" italics="true" pageId="6" pageNumber="327">ag</emphasis>
) setae.
</paragraph>
<paragraph id="7E68F934CA71F33510F5EE98D57FFC8B" blockId="6.[151,1436,151,856]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F33510F5EE98D6D2FCD3" box="[199,252,760,783]" italics="true" pageId="6" pageNumber="327">Legs</emphasis>
. Leg I slightly shorter than body length, legs II to IV less than half the body length. Leg I 440 long (measured from femur to tarsus), leg II 172, leg III 176, leg IV 190. Length of segments of leg I as follows: femur 172, genu 32, tibia 144, tarsus 91. Leg setal counts as follows:
</paragraph>
<paragraph id="7E68F934CA71F33510A5EFE8D4E9FC43" blockId="6.[151,711,903,1035]" box="[151,711,903,927]" pageId="6" pageNumber="327">
<date id="0A69DFF4CA71F33510A5EFE8D6FBFC43" box="[151,213,903,927]" pageId="6" pageNumber="327">I 2</date>
¯ 1 ¯ 6 ¯ 2 9 + (1) 9 + (1) + 2 duplexes;
</paragraph>
<paragraph id="7E68F934CA71F33510A5EFCBD402FC1F" blockId="6.[151,711,903,1035]" box="[151,556,939,963]" pageId="6" pageNumber="327">
<date id="0A69DFF4CA71F33510A5EFCBD6FBFC1F" box="[151,213,939,963]" pageId="6" pageNumber="327">II 2</date>
¯ 0 3 ¯ 2 3 ¯ 7 + 1 duplex;
</paragraph>
<paragraph id="7E68F934CA71F33510A5EFB0D7DCFC3B" blockId="6.[151,711,903,1035]" box="[151,498,975,999]" pageId="6" pageNumber="327">
<date id="0A69DFF4CA71F33510A5EFB0D6FBFC3B" box="[151,213,975,999]" pageId="6" pageNumber="327">III 1</date>
¯ 1 2 2 ¯ 3 ¯ 6 + (1);
</paragraph>
<paragraph id="7E68F934CA71F33510A5EF93D7F7FBD7" blockId="6.[151,711,903,1035]" box="[151,473,1010,1035]" pageId="6" pageNumber="327">
<date id="0A69DFF4CA71F33510A5EF93D6FBFBD7" box="[151,213,1011,1035]" pageId="6" pageNumber="327">IV 1</date>
¯ 0 1 2 [1] ¯ 3 6.
</paragraph>
<paragraph id="7E68F934CA71F33510F5E85BD40BFB3E" blockId="6.[151,1436,1082,1853]" pageId="6" pageNumber="327">
True claws and empodia pad-like as in female. First primiventral setae (
<emphasis id="4CA32526CA71F33513D4E85CD22EFB8F" box="[998,1024,1084,1107]" italics="true" pageId="6" pageNumber="327">pv</emphasis>
) absent on tarsus I, present on other tarsi. Tarsi without tectal setae (
<emphasis id="4CA32526CA71F335123EE800D40EFBAB" box="[524,544,1120,1143]" italics="true" pageId="6" pageNumber="327">tc</emphasis>
-
<emphasis id="4CA32526CA71F3351201E800D469FBAB" box="[563,583,1120,1143]" italics="true" pageId="6" pageNumber="327">tc</emphasis>
). Large strong serrate seta present on some leg segments (
<figureCitation id="E6ECE5B1CA71F3351534E83ED34DFBAB" box="[1286,1379,1118,1143]" captionStart="FIGURE 4" captionStartId="10.[151,250,1101,1123]" captionTargetBox="[150,1436,214,1066]" captionTargetId="figure@10.[150,1436,214,1066]" captionTargetPageId="10" captionText="FIGURE 4. Georgiobia skuthiae n. sp., Deutonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation, D. peritreme. Scale bars: 100 µm (A), 25 µm (BD)." httpUri="https://zenodo.org/record/2627016/files/figure.png" pageId="6" pageNumber="327">Fig. 4A</figureCitation>
), the longest on femur I, half the length on other segments: on femur I 120, on genu
<date id="0A69DFF4CA71F335143CE8E3D215FB47" box="[1038,1083,1154,1179]" pageId="6" pageNumber="327" value="1966-01">I 66</date>
(twice as long as leg segment), on tibia
<date id="0A69DFF4CA71F33510C0E8C8D7ADFB63" box="[242,387,1190,1215]" pageId="6" pageNumber="327" value="1940-03" valueMax="1940-04-31" valueMin="1940-03-00">III and IV 40</date>
and 56 (slightly longer than leg segment), respectively. Tarsus III with short solenidion, about 1/3 the length of distal tactile setae.
</paragraph>
<paragraph id="7E68F934CA71F33510F5E88ED766FAF2" blockId="6.[151,1436,1082,1853]" pageId="6" pageNumber="327">
Protonymph (n=1), (
<figureCitation id="E6ECE5B1CA71F335119CE88ED7DCFADB" box="[430,498,1262,1287]" captionStart="FIGURE 5" captionStartId="11.[151,250,1093,1115]" captionTargetBox="[150,1436,195,1071]" captionTargetId="figure@11.[150,1436,195,1071]" captionTargetPageId="11" captionText="FIGURE 5. Georgiobia skuthiae n. sp., Protonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation. Scale bars: 50 µm (A), 25 µm (B, C)" httpUri="https://zenodo.org/record/2627018/files/figure.png" pageId="6" pageNumber="327">Fig. 5</figureCitation>
). Body length 370 including gnathosoma, width 220, gnathosoma
<date id="0A69DFF4CA71F33514D2E88FD330FAD7" box="[1248,1310,1263,1291]" pageId="6" pageNumber="327" value="1970-05-02">
70,
<emphasis id="4CA32526CA71F3351538E890D338FADB" box="[1290,1302,1264,1287]" italics="true" pageId="6" pageNumber="327">v</emphasis>
<subScript id="E253FB71CA71F3351524E89DD330FAD7" attach="left" box="[1302,1310,1277,1291]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
</date>
-
<emphasis id="4CA32526CA71F3351515E88ED31BFADB" box="[1319,1333,1262,1287]" italics="true" pageId="6" pageNumber="327">h</emphasis>
<subScript id="E253FB71CA71F3351507E89DD313FAD7" attach="left" box="[1333,1341,1277,1291]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
280,
<emphasis id="4CA32526CA71F335154FE890D3BAFADB" box="[1405,1428,1264,1287]" italics="true" pageId="6" pageNumber="327">sc</emphasis>
<subScript id="E253FB71CA71F33515A6E89DD3B2FAD7" attach="left" box="[1428,1436,1277,1291]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
153,
<emphasis id="4CA32526CA71F33510FDE976D6F5FAF1" box="[207,219,1302,1325]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33510E9E944D6CDFAEE" attach="left" box="[219,227,1316,1330]" fontSize="6" pageId="6" pageNumber="327">3</subScript>
<emphasis id="4CA32526CA71F33510CDE976D725FAF1" box="[255,267,1302,1325]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F3351139E944D73DFAEE" attach="left" box="[267,275,1316,1330]" fontSize="6" pageId="6" pageNumber="327">3</subScript>
164.
