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<document id="B46CEFBA2610464AA407FCB1576F9219" ID-DOI="10.5281/zenodo.193288" ID-GBIF-Dataset="17698376-7321-4993-b9b8-b1e3530ec632" ID-ISSN="1175-5326" ID-Zenodo-Dep="193288" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460130715195" checkinUser="plazi" docAuthor="Barej, Michael F., Rödel, Mark-Oliver, Gonwouo, Legrand Nono, Pauwels, Olivier S. G., Böhme, Wolfgang &amp; Schmitz, Andreas" docDate="2010" docId="B479090FFFDFFF83D7B4FA2943DF0EF0" docLanguage="en" docName="zt02340p049.pdf" docOrigin="Zootaxa 2340" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Petropedetes parkeri Amiet 1983" docType="treatment" docVersion="9" lastPageNumber="22" masterDocId="48407177FFCFFF96D723FFDB415F097C" masterDocTitle="Review of the genus Petropedetes Reichenow, 1874 in Central Africa with the description of three new species (Amphibia: Anura: Petropedetidae)" masterLastPageNumber="49" masterPageNumber="1" pageNumber="17" updateTime="1698235177974" updateUser="plazi">
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<mods:title id="BC6A0AEB3F44C84800F16496656BC78A">Review of the genus Petropedetes Reichenow, 1874 in Central Africa with the description of three new species (Amphibia: Anura: Petropedetidae)</mods:title>
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<mods:namePart id="E38B0F8124B8323719444A7238326A66">Barej, Michael F.</mods:namePart>
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<mods:namePart id="A11127C32D3DF60B75E45EBDF050EAAA">Rödel, Mark-Oliver</mods:namePart>
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<mods:namePart id="74570D6EF93F253054870C3746F1C954">Gonwouo, Legrand Nono</mods:namePart>
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<mods:namePart id="403B0D0598B75011A30436DCF5CB1690">Pauwels, Olivier S. G.</mods:namePart>
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<mods:namePart id="CE3833E554D21F785A0DF9143169FF35">Böhme, Wolfgang</mods:namePart>
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<mods:namePart id="E680EF56969D46EEBCE7E1E91074434A">Schmitz, Andreas</mods:namePart>
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<mods:date id="85E2E1DB5F1821C6C7CB0750558E6F6D">2010</mods:date>
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<treatment id="B479090FFFDFFF83D7B4FA2943DF0EF0" ID-DOI="http://doi.org/10.5281/zenodo.6199590" ID-GBIF-Taxon="124955649" ID-Zenodo-Dep="6199590" LSID="urn:lsid:plazi:treatment:B479090FFFDFFF83D7B4FA2943DF0EF0" httpUri="http://treatment.plazi.org/id/B479090FFFDFFF83D7B4FA2943DF0EF0" lastPageId="21" lastPageNumber="22" pageId="16" pageNumber="17">
<subSubSection id="74CAEB92FFDFFF86D7B4FA2940E70F53" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="3C6FB819FFDFFF86D7B4FA2943730F70" blockId="16.[151,556,1522,1583]" box="[151,556,1522,1548]" pageId="16" pageNumber="17">
<heading id="67270F75FFDFFF86D7B4FA2943730F70" bold="true" box="[151,556,1522,1548]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="FBD0C39AFFDFFF86D7B4FA2943730F70" ID-CoL="776CY" authority="Amiet, 1983" authorityName="Amiet" authorityYear="1983" box="[151,556,1522,1548]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDFFF86D7B4FA2943730F70" bold="true" box="[151,556,1522,1548]" pageId="16" pageNumber="17">
<emphasis id="0EA4640BFFDFFF86D7B4FA2940D00F70" bold="true" box="[151,399,1522,1548]" italics="true" pageId="16" pageNumber="17">Petropedetes parkeri</emphasis>
<bibRefCitation id="5841C5E8FFDFFF86D6B5FA2943730F70" author="Amiet" box="[406,556,1522,1548]" pageId="16" pageNumber="45" refString="Amiet, J. - L. (1983) Une espece meconnue de Petropedetes du Cameroun: Petropedetes parkeri n. sp. (Amphibia Anura: Ranidae, Phrynobatrachinae). Revue Suisse de Zoologie, 90, 457 - 468." type="journal article" year="1983">Amiet, 1983</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="3C6FB819FFDFFF86D7B4F9CE40E70F53" blockId="16.[151,556,1522,1583]" box="[151,440,1557,1583]" pageId="16" pageNumber="17">
<figureCitation id="A4EBA49CFFDFFF86D7B4F9CE41BB0F53" box="[151,228,1557,1583]" captionStart="FIGURE 1" captionStartId="4.[151,255,1795,1819]" captionTargetBox="[214,1373,207,1755]" captionTargetId="figure@4.[214,1374,207,1755]" captionTargetPageId="4" captionText="FIGURE 1. Life coloration of rudimentary-webbed Central African Petropedetes. a) Petropedetes johnstoni (male and female: Nkoelon, Cameroon), b) P. vulpiae sp. nov. (male and female: Nyasoso, Mt. Kupe, Cameroon), c) P. parkeri (male Nguti, Cameroon; female: Amebishu, Cameroon) d) P. euskircheni sp. nov. (male: Nyasoso, Mt. Kupe, Cameroon; female: Nguengue, Mt. Nlonako, Cameroon)." httpUri="https://zenodo.org/record/193289/files/figure.png" pageId="16" pageNumber="17">Figs. 1</figureCitation>