</paragraph>
<paragraph id="7E68F934CA71F33510F5E95DD32AF95A" blockId="6.[151,1436,1082,1853]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F33510F5E95DD70FFA88" box="[199,289,1341,1364]" italics="true" pageId="6" pageNumber="327">Dorsum</emphasis>
. Prodorsal striation longitudinal becoming punctate close to setae
<date id="0A69DFF4CA71F3351415E95DD212FA85" box="[1063,1084,1341,1369]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F3351415E95DD21DFA88" box="[1063,1075,1341,1364]" italics="true" pageId="6" pageNumber="327">v</emphasis>
<subScript id="E253FB71CA71F3351406E92BD212FA85" attach="right" box="[1076,1084,1355,1369]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
</date>
, medial alveoli pattern almost absent (difficult to see), lateral striation longitudinal (
<figureCitation id="E6ECE5B1CA71F33512C9E903D573FAA7" box="[763,861,1379,1404]" captionStart="FIGURE 5" captionStartId="11.[151,250,1093,1115]" captionTargetBox="[150,1436,195,1071]" captionTargetId="figure@11.[150,1436,195,1071]" captionTargetPageId="11" captionText="FIGURE 5. Georgiobia skuthiae n. sp., Protonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation. Scale bars: 50 µm (A), 25 µm (B, C)" httpUri="https://zenodo.org/record/2627018/files/figure.png" pageId="6" pageNumber="327">Fig. 5 B</figureCitation>
). Overall hysterosomal striation pattern transverse between members of setae
<emphasis id="4CA32526CA71F33511FEE9E8D7F6FA43" box="[460,472,1416,1439]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33511EAE9F5D7CEFA7F" attach="left" box="[472,480,1429,1443]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
to
<emphasis id="4CA32526CA71F3351234E9E7D420FA43" box="[518,526,1415,1439]" italics="true" pageId="6" pageNumber="327">f</emphasis>
<subScript id="E253FB71CA71F335123CE9F5D438FA7F" attach="left" box="[526,534,1429,1443]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
, longitudinal laterally. Humeral setae (
<emphasis id="4CA32526CA71F33513E1E9E8D5F1FA43" box="[979,991,1416,1439]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33513EDE9F5D5C9FA7F" attach="left" box="[991,999,1429,1443]" fontSize="6" pageId="6" pageNumber="327">3</subScript>
) not in lateral position, moved behind first dorsolateral setae
<emphasis id="4CA32526CA71F33511A5E9CFD78DFA1A" box="[407,419,1455,1478]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F3351191E9DCD785FA16" attach="left" box="[419,427,1468,1482]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
. Dorsal setae relatively long, linear lanceolate, serrate, inserted on strong tubercles (
<figureCitation id="E6ECE5B1CA71F3351556E9CDD6C8FA31" captionStart="FIGURE 5" captionStartId="11.[151,250,1093,1115]" captionTargetBox="[150,1436,195,1071]" captionTargetId="figure@11.[150,1436,195,1071]" captionTargetPageId="11" captionText="FIGURE 5. Georgiobia skuthiae n. sp., Protonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation. Scale bars: 50 µm (A), 25 µm (B, C)" httpUri="https://zenodo.org/record/2627018/files/figure.png" pageId="6" pageNumber="327">Figs. 5 A, C</figureCitation>
). Dorsal tubercles cone shaped with irregularly broken striation (
<figureCitation id="E6ECE5B1CA71F33513FBE9B4D205FA31" box="[969,1067,1492,1517]" captionStart="FIGURE 5" captionStartId="11.[151,250,1093,1115]" captionTargetBox="[150,1436,195,1071]" captionTargetId="figure@11.[150,1436,195,1071]" captionTargetPageId="11" captionText="FIGURE 5. Georgiobia skuthiae n. sp., Protonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation. Scale bars: 50 µm (A), 25 µm (B, C)" httpUri="https://zenodo.org/record/2627018/files/figure.png" pageId="6" pageNumber="327">Fig. 5 C</figureCitation>
). First, second and third pairs of dorsocentral hysterosomal setae surpass insertions of consecutive setae, setae
<emphasis id="4CA32526CA71F3351423E99AD233F9CD" box="[1041,1053,1530,1553]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F335142FEA67D20BF9C9" attach="left" box="[1053,1061,1543,1557]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
the longest, setae
<emphasis id="4CA32526CA71F33514C9E998D327F9CD" box="[1275,1289,1528,1553]" italics="true" pageId="6" pageNumber="327">h</emphasis>
<subScript id="E253FB71CA71F335153BEA67D33FF9C9" attach="left" box="[1289,1297,1543,1557]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
the shortest (
<figureCitation id="E6ECE5B1CA71F33510ADEA7FD72AF9E4" box="[159,260,1567,1592]" captionStart="FIGURE 5" captionStartId="11.[151,250,1093,1115]" captionTargetBox="[150,1436,195,1071]" captionTargetId="figure@11.[150,1436,195,1071]" captionTargetPageId="11" captionText="FIGURE 5. Georgiobia skuthiae n. sp., Protonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation. Scale bars: 50 µm (A), 25 µm (B, C)" httpUri="https://zenodo.org/record/2627018/files/figure.png" pageId="6" pageNumber="327">Fig. 5 A</figureCitation>
). Setae lengths:
<date id="0A69DFF4CA71F33511F3EA40D7D2F9E4" box="[449,508,1567,1596]" pageId="6" pageNumber="327" value="1962-05-02">
<emphasis id="4CA32526CA71F33511F3EA40D7E3F9EB" box="[449,461,1568,1591]" italics="true" pageId="6" pageNumber="327">v</emphasis>
<subScript id="E253FB71CA71F33511FFEA4ED7FBF9E0" attach="left" box="[461,469,1582,1596]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
62
</date>
;
<emphasis id="4CA32526CA71F3351238EA40D40FF9EB" box="[522,545,1568,1591]" italics="true" pageId="6" pageNumber="327">sc</emphasis>
<subScript id="E253FB71CA71F3351213EA4ED407F9E0" attach="left" box="[545,553,1582,1596]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
80;
<emphasis id="4CA32526CA71F335126DEA40D458F9EB" box="[607,630,1568,1591]" italics="true" pageId="6" pageNumber="327">sc</emphasis>
<subScript id="E253FB71CA71F3351244EA4ED450F9E0" attach="left" box="[630,638,1582,1596]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
84;
<emphasis id="4CA32526CA71F3351281EA40D491F9EB" box="[691,703,1568,1591]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F335128DEA4ED4E9F9E0" attach="left" box="[703,711,1582,1596]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
126;
<emphasis id="4CA32526CA71F3351339EA40D539F9EB" box="[779,791,1568,1591]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F3351325EA4ED531F9E0" attach="left" box="[791,799,1582,1596]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
74;
<emphasis id="4CA32526CA71F3351366EA40D54EF9EB" box="[852,864,1568,1591]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F3351353EA4ED547F9E0" attach="left" box="[865,873,1582,1596]" fontSize="6" pageId="6" pageNumber="327">3</subScript>
62;
<emphasis id="4CA32526CA71F33513ACEA7FD582F9E4" box="[926,940,1567,1592]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F335139EEA4ED59AF9E0" attach="left" box="[940,948,1582,1596]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
114;
<emphasis id="4CA32526CA71F33513C4EA7FD22AF9E4" box="[1014,1028,1567,1592]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F3351436EA4ED222F9E0" attach="left" box="[1028,1036,1582,1596]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
82;
<emphasis id="4CA32526CA71F3351473EA40D263F9EB" box="[1089,1101,1568,1591]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F335147FEA4ED27BF9E0" attach="left" box="[1101,1109,1582,1596]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
98;
<emphasis id="4CA32526CA71F33514B9EA40D2B9F9EB" box="[1163,1175,1568,1591]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F33514A5EA4ED2B1F9E0" attach="left" box="[1175,1183,1582,1596]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
76;
<emphasis id="4CA32526CA71F33514E6EA40D2F2F9E4" box="[1236,1244,1568,1592]" italics="true" pageId="6" pageNumber="327">f</emphasis>
<subScript id="E253FB71CA71F33514EEEA4ED2CAF9E0" attach="left" box="[1244,1252,1582,1596]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
74;
<emphasis id="4CA32526CA71F335152BEA40D30FF9E4" box="[1305,1313,1568,1592]" italics="true" pageId="6" pageNumber="327">f</emphasis>
<subScript id="E253FB71CA71F3351513EA4ED307F9E0" attach="left" box="[1313,1321,1582,1596]" fontSize="6" pageId="6" pageNumber="327">2</subScript>
64;
<emphasis id="4CA32526CA71F3351568EA7FD346F9E4" box="[1370,1384,1567,1592]" italics="true" pageId="6" pageNumber="327">h</emphasis>
<subScript id="E253FB71CA71F335155AEA4ED35EF9E0" attach="left" box="[1384,1392,1582,1596]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
46. Distances between setae:
<emphasis id="4CA32526CA71F33511F2EA27D7E2F982" box="[448,460,1607,1630]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F33511FEEA34D7FAF9BE" attach="left" box="[460,468,1620,1634]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
-
<emphasis id="4CA32526CA71F33511EFEA26D7C5F983" box="[477,491,1606,1631]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F33511D9EA34D7DDF9BE" attach="left" box="[491,499,1620,1634]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
44,
<emphasis id="4CA32526CA71F3351219EA26D417F983" box="[555,569,1606,1631]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F335120BEA34D46FF9BE" attach="left" box="[569,577,1620,1634]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
-
<emphasis id="4CA32526CA71F3351278EA27D478F982" box="[586,598,1607,1630]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F3351264EA34D470F9BE" attach="left" box="[598,606,1620,1634]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
64,
<emphasis id="4CA32526CA71F33512A4EA27D48CF982" box="[662,674,1607,1630]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F3351290EA34D484F9BE" attach="left" box="[674,682,1620,1634]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
-
<emphasis id="4CA32526CA71F3351281EA26D495F982" box="[691,699,1606,1630]" italics="true" pageId="6" pageNumber="327">f</emphasis>
<subScript id="E253FB71CA71F3351289EA34D4EDF9BE" attach="left" box="[699,707,1620,1634]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
66 (distance between members of
<emphasis id="4CA32526CA71F335146CEA27D244F982" box="[1118,1130,1607,1630]" italics="true" pageId="6" pageNumber="327">c</emphasis>
<subScript id="E253FB71CA71F3351458EA34D25CF9BE" attach="left" box="[1130,1138,1620,1634]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
,
<emphasis id="4CA32526CA71F33514B1EA26D2BFF983" box="[1155,1169,1606,1631]" italics="true" pageId="6" pageNumber="327">d</emphasis>
<subScript id="E253FB71CA71F33514A3EA34D2B7F9BE" attach="left" box="[1169,1177,1620,1634]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
and
<emphasis id="4CA32526CA71F33514E4EA27D2CCF982" box="[1238,1250,1607,1630]" italics="true" pageId="6" pageNumber="327">e</emphasis>
<subScript id="E253FB71CA71F33514D1EA34D2C5F9BE" attach="left" box="[1251,1259,1620,1634]" fontSize="6" pageId="6" pageNumber="327">1</subScript>
setae were not assessed because of the mounting and setal insertions on strong tubercles that cause large variations).
</paragraph>
<paragraph id="7E68F934CA71F33510F5EAF1D565F976" blockId="6.[151,1436,1082,1853]" box="[199,843,1681,1706]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F33510F5EAF1D778F976" box="[199,342,1681,1706]" italics="true" pageId="6" pageNumber="327">Gnathosoma</emphasis>
. Peritreme ending in a reduced anastomosis.
</paragraph>
<paragraph id="7E68F934CA71F33510F5EAD9D2F4F913" blockId="6.[151,1436,1082,1853]" box="[199,1242,1716,1744]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F33510F5EAD9D722F90C" box="[199,268,1721,1744]" italics="true" pageId="6" pageNumber="327">Venter</emphasis>
. Ventral striation entirely transverse between 1
<superScript id="89A2547CCA71F335132FEAD4D509F91E" attach="left" box="[797,807,1716,1730]" fontSize="6" pageId="6" pageNumber="327">st</superScript>
(
<emphasis id="4CA32526CA71F3351305EAD8D57CF90C" box="[823,850,1720,1744]" italics="true" pageId="6" pageNumber="327">1a</emphasis>
) and 3
<superScript id="89A2547CCA71F3351393EAD4D580F91E" attach="left" box="[929,942,1716,1730]" fontSize="6" pageId="6" pageNumber="327">rd</superScript>
(
<emphasis id="4CA32526CA71F335138CEAD9D5F4F90C" box="[958,986,1721,1744]" italics="true" pageId="6" pageNumber="327">ag</emphasis>
) pairs of ventral setae.