c, 2c, 4, 7a, 7b, 7f
</paragraph>
</subSubSection>
<subSubSection id="74CAEB92FFDFFF87D7B4F984429B08CA" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="3C6FB819FFDFFF86D7B4F98443C30EB5" blockId="16.[151,1437,1631,1993]" pageId="16" pageNumber="17">
<emphasis id="0EA4640BFFDFFF86D7B4F98440CE0F05" bold="true" box="[151,401,1631,1657]" pageId="16" pageNumber="17">Material examined.</emphasis>
BM 1936.3.4.124 (
<typeStatus id="E36B06BBFFDFFF86D55EF98443B60F05" box="[637,745,1631,1657]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
) adult male,
<collectingCountry id="44C7F889FFDFFF86D4AAF98445560F05" box="[905,1033,1631,1657]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
, Mamfe region, Atolo, Assumbo,
<date id="486E9ED9FFDFFF86D7B4F95E407F0FE3" box="[151,288,1669,1695]" pageId="16" pageNumber="17" value="1933-04-12">12.IV.1933</date>
, coll. I. Sanderson; BM 1936.3.4.123 (male),
<collectingCountry id="44C7F889FFDFFF86D474F95E42870FE3" box="[855,984,1669,1695]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
, Mamfe; BM 1936.3.4.127 (female), 1936.3.4.136 (male),
<collectingCountry id="44C7F889FFDFFF86D6B5F971434C0FB8" box="[406,531,1706,1732]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
, Bachor II; 1936.3.4.128 (female), 1936.3.4.131 (male),
<collectingCountry id="44C7F889FFDFFF86D397F971446D0FB8" box="[1204,1330,1706,1732]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
, Bashau,
<date id="486E9ED9FFDFFF86D7B4F914407E0F95" box="[151,289,1743,1769]" pageId="16" pageNumber="17" value="1932" valueMax="1933">1932-1933</date>
, coll. I. Sanderson;
<collectionCode id="5AC120DCFFDFFF86D502F91443200F95" box="[545,639,1743,1769]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="16" pageNumber="17">MHNG</collectionCode>
961.66 (male, formerly BM 1936.3.4.133),
<collectingCountry id="44C7F889FFDFFF86D393F914446A0F95" box="[1200,1333,1743,1769]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
, Mamfe Division, Bashau,
<date id="486E9ED9FFDFFF86D65EF92E43550E73" box="[381,522,1781,1807]" pageId="16" pageNumber="17" value="1933-03-10">10.III.1933</date>
, coll. I. Sanderson;
<collectionCode id="5AC120DCFFDFFF86D427F92E421D0E73" box="[772,834,1781,1807]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="16" pageNumber="17">PEM</collectionCode>
A 9123-9124 (
<specimenCount id="2AD67390FFDFFF86D323F92E45390E73" box="[1024,1126,1781,1807]" pageId="16" pageNumber="17" type="male">2 males</specimenCount>
), A 9126 (male), A 9127 (female),
<collectingCountry id="44C7F889FFDFFF86D636F8C140C30E48" box="[277,412,1818,1844]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
, Nguti,
<date id="486E9ED9FFDFFF86D527F8C143FC0E48" box="[516,675,1818,1844]" pageId="16" pageNumber="17" value="1996-07-15">15.VII.1996</date>
, coll. L. Minter;
<collectionCode id="5AC120DCFFDFFF86D4A2F8C1429E0E48" box="[897,961,1818,1844]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="16" pageNumber="17">PEM</collectionCode>
A 9125 (male), A 9128 (male),
<date id="486E9ED9FFDFFF86D240F8C144240E48" box="[1379,1403,1818,1844]" pageId="16" pageNumber="17" value="1996-07">5.</date>
&amp;
<date id="486E9ED9FFDFFF86D7B4F8E440440E25" box="[151,283,1855,1881]" pageId="16" pageNumber="17" value="1996-07-09">9.VII.1996</date>
, other data as A 9123;
<collectionCode id="5AC120DCFFDFFF86D50CF8E443DE0E25" LSID="urn:lsid:biocol.org:col:34613" box="[559,641,1855,1881]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
61337-338 (
<specimenCount id="2AD67390FFDFFF86D439F8E442CE0E25" box="[794,913,1855,1881]" pageId="16" pageNumber="17" type="female">2 females</specimenCount>
), 61342 (male),
<collectingCountry id="44C7F889FFDFFF86D376F8E4458C0E25" box="[1109,1235,1855,1881]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
, Ekundu-Kundu,
<date id="486E9ED9FFDFFF86D7B4F8BE40730E03" box="[151,300,1893,1919]" pageId="16" pageNumber="17" value="1989-02-03" valueMax="1989-02-27" valueMin="1989-02-03">3-27.II.1989</date>
, coll. M.T. OShea;
<collectionCode id="5AC120DCFFDFFF86D538F8BE43330E03" LSID="urn:lsid:biocol.org:col:34613" box="[539,620,1893,1919]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="16" pageNumber="17">ZFMK</collectionCode>
87702 (female),
<collectingCountry id="44C7F889FFDFFF86D415F8BE42EC0E03" box="[822,947,1893,1919]" name="Cameroon" pageId="16" pageNumber="17">Cameroon</collectingCountry>
, Mamfe region, near Amebishu, app.