</paragraph>
<paragraph id="7E68F934CA71F33510F5EABDD73FF8E1" blockId="6.[151,1436,1082,1853]" pageId="6" pageNumber="327">
<emphasis id="4CA32526CA71F33510F5EABDD6D2F928" box="[199,252,1757,1780]" italics="true" pageId="6" pageNumber="327">Legs</emphasis>
. Leg I obviously shorter than body length, 224 long; legs II to IV about ¼ the body length: leg II 130, leg III 125, leg IV 135. Length of segments of leg I as follows: femur 80, genu 28, tibia 60, tarsus 56. Leg setal counts as follows:
</paragraph>
<paragraph id="7E68F934CA71F33510A5EB0DD4A4F858" blockId="6.[151,650,1900,2032]" box="[151,650,1900,1924]" pageId="6" pageNumber="327">
<date id="0A69DFF4CA71F33510A5EB0DD6FBF858" box="[151,213,1900,1924]" pageId="6" pageNumber="327">I 2</date>
¯ 0 ¯ 3 ¯ 2 ¯ 5 + (1) 5 + 2 duplexes;
</paragraph>
<paragraph id="7E68F934CA71F33510A5EBF0D402F874" blockId="6.[151,650,1900,2032]" box="[151,556,1936,1960]" pageId="6" pageNumber="327">
<date id="0A69DFF4CA71F33510A5EBF0D6FBF874" box="[151,213,1936,1960]" pageId="6" pageNumber="327">II 1</date>
¯ 0 2 ¯ 2 3 ¯ 7 + 1 duplex;
</paragraph>
<paragraph id="7E68F934CA71F33510A5EBD5D79AF810" blockId="6.[151,650,1900,2032]" box="[151,436,1972,1996]" pageId="6" pageNumber="327">
<date id="0A69DFF4CA71F33510A5EBD5D6FBF810" box="[151,213,1972,1996]" pageId="6" pageNumber="327">III 1</date>
¯ 0 2 1 ¯ 3 ¯ 6;
</paragraph>
<paragraph id="7E68F934CA71F33510A5EBB8D79FF82C" blockId="6.[151,650,1900,2032]" box="[151,433,2007,2032]" pageId="6" pageNumber="327">IV 0 ¯ 0 2 1 ¯ 3 6.</paragraph>
<caption id="2AA8A9BCCA70F33410A5EBD3D4CCF834" httpUri="https://zenodo.org/record/2627010/files/figure.png" pageId="7" pageNumber="328" startId="7.[151,250,1971,1993]" targetBox="[179,1415,195,1955]" targetPageId="7">
<paragraph id="7E68F934CA70F33410A5EBD3D4CCF834" blockId="7.[151,1436,1971,2027]" pageId="7" pageNumber="328">
<emphasis id="4CA32526CA70F33410A5EBD3D720F814" bold="true" box="[151,270,1971,1993]" pageId="7" pageNumber="328">FIGURE 1</emphasis>
.
<taxonomicName id="B9D782B7CA70F334112EEBD3D7C5F815" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[284,491,1971,1993]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="328" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA70F334112EEBD3D7C5F815" box="[284,491,1971,1993]" italics="true" pageId="7" pageNumber="328">Georgiobia skuthiae</emphasis>
</taxonomicName>
<emphasis id="4CA32526CA70F33411C6EBD3D400F814" bold="true" box="[500,558,1971,1992]" pageId="7" pageNumber="328">
<taxonomicNameLabel id="5790985DCA70F33411C6EBD3D400F814" box="[500,558,1971,1992]" pageId="7" pageNumber="328" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female. A. dorsal aspect, B. detail of medial propodosomal striation, C. dorsal body seta (
<emphasis id="4CA32526CA70F33410FFEBB3D6F6F834" box="[205,216,2003,2024]" italics="true" pageId="7" pageNumber="328">e</emphasis>
<subScript id="E253FB71CA70F33410EAEBBFD6F1F837" attach="left" box="[216,223,2015,2027]" fontSize="5" pageId="7" pageNumber="328">2</subScript>
). Scale bars = 100 µm (A), 10 µm (B), 25 µm (C).
</paragraph>
</caption>
<caption id="2AA8A9BCCA7FF33B10A5EBD3D5DBF83B" httpUri="https://zenodo.org/record/2627012/files/figure.png" pageId="8" pageNumber="329" startId="8.[151,250,1971,1993]" targetBox="[279,1436,238,1924]" targetPageId="8">
<paragraph id="7E68F934CA7FF33B10A5EBD3D5DBF83B" blockId="8.[151,1436,1971,2023]" pageId="8" pageNumber="329">
<emphasis id="4CA32526CA7FF33B10A5EBD3D722F814" bold="true" box="[151,268,1971,1993]" pageId="8" pageNumber="329">FIGURE 2</emphasis>
.
<taxonomicName id="B9D782B7CA7FF33B112AEBD3D7C8F815" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[280,486,1971,1993]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="329" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7FF33B112AEBD3D7C8F815" box="[280,486,1971,1993]" italics="true" pageId="8" pageNumber="329">Georgiobia skuthiae</emphasis>
</taxonomicName>
<emphasis id="4CA32526CA7FF33B11DFEBD3D408F814" bold="true" box="[493,550,1971,1992]" pageId="8" pageNumber="329">
<taxonomicNameLabel id="5790985DCA7FF33B11DFEBD3D408F814" box="[493,550,1971,1992]" pageId="8" pageNumber="329" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female. A. tarsus I, B. tibia I, C. empodium and claws I, D. empodium and claws II, E. coxisternal setae
<emphasis id="4CA32526CA7FF33B1177EBB1D770F83B" box="[325,350,2001,2023]" italics="true" pageId="8" pageNumber="329">1b</emphasis>
,
<emphasis id="4CA32526CA7FF33B1159EBB1D7ACF83B" box="[363,386,2001,2023]" italics="true" pageId="8" pageNumber="329">1c</emphasis>
(from left to right). Scale bars = 25 µm (AB), 10 µm (CE).
</paragraph>
</caption>
<caption id="2AA8A9BCCA7EF33A10A5EBD7D3A8F837" httpUri="https://zenodo.org/record/2627014/files/figure.png" pageId="9" pageNumber="330" startId="9.[151,250,1975,1997]" targetBox="[190,1400,196,1958]" targetPageId="9">
<paragraph id="7E68F934CA7EF33A10A5EBD7D3A8F837" blockId="9.[151,1436,1975,2027]" pageId="9" pageNumber="330">
<emphasis id="4CA32526CA7EF33A10A5EBD7D73FF810" bold="true" box="[151,273,1975,1997]" pageId="9" pageNumber="330">FIGURE 3</emphasis>
.
<taxonomicName id="B9D782B7CA7EF33A1113EBD7D7DAF811" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[289,500,1975,1997]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="330" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7EF33A1113EBD7D7DAF811" box="[289,500,1975,1997]" italics="true" pageId="9" pageNumber="330">Georgiobia skuthiae</emphasis>
</taxonomicName>
<emphasis id="4CA32526CA7EF33A11CDEBD7D412F810" bold="true" box="[511,572,1975,1996]" pageId="9" pageNumber="330">
<taxonomicNameLabel id="5790985DCA7EF33A11CDEBD7D412F810" box="[511,572,1975,1996]" pageId="9" pageNumber="330" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female. A. tarsus and tibia II, B. tarsus and tibia III, C. tarsus and tibia IV, D. palptarsus and palptibia, E. peritremal enlargement, F. ventral striation. Scale bars = 25 µm (AC), 10 µm (DE), 50 µm (F).
</paragraph>
</caption>
<caption id="2AA8A9BCCA7DF33910A5E82DD4A0FB5E" httpUri="https://zenodo.org/record/2627016/files/figure.png" pageId="10" pageNumber="331" startId="10.[151,250,1101,1123]" targetBox="[150,1436,214,1066]" targetPageId="10">
<paragraph id="7E68F934CA7DF33910A5E82DD4A0FB5E" blockId="10.[151,1436,1101,1154]" pageId="10" pageNumber="331">
<emphasis id="4CA32526CA7DF33910A5E82DD723FBBF" bold="true" box="[151,269,1101,1123]" pageId="10" pageNumber="331">FIGURE 4</emphasis>
.
<taxonomicName id="B9D782B7CA7DF339112BE82DD7C6FBBF" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[281,488,1101,1123]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7DF339112BE82DD7C6FBBF" box="[281,488,1101,1123]" italics="true" pageId="10" pageNumber="331">Georgiobia skuthiae</emphasis>
</taxonomicName>
<emphasis id="4CA32526CA7DF33911DCE82ED406FBBF" bold="true" box="[494,552,1102,1123]" pageId="10" pageNumber="331">
<taxonomicNameLabel id="5790985DCA7DF33911DCE82ED406FBBF" box="[494,552,1102,1123]" pageId="10" pageNumber="331" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Deutonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation, D. peritreme. Scale bars: 100 µm (A), 25 µm (BD).
</paragraph>
</caption>
<paragraph id="7E68F934CA7DF33910F5E8D0D283FAE8" blockId="10.[151,1436,1199,1980]" pageId="10" pageNumber="331">
True claws and empodia as in deutonymph. First primiventral setae (
<emphasis id="4CA32526CA7DF33913E6E8D0D5D9FB1B" box="[980,1015,1200,1223]" italics="true" pageId="10" pageNumber="331">pv</emphasis>
) absent on tarsus I, tarsi I to tarsi III without tectal setae (
<emphasis id="4CA32526CA7DF33911B2E8B4D7BAFB37" box="[384,404,1236,1259]" italics="true" pageId="10" pageNumber="331">tc</emphasis>
-
<emphasis id="4CA32526CA7DF3391195E8B4D795FB37" box="[423,443,1236,1259]" italics="true" pageId="10" pageNumber="331">tc</emphasis>
). Large strong serrate setae present on some leg segments, the longest on femur I, half the length on genu I and far smaller on tibia III and IV (
<figureCitation id="E6ECE5B1CA7DF339132FE897D5ACFAD3" box="[797,898,1271,1296]" captionStart="FIGURE 5" captionStartId="11.[151,250,1093,1115]" captionTargetBox="[150,1436,195,1071]" captionTargetId="figure@11.[150,1436,195,1071]" captionTargetPageId="11" captionText="FIGURE 5. Georgiobia skuthiae n. sp., Protonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation. Scale bars: 50 µm (A), 25 µm (B, C)" httpUri="https://zenodo.org/record/2627018/files/figure.png" pageId="10" pageNumber="331">Fig. 5 A</figureCitation>
): on femur I 108 (longer than leg segment), on genu
<date id="0A69DFF4CA7DF33910E6E97CD72FFAEF" box="[212,257,1307,1331]" pageId="10" pageNumber="331" value="1948-01">I 48</date>
(less than twice as long as leg segment), on tibia
<date id="0A69DFF4CA7DF339131EE97CD23DFAEF" box="[812,1043,1307,1332]" pageId="10" pageNumber="331">III and IV 18 and 24</date>
, respectively.
</paragraph>
<paragraph id="7E68F934CA7DF33910F5E95FD7A5FA84" blockId="10.[151,1436,1199,1980]" box="[199,395,1343,1368]" pageId="10" pageNumber="331">Male. Not known</paragraph>
</subSubSection>
<subSubSection id="36CDAABFCA7DF33910F5E903D735FA18" pageId="10" pageNumber="331" type="etymology">
<paragraph id="7E68F934CA7DF33910F5E903D735FA18" blockId="10.[151,1436,1199,1980]" pageId="10" pageNumber="331">
<emphasis id="4CA32526CA7DF33910F5E903D760FAA0" bold="true" box="[199,334,1379,1404]" pageId="10" pageNumber="331">Etymology.</emphasis>
The specific epithet “skuthiae” refers to the old Greek name of the region located in the north of the black sea, ΣκΥΘΊα (= Scythia), the location were the new species was collected, meaning from Skuthia after latinisation.
</paragraph>
</subSubSection>
<subSubSection id="36CDAABFCA7DF33810F5E9AFD449F907" lastPageId="11" lastPageNumber="332" pageId="10" pageNumber="331" type="discussion">
<paragraph id="7E68F934CA7DF33910F5E9AFD782F940" blockId="10.[151,1436,1199,1980]" pageId="10" pageNumber="331">
<emphasis id="4CA32526CA7DF33910F5E9AFD715FA34" bold="true" box="[199,315,1487,1512]" pageId="10" pageNumber="331">Remarks.</emphasis>
Initially we were reluctant to create a new species because of the similarities between the leg setal counts and that of the striation patterns of the medial part of the propodosoma found in
<taxonomicName id="B9D782B7CA7DF339145CE994D2C4F9D0" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[1134,1258,1523,1548]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7DF339145CE994D2C4F9D0" box="[1134,1258,1523,1548]" italics="true" pageId="10" pageNumber="331">G. skuthiae</emphasis>
</taxonomicName>
<emphasis id="4CA32526CA7DF33914C3E993D31EF9D0" bold="true" box="[1265,1328,1523,1548]" pageId="10" pageNumber="331">
<taxonomicNameLabel id="5790985DCA7DF33914C3E993D31EF9D0" box="[1265,1328,1523,1548]" pageId="10" pageNumber="331" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and in
<taxonomicName id="B9D782B7CA7DF33915B6E994D73BF9EC" baseAuthorityName="Reck" baseAuthorityYear="1956" class="Arachnida" family="Tetranychidae" genus="Petrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="shirakensis">
<emphasis id="4CA32526CA7DF33915B6E994D73BF9EC" italics="true" pageId="10" pageNumber="331">G. shirakensis</emphasis>
</taxonomicName>
despite other obvious morphological differences. Following a careful search of the descriptions of other species belonging to the genus
<taxonomicName id="B9D782B7CA7DF3391238EA5BD4A4F988" authorityName="sensu Wainstein" authorityYear="1960" box="[522,650,1595,1620]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="4CA32526CA7DF3391238EA5BD4A4F988" box="[522,650,1595,1620]" italics="true" pageId="10" pageNumber="331">Georgiobia</emphasis>
</taxonomicName>
, particularly African ones, and after a comparison with some
<typeStatus id="A16C4796CA7DF339155EEA5CD3B2F988" box="[1388,1436,1596,1620]" pageId="10" pageNumber="331">type</typeStatus>
specimens of
<taxonomicName id="B9D782B7CA7DF339110BEA00D7F7F9A4" baseAuthorityName="Reck" baseAuthorityYear="1956" box="[313,473,1631,1656]" class="Arachnida" family="Tetranychidae" genus="Petrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="shirakensis">
<emphasis id="4CA32526CA7DF339110BEA00D7F7F9A4" box="[313,473,1631,1656]" italics="true" pageId="10" pageNumber="331">G. shirakensis</emphasis>
</taxonomicName>
from Recks collection, we concluded that the Ukrainian specimens reported here constitute a new species.