<quantity id="FB2815FCFFDFFF86D253F8BE41F10ED8" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="16" pageNumber="17" unit="m" value="720.0">720 m</quantity>
a.s.l.,
<date id="486E9ED9FFDFFF86D7DCF85140D60ED8" box="[255,393,1930,1956]" pageId="16" pageNumber="17" value="2007-09-14">14.IX.2007</date>
, coll. J.A.M. Wurstner &amp; M.F. Barej;
<collectionCode id="5AC120DCFFDFFF86D475F85142C90ED8" box="[854,918,1930,1956]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="16" pageNumber="17">ZMB</collectionCode>
73739 (female),
<collectingCountry id="44C7F889FFDFFF86D344F851459B0ED8" box="[1127,1220,1930,1956]" name="Nigeria" pageId="16" pageNumber="17">Nigeria</collectingCountry>
, Okwangwo, app.
<quantity id="FB2815FCFFDFFF86D7B4F87441BE0EB5" box="[151,225,1967,1993]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="17" unit="m" value="900.0">900 m</quantity>
a.s.l.,
<date id="486E9ED9FFDFFF86D60EF87440EE0EB5" box="[301,433,1967,1993]" pageId="16" pageNumber="17" value="2004-04-15">15.IV.2004</date>
, coll. M. Gartshore.
</paragraph>
<paragraph id="3C6FB819FFDEFF87D7E7FF4C429B08CA" blockId="17.[151,1437,151,2038]" pageId="17" pageNumber="18">
<emphasis id="0EA4640BFFDEFF87D7E7FF4C40BC09CD" bold="true" box="[196,483,151,177]" pageId="17" pageNumber="18">Problematic material</emphasis>
(see systematic remarks). IRSNB-KBIN 13.842 (female),
<collectingCountry id="44C7F889FFDEFF87D3F3FF43447809CE" box="[1232,1319,152,178]" name="Gabon" pageId="17" pageNumber="18">Gabon</collectingCountry>
, Estuaire Province, Kango Department, 15 north of Alen-Nkoma, right of route L107, 2004, coll. O.S.G. Pauwels; IRSNB-KBIN 14.941 (female),
<collectingCountry id="44C7F889FFDEFF87D537FF39433A0980" box="[532,613,226,252]" name="Gabon" pageId="17" pageNumber="18">Gabon</collectingCountry>
, Province du Woleu-Ntem, Tchimbélé, app.