</paragraph>
<paragraph id="7E68F934CA7DF33910F5EAC7D466F860" blockId="10.[151,1436,1199,1980]" pageId="10" pageNumber="331">
<taxonomicName id="B9D782B7CA7DF33910F5EAC7D785F91C" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[199,427,1703,1728]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7DF33910F5EAC7D785F91C" box="[199,427,1703,1728]" italics="true" pageId="10" pageNumber="331">Georgiobia skuthiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5790985DCA7DF3391180EAC7D7DEF963" box="[434,496,1703,1728]" pageId="10" pageNumber="331" rank="species">
<emphasis id="4CA32526CA7DF3391180EAC7D7C7F91C" bold="true" box="[434,489,1703,1728]" pageId="10" pageNumber="331">n. sp</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="B9D782B7CA7DF3391214EAC8D4EDF91C" baseAuthorityName="Reck" baseAuthorityYear="1956" box="[550,707,1703,1728]" class="Arachnida" family="Tetranychidae" genus="Petrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="shirakensis">
<emphasis id="4CA32526CA7DF3391214EAC8D4EDF91C" box="[550,707,1703,1728]" italics="true" pageId="10" pageNumber="331">G. shirakensis</emphasis>
</taxonomicName>
have identical leg setal counts but the latter is also identical to
<taxonomicName id="B9D782B7CA7DF33915B6EAC8D6D8F93F" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" baseAuthorityName="Smith Meyer" baseAuthorityYear="1974" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="crassata">
<emphasis id="4CA32526CA7DF33915B6EAC8D6D8F93F" italics="true" pageId="10" pageNumber="331">G. crassata</emphasis>
</taxonomicName>
regarding this criterion. However,
<taxonomicName id="B9D782B7CA7DF33912B4EAACD50AF938" baseAuthorityName="Reck" baseAuthorityYear="1956" box="[646,804,1739,1764]" class="Arachnida" family="Tetranychidae" genus="Petrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="shirakensis">
<emphasis id="4CA32526CA7DF33912B4EAACD50AF938" box="[646,804,1739,1764]" italics="true" pageId="10" pageNumber="331">G. shirakensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B9D782B7CA7DF339136FEAACD5F2F93F" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" baseAuthorityName="Smith Meyer" baseAuthorityYear="1974" box="[861,988,1740,1763]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="crassata">
<emphasis id="4CA32526CA7DF339136FEAACD5F2F93F" box="[861,988,1740,1763]" italics="true" pageId="10" pageNumber="331">G. crassata</emphasis>
</taxonomicName>
clearly differ according to the size and shape of their dorsal setae. Moreover,
<taxonomicName id="B9D782B7CA7DF3391270EA90D490F8D4" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[578,702,1775,1800]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7DF3391270EA90D490F8D4" box="[578,702,1775,1800]" italics="true" pageId="10" pageNumber="331">G. skuthiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5790985DCA7DF33912F7EA8FD52AF8DB" box="[709,772,1775,1800]" pageId="10" pageNumber="331" rank="species">
<emphasis id="4CA32526CA7DF33912F7EA8FD4D2F8D4" bold="true" box="[709,764,1775,1800]" pageId="10" pageNumber="331">n. sp</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="B9D782B7CA7DF3391308EA90D5F8F8D4" baseAuthorityName="Reck" baseAuthorityYear="1956" box="[826,982,1775,1800]" class="Arachnida" family="Tetranychidae" genus="Petrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="species" species="shirakensis">
<emphasis id="4CA32526CA7DF3391308EA90D5F8F8D4" box="[826,982,1775,1800]" italics="true" pageId="10" pageNumber="331">G. shirakensis</emphasis>
</taxonomicName>
are also very close to African species of
<taxonomicName id="B9D782B7CA7DF33910A5EB73D739F8F0" authorityName="sensu Wainstein" authorityYear="1960" box="[151,279,1811,1836]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="4CA32526CA7DF33910A5EB73D739F8F0" box="[151,279,1811,1836]" italics="true" pageId="10" pageNumber="331">Georgiobia</emphasis>
</taxonomicName>
according to the medial propodosomal striation pattern (geometrical alveoli pattern present). Therefore we concluded that both leg setal counts and medial prodorsal striation are too consistent for species separation in this genus, but other morphological traits can distinguish species of
<taxonomicName id="B9D782B7CA7DF33913ABEB3BD237F8A8" authorityName="sensu Wainstein" authorityYear="1960" box="[921,1049,1883,1908]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="4CA32526CA7DF33913ABEB3BD237F8A8" box="[921,1049,1883,1908]" italics="true" pageId="10" pageNumber="331">Georgiobia</emphasis>
</taxonomicName>
: here, the body size, the length of the forelegs, the length and the relative lengths of tibial macrosetae III and IV, the lengths of dorsal setae, and the shape of the stylophore are distinctive.
</paragraph>
<caption id="2AA8A9BCCA7CF33810A5E825D424FBA5" httpUri="https://zenodo.org/record/2627018/files/figure.png" pageId="11" pageNumber="332" startId="11.[151,250,1093,1115]" targetBox="[150,1436,195,1071]" targetPageId="11">
<paragraph id="7E68F934CA7CF33810A5E825D424FBA5" blockId="11.[151,1437,1092,1145]" pageId="11" pageNumber="332">
<emphasis id="4CA32526CA7CF33810A5E825D721FB86" bold="true" box="[151,271,1093,1115]" pageId="11" pageNumber="332">FIGURE 5</emphasis>
.
<taxonomicName id="B9D782B7CA7CF338112CE824D7C0FB86" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[286,494,1092,1114]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7CF338112CE824D7C0FB86" box="[286,494,1092,1114]" italics="true" pageId="11" pageNumber="332">Georgiobia skuthiae</emphasis>
</taxonomicName>
<emphasis id="4CA32526CA7CF33811C5E825D41DFB86" bold="true" box="[503,563,1093,1114]" pageId="11" pageNumber="332">
<taxonomicNameLabel id="5790985DCA7CF33811C5E825D41DFB86" box="[503,563,1093,1114]" pageId="11" pageNumber="332" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Protonymph. A. dorsal view, B. detail of prodorsal striation, C. tubercles striation. Scale bars: 50 µm (A), 25 µm (B, C)
</paragraph>
</caption>
<paragraph id="7E68F934CA7CF33810F5E8A4D25EFA4B" blockId="11.[151,1437,1220,1755]" pageId="11" pageNumber="332">
Body and leg I are longer in
<taxonomicName id="B9D782B7CA7CF338123AE8A6D4ADFB01" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[520,643,1220,1245]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7CF338123AE8A6D4ADFB01" box="[520,643,1220,1245]" italics="true" pageId="11" pageNumber="332">G. skuthiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5790985DCA7CF33812B9E8A4D4E4FB01" box="[651,714,1220,1245]" pageId="11" pageNumber="332" rank="species">
<emphasis id="4CA32526CA7CF33812B9E8A4D4EDFB01" bold="true" box="[651,707,1220,1245]" pageId="11" pageNumber="332">n. sp</emphasis>
.