<quantity id="FB2815FCFFDEFF87D357FF39459F0980" box="[1140,1216,226,252]" metricMagnitude="2" metricUnit="m" metricValue="4.6899999999999995" pageId="17" pageNumber="18" unit="m" value="469.0">469 m</quantity>
a.s.l.,
<date id="486E9ED9FFDEFF87D22DFF3944C80980" box="[1294,1431,226,252]" pageId="17" pageNumber="18" value="2001-06-20">20.VI.2001</date>
, coll. O.S.G. Pauwels;
<collectionCode id="5AC120DCFFDEFF87D693FEDC4351085D" box="[432,526,263,289]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="17" pageNumber="18">MHNG</collectionCode>
1521.60 (male), 1521.61 (female), 1521.64-65 (
<specimenCount id="2AD67390FFDEFF87D359FEDC45A3085D" box="[1146,1276,263,289]" pageId="17" pageNumber="18" type="female">2 females</specimenCount>
),
<collectingCountry id="44C7F889FFDEFF87D231FEDC44C7085D" box="[1298,1432,263,289]" name="Cameroon" pageId="17" pageNumber="18">Cameroon</collectingCountry>
, Bafoussam, Bangwa,
<date id="486E9ED9FFDEFF87D6BEFEF7434A083A" box="[413,533,300,326]" pageId="17" pageNumber="18" value="1973-04-07">7.IV.1973</date>
, coll. J.-L. Perret;
<collectionCode id="5AC120DCFFDEFF87D5D4FEF7420D083A" box="[759,850,300,326]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="17" pageNumber="18">MHNG</collectionCode>
1521.70-71 (
<specimenCount id="2AD67390FFDEFF87D4D6FEF74508083A" box="[1013,1111,300,326]" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
),
<collectingCountry id="44C7F889FFDEFF87D34EFEF745B3083A" box="[1133,1260,300,326]" name="Cameroon" pageId="17" pageNumber="18">Cameroon</collectingCountry>
, Yaoundé, Mt. Kala,
<date id="486E9ED9FFDEFF87D7FDFE8940670810" box="[222,312,338,364]" pageId="17" pageNumber="18" value="1972-10">X.1972</date>
, coll. J.-L. Perret;
<collectionCode id="5AC120DCFFDEFF87D536FE8943300810" box="[533,623,338,364]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="17" pageNumber="18">MHNG</collectionCode>
1521.78 (male),
<collectingCountry id="44C7F889FFDEFF87D41FFE8942E50810" box="[828,954,338,364]" name="Cameroon" pageId="17" pageNumber="18">Cameroon</collectingCountry>
, Dschang, Falaise de Mbos,
<date id="486E9ED9FFDEFF87D22DFE8944C80810" box="[1294,1431,338,364]" pageId="17" pageNumber="18" value="1972-11-25">25.XI.1972</date>
, coll. J.-L. Perret;
<collectionCode id="5AC120DCFFDEFF87D644FEAC40E608ED" LSID="urn:lsid:biocol.org:col:34613" box="[359,441,375,401]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="17" pageNumber="18">ZFMK</collectionCode>
73209 (female), 73210 (male),
<collectingCountry id="44C7F889FFDEFF87D410FEAC42DB08ED" box="[819,900,375,401]" name="Gabon" pageId="17" pageNumber="18">Gabon</collectingCountry>
, Barrage de Tchimbélé, coll. V. Gossmann &amp; S. Lötters;
<collectionCode id="5AC120DCFFDEFF87D63BFE47400808CA" box="[280,343,412,438]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="17" pageNumber="18">ZMB</collectionCode>
73892 (female), same data as IRSNB-KBIN 14.941.
</paragraph>
</subSubSection>
<subSubSection id="74CAEB92FFDEFF87D7E7FE1A44D80B0C" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="3C6FB819FFDEFF87D7E7FE1A44D80B0C" blockId="17.[151,1437,151,2038]" pageId="17" pageNumber="18">
<emphasis id="0EA4640BFFDEFF87D7E7FE1A401908A7" bold="true" box="[196,326,449,475]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
Large sized
<taxonomicName id="FBD0C39AFFDEFF87D6C1FE1A432208A6" box="[482,637,449,474]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="0EA4640BFFDEFF87D6C1FE1A432208A6" box="[482,637,449,474]" italics="true" pageId="17" pageNumber="18">Petropedetes</emphasis>
</taxonomicName>
; robust body shape; tympanum round, larger than eye diameter in males, smaller in females; characters of breeding males: tympanic papilla present (broad and fleshy); papilla close to the upper border of the tympanum; forearm hypertrophy strongly developed in males; carpal spike present; tegumental spinosities well developed on throat, forearms, scattered on flanks and dorsum; femoral glands small, coloured similar to limb, shifted slightly to the posterior side of the leg; webbing rudimentary.
</paragraph>
</subSubSection>
<subSubSection id="74CAEB92FFDEFF87D7E7FDA043500E8A" pageId="17" pageNumber="18" type="description">
<paragraph id="3C6FB819FFDEFF87D7E7FDA045A20DBF" blockId="17.[151,1437,151,2038]" pageId="17" pageNumber="18">
<emphasis id="0EA4640BFFDEFF87D7E7FDA0403F0BE9" bold="true" box="[196,352,635,661]" pageId="17" pageNumber="18">Description.</emphasis>
Large sized
<taxonomicName id="FBD0C39AFFDEFF87D6DEFDA043C60BE8" box="[509,665,635,660]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="0EA4640BFFDEFF87D6DEFDA043C60BE8" box="[509,665,635,660]" italics="true" pageId="17" pageNumber="18">Petropedetes</emphasis>
</taxonomicName>
with robust body; males distinctly larger than females (SUL in males: 38.0
<quantity id="FB2815FCFFDEFF87D60DFD7A40C50BC7" box="[302,410,673,699]" metricMagnitude="-2" metricUnit="m" metricValue="7.43" pageId="17" pageNumber="18" unit="mm" value="74.3">74.3 mm</quantity>
, in females:
<quantity id="FB2815FCFFDEFF87D512FD7A43BF0BC7" box="[561,736,673,699]" metricMagnitude="-2" metricUnit="m" metricValue="4.77" metricValueMax="6.13" metricValueMin="3.41" pageId="17" pageNumber="18" unit="mm" value="47.7" valueMax="61.3" valueMin="34.1">34.161.3 mm</quantity>
); mean head width in males about 44% of SUL, in females about 42%; snout in lateral view short, rounded;
<emphasis id="0EA4640BFFDEFF87D5F8FD1D42FE0BA3" box="[731,929,710,735]" italics="true" pageId="17" pageNumber="18">canthus rostralis</emphasis>
slightly rounded but distinct; loreal region concave; eye diameter about 1.3 times eye-narial distance; nose closer to snout-tip than to eye; distinct tympanum, usually larger than eye in males, always smaller in females (tympanum / eye in males: 0.73-1.39, in females: 0.600.89; the lowest value of
<quantity id="FB2815FCFFDEFF87D5B3FCEE43BC0A33" box="[656,739,821,847]" metricMagnitude="-2" metricUnit="m" metricValue="1.8542" pageId="17" pageNumber="18" unit="in" value="0.73">0.73 in</quantity>
males measured in the smallest specimen, second smallest value: 0.92; tympanic papilla closer to upper border than centre of tympanum; papilla broad and fleshy with oval basis; supratympanic fold distinct; fingers slender, with typically T-shaped fingertips; relative length of fingers: III&gt; IV&gt; II&gt; I; manual subarticular tubercles single; palmar tubercle and thenar tubercle present, palmar tubercle sometimes indistinct; forearm hypertrophy strongly developed in males; carpal spike present in males; tegumental spinosities well developed on throat, forearms, scattered on flanks and dorsum in males; dorsal tegument with smaller warts than flanks; few larger, elongated warts on dorsum; ventral skin smooth; femora long, mean femur length in males 50% of SUL, in females 51%; mean foot length in both sexes 74% of SUL; upper hind limbs of moderate width, lower hind limbs slender; femoral glands small in both sexes, slightly bigger in males (femoral gland / femur in males: 0.180.27, in females: 0.160.22); relative length of toes: IV&gt; III&gt; V&gt; II&gt; I; webbing rudimentary: 1 (1) 2 (1-1) or 2 (1.25-1) 3 (2-2) 4 (3-3) 5 (2).
</paragraph>
<paragraph id="3C6FB819FFDEFF87D7E7FB1543960CB4" blockId="17.[151,1437,151,2038]" pageId="17" pageNumber="18">
<emphasis id="0EA4640BFFDEFF87D7E7FB15400D0D94" bold="true" box="[196,338,1230,1256]" pageId="17" pageNumber="18">Coloration.</emphasis>
Dorsum and flanks olive or brownish and dark marbled with diffuse brown-olive spots (
<figureCitation id="A4EBA49CFFDEFF87D24FFB1541FC0C72" captionStart="FIGURE 1" captionStartId="4.[151,255,1795,1819]" captionTargetBox="[214,1373,207,1755]" captionTargetId="figure@4.[214,1374,207,1755]" captionTargetPageId="4" captionText="FIGURE 1. Life coloration of rudimentary-webbed Central African Petropedetes. a) Petropedetes johnstoni (male and female: Nkoelon, Cameroon), b) P. vulpiae sp. nov. (male and female: Nyasoso, Mt. Kupe, Cameroon), c) P. parkeri (male Nguti, Cameroon; female: Amebishu, Cameroon) d) P. euskircheni sp. nov. (male: Nyasoso, Mt. Kupe, Cameroon; female: Nguengue, Mt. Nlonako, Cameroon)." httpUri="https://zenodo.org/record/193289/files/figure.png" pageId="17" pageNumber="18">Fig. 1</figureCitation>
c); throat dirty whitish; belly whitish, slightly translucent; throat can be darker than belly; femora and lower legs with large darker spots, darker spots divided by thin bright coloured transversal bars; ventral surfaces of limbs of the same coloration as belly, pale greenish; iris golden with white-green shades; femoral glands coloured as hind limbs, pale orange or greenish. Coloration in preservation: similar as in life; dorsum uniform brown; belly whitish, with minuscule speckles (less dense than in former species, only recognizable under microscope); throat dirty whitish or pale brown.