</taxonomicNameLabel>
compared to
<taxonomicName id="B9D782B7CA7CF3381351E8A6D5D1FB01" baseAuthorityName="Reck" baseAuthorityYear="1956" box="[867,1023,1220,1245]" class="Arachnida" family="Tetranychidae" genus="Petrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="shirakensis">
<emphasis id="4CA32526CA7CF3381351E8A6D5D1FB01" box="[867,1023,1220,1245]" italics="true" pageId="11" pageNumber="332">G. shirakensis</emphasis>
</taxonomicName>
. These species can also be separated by the length of their dorsal setae: except for sacrals (
<emphasis id="4CA32526CA7CF33812DEE889D4DAFADD" box="[748,756,1257,1281]" italics="true" pageId="11" pageNumber="332">f</emphasis>
<subScript id="E253FB71CA7CF33812C6E897D4D2FAD9" attach="left" box="[756,764,1271,1285]" fontSize="6" pageId="11" pageNumber="332">1</subScript>
and
<emphasis id="4CA32526CA7CF3381300E889D514FADD" box="[818,826,1257,1281]" italics="true" pageId="11" pageNumber="332">f</emphasis>
<subScript id="E253FB71CA7CF3381308E897D56CFAD9" attach="left" box="[826,834,1271,1285]" fontSize="6" pageId="11" pageNumber="332">2</subScript>
) and clunals (
<emphasis id="4CA32526CA7CF33813D2E888D5C0FADD" box="[992,1006,1256,1281]" italics="true" pageId="11" pageNumber="332">h</emphasis>
<subScript id="E253FB71CA7CF33813DCE897D5D8FAD9" attach="left" box="[1006,1014,1271,1285]" fontSize="6" pageId="11" pageNumber="332">1</subScript>
) which are subequal in length in both species, the dorsal body setae are obviously longer in
<taxonomicName id="B9D782B7CA7CF33812CBE970D558FAF4" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[761,886,1295,1320]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7CF33812CBE970D558FAF4" box="[761,886,1295,1320]" italics="true" pageId="11" pageNumber="332">G. skuthiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5790985DCA7CF338134CE96FD590FAFB" box="[894,958,1295,1320]" pageId="11" pageNumber="332" rank="species">
<emphasis id="4CA32526CA7CF338134CE96FD599FAF4" bold="true" box="[894,951,1295,1320]" pageId="11" pageNumber="332">n. sp</emphasis>
.
</taxonomicNameLabel>
(setae
<emphasis id="4CA32526CA7CF338143CE970D20BFAFB" box="[1038,1061,1296,1319]" italics="true" pageId="11" pageNumber="332">sc</emphasis>
<subScript id="E253FB71CA7CF3381414E97ED200FAF0" attach="left" box="[1062,1070,1310,1324]" fontSize="6" pageId="11" pageNumber="332">2</subScript>
more than ⅓ longer). Moreover, in
<taxonomicName id="B9D782B7CA7CF3381081E957D701FA93" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[179,303,1334,1359]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7CF3381081E957D701FA93" box="[179,303,1334,1359]" italics="true" pageId="11" pageNumber="332">G. skuthiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5790985DCA7CF3381104E956D75BFA92" box="[310,373,1334,1359]" pageId="11" pageNumber="332" rank="species">
<emphasis id="4CA32526CA7CF3381104E956D740FA93" bold="true" box="[310,366,1334,1359]" pageId="11" pageNumber="332">n. sp</emphasis>
.
</taxonomicNameLabel>
the stout macrosetae present on tibia III and IV are about twice as long as in
<taxonomicName id="B9D782B7CA7CF33814E1E957D35EFA93" baseAuthorityName="Reck" baseAuthorityYear="1956" box="[1235,1392,1334,1359]" class="Arachnida" family="Tetranychidae" genus="Petrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="shirakensis">
<emphasis id="4CA32526CA7CF33814E1E957D35EFA93" box="[1235,1392,1334,1359]" italics="true" pageId="11" pageNumber="332">G. shirakensis</emphasis>
</taxonomicName>
(80
<emphasis id="4CA32526CA7CF33810A5E93BD680FAAE" box="[151,174,1371,1394]" italics="true" pageId="11" pageNumber="332">vs</emphasis>
. 41); macrosetae are as long as tibia III and IV,
<emphasis id="4CA32526CA7CF33812C1E93BD524FAAE" box="[755,778,1371,1394]" italics="true" pageId="11" pageNumber="332">vs</emphasis>
. ½ and ¾ to ¼ the length, in
<taxonomicName id="B9D782B7CA7CF3381445E93BD2D9FAAF" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[1143,1271,1370,1395]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7CF3381445E93BD2D9FAAF" box="[1143,1271,1370,1395]" italics="true" pageId="11" pageNumber="332">G. skuthiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5790985DCA7CF3381530E93AD36BFAAE" box="[1282,1349,1370,1395]" pageId="11" pageNumber="332" rank="species">
<emphasis id="4CA32526CA7CF3381530E93AD310FAAF" bold="true" box="[1282,1342,1370,1395]" pageId="11" pageNumber="332">n. sp</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="B9D782B7CA7CF33815B6E93BD73BFA4B" baseAuthorityName="Reck" baseAuthorityYear="1956" class="Arachnida" family="Tetranychidae" genus="Petrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="shirakensis">
<emphasis id="4CA32526CA7CF33815B6E93BD3BBFAAE" box="[1412,1429,1371,1394]" italics="true" pageId="11" pageNumber="332">G</emphasis>
.
<emphasis id="4CA32526CA7CF33810A5E91ED73BFA4B" box="[151,277,1406,1431]" italics="true" pageId="11" pageNumber="332">shirakensis</emphasis>
</taxonomicName>
, respectively. Finally, the stylophore is emarginated only in the new species.
</paragraph>
<paragraph id="7E68F934CA7CF33810F5E9C2D449F907" blockId="11.[151,1437,1220,1755]" pageId="11" pageNumber="332">
The tarsal leg chaetotaxy of
<emphasis id="4CA32526CA7CF338122BE9C2D48EFA67" box="[537,672,1442,1467]" italics="true" pageId="11" pageNumber="332">
<taxonomicName id="B9D782B7CA7CF338122BE9C2D4B2FA67" authorityName="sensu Wainstein" authorityYear="1960" box="[537,668,1442,1467]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="genus">Georgiobia</taxonomicName>
,
</emphasis>
as seen in
<taxonomicName id="B9D782B7CA7CF338131EE9C3D582FA67" authority="Zhovnerchuk &amp; Auger, 2019" authorityName="Zhovnerchuk &amp; Auger" authorityYear="2019" box="[812,940,1442,1467]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="species" species="skuthiae" status="sp. nov.">
<emphasis id="4CA32526CA7CF338131EE9C3D582FA67" box="[812,940,1442,1467]" italics="true" pageId="11" pageNumber="332">G. skuthiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5790985DCA7CF3381385E9C2D5D7FA66" box="[951,1017,1442,1467]" pageId="11" pageNumber="332" rank="species">
<emphasis id="4CA32526CA7CF3381385E9C2D5DAFA67" bold="true" box="[951,1012,1442,1467]" pageId="11" pageNumber="332">n. sp</emphasis>
.