</paragraph>
<paragraph id="3C6FB819FFDEFF87D7E7FA0942C00E6A" blockId="17.[151,1437,151,2038]" pageId="17" pageNumber="18">
<emphasis id="0EA4640BFFDEFF87D7E7FA0940D30C90" bold="true" box="[196,396,1490,1516]" pageId="17" pageNumber="18">Natural history.</emphasis>
After the first work on the ecology of the species by
<bibRefCitation id="5841C5E8FFDEFF87D329FA08404F0F6E" author="Sanderson" pageId="17" pageNumber="47" refString="Sanderson, I. T. (1936) The amphibians of the Mamfe Division, Cameroon. - II. Ecology of the frogs. Proceedings of the Zoological Society of London, 165 - 208." type="proceedings paper" year="1936">
Sanderson (1936, referred to as
<taxonomicName id="FBD0C39AFFDEFF87D2A5FA08405A0F6D" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="johnstoni">
<emphasis id="0EA4640BFFDEFF87D2A5FA08405A0F6D" italics="true" pageId="17" pageNumber="18">P. johnstoni</emphasis>
</taxonomicName>
)
</bibRefCitation>
, additional data have been gathered by Amiet. However, some observations by
<bibRefCitation id="5841C5E8FFDEFF87D3E9FA2344C80F6E" author="Amiet" box="[1226,1431,1528,1554]" pageId="17" pageNumber="44" refString="Amiet, J. - L. (1975) Ecologie et distribution des Amphibiens Anoures de la region de Nkongsamba. Annales de la Faculte des Sciences de Yaounde, 20, 33 - 107." type="journal article" year="1975">Amiet (e.g. 1975</bibRefCitation>
,
<bibRefCitation id="5841C5E8FFDEFF87D7B4F9C6418A0F4B" author="Amiet" box="[151,213,1565,1591]" pageId="17" pageNumber="45" refString="Amiet, J. - L. (1983) Une espece meconnue de Petropedetes du Cameroun: Petropedetes parkeri n. sp. (Amphibia Anura: Ranidae, Phrynobatrachinae). Revue Suisse de Zoologie, 90, 457 - 468." type="journal article" year="1983">1983</bibRefCitation>
,
<bibRefCitation id="5841C5E8FFDEFF87D7C2F9C640420F4B" author="Amiet" box="[225,285,1565,1591]" pageId="17" pageNumber="45" refString="Amiet, J. - L. (1986) La batrachofaune sylvicole d'un secteur forestier du Cameroun: la region de Yaounde. Memoires du Museum National d'histoire Naturelle, 132, 29 - 42." type="journal article" year="1986">1986</bibRefCitation>
) are based on populations of “
<taxonomicName id="FBD0C39AFFDEFF87D5A9F9C543A10F4B" box="[650,766,1566,1591]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDEFF87D5A9F9C543A10F4B" box="[650,766,1566,1591]" italics="true" pageId="17" pageNumber="18">P. parkeri</emphasis>
</taxonomicName>
” from Mt. Kala, the Bamiléké Plateau and the region of Nkongsamba. The taxonomic status of these populations at present is not clear (see below).
<bibRefCitation id="5841C5E8FFDEFF87D3EFF99944C40F20" author="Sanderson" box="[1228,1435,1602,1628]" pageId="17" pageNumber="47" refString="Sanderson, I. T. (1936) The amphibians of the Mamfe Division, Cameroon. - II. Ecology of the frogs. Proceedings of the Zoological Society of London, 165 - 208." type="proceedings paper" year="1936">Sanderson (1936)</bibRefCitation>
calls this species arboreal, because he found them on lower, broadened leafs, in shade. The breeding season is said to take place in the rainy season (
<bibRefCitation id="5841C5E8FFDEFF87D571F95645420FDA" author="Parker" box="[594,1053,1677,1703]" pageId="17" pageNumber="47" refString="Parker, H. W. (1936) The amphibians of the Mamfe Division, Cameroons. - I. Zoogeography and systematics. Proceedings of the Zoological Society of London, 135 - 163." type="proceedings paper" year="1936">
Parker 1936; referred to as
<taxonomicName id="FBD0C39AFFDEFF87D4B0F95645420FDA" box="[915,1053,1677,1702]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="johnstoni">
<emphasis id="0EA4640BFFDEFF87D4B0F95645420FDA" box="[915,1053,1677,1702]" italics="true" pageId="17" pageNumber="18">P. johnstoni</emphasis>
</taxonomicName>
</bibRefCitation>
). The species has been found on wet, mossy rocks in the forest; the large rocks were crossed by runlets (