</taxonomicNameLabel>
, is singular for two reasons: 1) the fundamental tectal setae (
<emphasis id="4CA32526CA7CF3381185E9A7D7E5FA02" box="[439,459,1479,1502]" italics="true" pageId="11" pageNumber="332">tc</emphasis>
-
<emphasis id="4CA32526CA7CF33811ECE9A7D7DCFA02" box="[478,498,1479,1502]" italics="true" pageId="11" pageNumber="332">tc</emphasis>
) are absent on all the tarsi (
<figureCitation id="E6ECE5B1CA7CF338130EE9A6D5A4FA03" box="[828,906,1478,1503]" captionStart="FIGURE 3" captionStartId="9.[151,250,1975,1997]" captionTargetBox="[190,1400,196,1958]" captionTargetId="figure@9.[190,1400,193,1958]" captionTargetPageId="9" captionText="FIGURE 3. Georgiobia skuthiae n. sp., female. A. tarsus and tibia II, B. tarsus and tibia III, C. tarsus and tibia IV, D. palptarsus and palptibia, E. peritremal enlargement, F. ventral striation. Scale bars = 25 µm (AC), 10 µm (DE), 50 µm (F)." httpUri="https://zenodo.org/record/2627014/files/figure.png" pageId="11" pageNumber="332">Figs. 3</figureCitation>
AC). This
<typeStatus id="A16C4796CA7CF3381425E9A7D269FA03" box="[1047,1095,1479,1503]" pageId="11" pageNumber="332">type</typeStatus>
of setal reduction is not a real novelty as it is observed in the closely related genus
<taxonomicName id="B9D782B7CA7CF33812DAE98AD573F9DF" authorityName=", sensu Gutierrez" authorityYear="1985" box="[744,861,1514,1539]" class="Arachnida" family="Tetranychidae" genus="Aplonobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="4CA32526CA7CF33812DAE98AD573F9DF" box="[744,861,1514,1539]" italics="true" pageId="11" pageNumber="332">Aplonobia</emphasis>
</taxonomicName>
and was previously reported by
<bibRefCitation id="1A4684C5CA7CF33814FFE98AD3B2F9DF" author="Wainstein B. A." box="[1229,1436,1514,1539]" pageId="11" pageNumber="332" pagination="1476 - 1487" refId="ref10852" refString="Wainstein B. A. (1958 b) Chaetom of the extremities of the Tetranychidae (Acariformes) and the systematics of this family. Zoologicheskii Zhurnal, 37, 1476 - 1487. [in Russian]" type="journal article" year="1958">Wainstein (1958b)</bibRefCitation>
in three species of
<taxonomicName id="B9D782B7CA7CF3381154EA6ED7C8F9FB" authorityName="sensu Wainstein" authorityYear="1960" box="[358,486,1550,1575]" class="Arachnida" family="Tetranychidae" genus="Georgiobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="4CA32526CA7CF3381154EA6ED7C8F9FB" box="[358,486,1550,1575]" italics="true" pageId="11" pageNumber="332">Georgiobia</emphasis>
</taxonomicName>
but also in other genera like
<taxonomicName id="B9D782B7CA7CF3381315EA6ED595F9FB" baseAuthorityName="Wainstein" baseAuthorityYear="1956" box="[807,955,1550,1575]" class="Arachnida" family="Tetranychidae" genus="Mesobryobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="4CA32526CA7CF3381315EA6ED595F9FB" box="[807,955,1550,1575]" italics="true" pageId="11" pageNumber="332">Mesobryobia</emphasis>
</taxonomicName>
and
<taxonomicName id="B9D782B7CA7CF33813C2EA6ED2DCF9FB" authority="Murray, 1877" authorityName="Murray" authorityYear="1877" box="[1008,1266,1550,1575]" class="Arachnida" family="Tetranychidae" genus="Petrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="4CA32526CA7CF33813C2EA6ED27FF9FB" box="[1008,1105,1550,1575]" italics="true" pageId="11" pageNumber="332">Petrobia</emphasis>
<bibRefCitation id="1A4684C5CA7CF3381465EA6FD2DCF9FB" author="Murray, A." box="[1111,1266,1551,1575]" pageId="11" pageNumber="332" refId="ref10070" refString="Murray, A. (1877) Economic Entomology, Aptera. Chapman and Hall, London, 433 pp. https: // doi. org / 10.5962 / bhl. title. 51877" type="book" year="1877">Murray, 1877</bibRefCitation>
</taxonomicName>
; 2) in the distal part of the ventral side of tarsus I, one seta is missing and one is unpaired (
<figureCitation id="E6ECE5B1CA7CF33813CEEA52D24CF997" box="[1020,1122,1586,1611]" captionStart="FIGURE 2" captionStartId="8.[151,250,1971,1993]" captionTargetBox="[279,1436,238,1924]" captionTargetId="figure@8.[151,1437,238,1924]" captionTargetPageId="8" captionText="FIGURE 2. Georgiobia skuthiae n. sp., female. A. tarsus I, B. tibia I, C. empodium and claws I, D. empodium and claws II, E. coxisternal setae 1b, 1c (from left to right). Scale bars = 25 µm (AB), 10 µm (CE)." httpUri="https://zenodo.org/record/2627012/files/figure.png" pageId="11" pageNumber="332">Fig. 2-A</figureCitation>
). According to the possible location of this missing seta and by homology with the setal ontogeny of tarsi I and II, it seems that the setal reduction concerns a fundamental primiventral seta. Indeed, on the underside of tarsus I, the eupathidial seta usually not closely paired with
<emphasis id="4CA32526CA7CF33811CBEAFFD43DF96A" box="[505,531,1695,1718]" italics="true" pageId="11" pageNumber="332">pv</emphasis>
and located near the level of the duplex setae is lacking (
<figureCitation id="E6ECE5B1CA7CF338149DEAFED2DAF96B" box="[1199,1268,1694,1719]" captionStart="FIGURE 6" captionStartId="12.[151,250,755,777]" captionTargetBox="[166,1433,196,713]" captionTargetId="figure@12.[166,1433,191,738]" captionTargetPageId="12" captionText="FIGURE 6. Schematic representation of the underside of tarsi I and II showing setal insertions in the protonymphs, deutonymphs and adult female of Georgiobia skuthiae n. sp." httpUri="https://zenodo.org/record/2627020/files/figure.png" pageId="11" pageNumber="332">Fig. 6</figureCitation>
). We conclude that the missing seta must be the seta
<emphasis id="4CA32526CA7CF338120EEAA3D479F906" box="[572,599,1731,1754]" italics="true" pageId="11" pageNumber="332">pv</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>