<figureCitation id="A4EBA49CFFDEFF87D326F969450F0FB0" box="[1029,1104,1714,1740]" captionStart="FIGURE 7" captionStartId="18.[151,259,1746,1770]" captionTargetBox="[215,1371,183,1725]" captionTargetId="figure@18.[215,1372,183,1725]" captionTargetPageId="18" captionText="FIGURE 7. Habitat pictures and specimens of uncertain determination: a) Petropedetes parkeri, Mamfe region, Cameroon; b) P. vulpiae sp. nov. biotope, waterfall at Itsila, Gabon Republic of Congo border; c) P. vulpiae sp. nov., cascade 15 km north of Alen-Nkoma, right of Route L. 107, Kango Department, Estuaire Province, Gabon (photo: J. P. Vande weghe); d) type locality of P. juliawurstnerae sp. nov. and P. euskircheni sp. nov., habitat of P. vulpiae sp. nov., Nyasoso, Mt. Kupe, Cameroon; e) P. vulpiae sp. nov., cascade at Big Massaka, Cameroon; f) P. aff. parkeri, Barrage de Tchimbélé, Gabon (ZFMK 73132; photo: V. Gossmann); g) P. cf. johnstoni near Langoué Bai, Gabon (no voucher; photo: J. P. Vande weghe):" httpUri="https://zenodo.org/record/193295/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
a). According to
<bibRefCitation id="5841C5E8FFDEFF87D23DF96941BB0F8D" author="Sanderson" pageId="17" pageNumber="47" refString="Sanderson, I. T. (1936) The amphibians of the Mamfe Division, Cameroon. - II. Ecology of the frogs. Proceedings of the Zoological Society of London, 165 - 208." type="proceedings paper" year="1936">Sanderson (1936)</bibRefCitation>
the adults aggregate in the breeding season on humid rocky surfaces in rough areas with torrent water, while they live in the forest on leaves outside the breeding season.
</paragraph>
<paragraph id="3C6FB819FFDEFF87D7E7F8F943500E8A" blockId="17.[151,1437,151,2038]" pageId="17" pageNumber="18">
The advertisement call of
<emphasis id="0EA4640BFFDEFF87D52BF8F943210E47" box="[520,638,1826,1851]" italics="true" pageId="17" pageNumber="18">P. p a r k e r i</emphasis>
has been published by
<bibRefCitation id="5841C5E8FFDEFF87D486F8F945C30E40" box="[933,1180,1826,1852]" pageId="17" pageNumber="47" refString="Narins, P. M., Lewis, E. R., Purgue, A. P., Bishop, P. J., Minter, L. R. &amp; Lawson, D. P. (2001) Functional consequences of a novel middle ear adaptation in the Central African frog Petropedetes parkeri (Ranidae). The Journal of Experimental Biology, 204, 1223 - 1232." type="journal article">
Narins
<emphasis id="0EA4640BFFDEFF87D4DCF8F945690E47" box="[1023,1078,1826,1851]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (2001)
</bibRefCitation>
.
<bibRefCitation id="5841C5E8FFDEFF87D38AF8F944C20E40" box="[1193,1437,1826,1852]" pageId="17" pageNumber="47" refString="Narins, P. M., Lewis, E. R., Purgue, A. P., Bishop, P. J., Minter, L. R. &amp; Lawson, D. P. (2001) Functional consequences of a novel middle ear adaptation in the Central African frog Petropedetes parkeri (Ranidae). The Journal of Experimental Biology, 204, 1223 - 1232." type="journal article">
Narins
<emphasis id="0EA4640BFFDEFF87D220F8F944650E47" box="[1283,1338,1826,1851]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (2001)
</bibRefCitation>
describe a part of the courtship behaviour, including the female striking the males head with her foreleg during amplexus. These authors speculate that this behaviour is connected with secretion of glands in the male tympanal papilla. The small eggs are deposited at the beginning of the rainy season on surfaces of stones within the splash zone. Metamorphosis in
<taxonomicName id="FBD0C39AFFDEFF87D5ABF86C43A40EAC" box="[648,763,1975,2000]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDEFF87D5ABF86C43A40EAC" box="[648,763,1975,2000]" italics="true" pageId="17" pageNumber="18">P. parkeri</emphasis>
</taxonomicName>
is finished before the end of the rains or at the beginning of the dry season (
<bibRefCitation id="5841C5E8FFDEFF87D64EF807435D0E8A" author="Parker" box="[365,514,2012,2038]" pageId="17" pageNumber="47" refString="Parker, H. W. (1936) The amphibians of the Mamfe Division, Cameroons. - I. Zoogeography and systematics. Proceedings of the Zoological Society of London, 135 - 163." type="proceedings paper" year="1936">Parker 1936</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="68AFE891FFDDFF84D7B4F90940F30ED6" httpUri="https://zenodo.org/record/193295/files/figure.png" pageId="18" pageNumber="19" targetBox="[215,1371,183,1725]" targetPageId="18">
<paragraph id="3C6FB819FFDDFF84D7B4F90940F30ED6" blockId="18.[151,1436,1746,1962]" pageId="18" pageNumber="19">
<emphasis id="0EA4640BFFDDFF84D7B4F909407E0F96" bold="true" box="[151,289,1746,1770]" pageId="18" pageNumber="19">FIGURE 7.</emphasis>
Habitat pictures and specimens of uncertain determination: a)
<taxonomicName id="FBD0C39AFFDDFF84D4D6F908441A0F96" authority="Mamfe" authorityName="Mamfe" box="[1013,1349,1747,1770]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDDFF84D4D6F90845B40F96" box="[1013,1259,1747,1770]" italics="true" pageId="18" pageNumber="19">Petropedetes parkeri,</emphasis>
Mamfe
</taxonomicName>
region, Cameroon; b)
<taxonomicName id="FBD0C39AFFDDFF84D610F92840C30E76" box="[307,412,1779,1802]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="vulpiae" status="sp. nov.">
<emphasis id="0EA4640BFFDDFF84D610F92840C30E76" box="[307,412,1779,1802]" italics="true" pageId="18" pageNumber="19">P. vulpiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDDFF84D687F92940A40E76" box="[420,507,1778,1802]" pageId="18" pageNumber="19" rank="species">
<emphasis id="0EA4640BFFDDFF84D687F92940A40E76" bold="true" box="[420,507,1778,1802]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
biotope, waterfall at Itsila, Gabon Republic of Congo border; c)
<taxonomicName id="FBD0C39AFFDDFF84D3EDF92844670E76" box="[1230,1336,1779,1802]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="vulpiae" status="sp. nov.">
<emphasis id="0EA4640BFFDDFF84D3EDF92844670E76" box="[1230,1336,1779,1802]" italics="true" pageId="18" pageNumber="19">P. vulpiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDDFF84D21CF92944CA0E76" box="[1343,1429,1778,1802]" pageId="18" pageNumber="19" rank="species">
<emphasis id="0EA4640BFFDDFF84D21CF92944CA0E76" bold="true" box="[1343,1429,1778,1802]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
, cascade 15 km north of Alen-Nkoma, right of Route L. 107, Kango Department, Estuaire Province, Gabon (photo: J.P. Vande weghe); d) type locality of
<taxonomicName id="FBD0C39AFFDDFF84D524F8E843940E36" box="[519,715,1843,1866]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="juliawurstnerae" status="sp. nov.">
<emphasis id="0EA4640BFFDDFF84D524F8E843940E36" box="[519,715,1843,1866]" italics="true" pageId="18" pageNumber="19">P. juliawurstnerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDDFF84D5F0F8E942760E36" box="[723,809,1842,1866]" pageId="18" pageNumber="19" rank="species">
<emphasis id="0EA4640BFFDDFF84D5F0F8E942760E36" bold="true" box="[723,809,1842,1866]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="FBD0C39AFFDDFF84D47CF8E842AA0E36" box="[863,1013,1843,1866]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="euskircheni" status="sp. nov.">
<emphasis id="0EA4640BFFDDFF84D47CF8E842AA0E36" box="[863,1013,1843,1866]" italics="true" pageId="18" pageNumber="19">P. euskircheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDDFF84D4DFF8E9450D0E36" box="[1020,1106,1842,1866]" pageId="18" pageNumber="19" rank="species">
<emphasis id="0EA4640BFFDDFF84D4DFF8E9450D0E36" bold="true" box="[1020,1106,1842,1866]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
, habitat of
<taxonomicName id="FBD0C39AFFDDFF84D3EDF8E844670E36" box="[1230,1336,1843,1866]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="vulpiae" status="sp. nov.">
<emphasis id="0EA4640BFFDDFF84D3EDF8E844670E36" box="[1230,1336,1843,1866]" italics="true" pageId="18" pageNumber="19">P. vulpiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDDFF84D21CF8E944CA0E36" box="[1343,1429,1842,1866]" pageId="18" pageNumber="19" rank="species">
<emphasis id="0EA4640BFFDDFF84D21CF8E944CA0E36" bold="true" box="[1343,1429,1842,1866]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
, Nyasoso, Mt. Kupe, Cameroon; e)
<taxonomicName id="FBD0C39AFFDDFF84D52DF88843280E16" box="[526,631,1875,1898]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="vulpiae" status="sp. nov.">
<emphasis id="0EA4640BFFDDFF84D52DF88843280E16" box="[526,631,1875,1898]" italics="true" pageId="18" pageNumber="19">P. vulpiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDDFF84D55DF889438B0E16" box="[638,724,1874,1898]" pageId="18" pageNumber="19" rank="species">
<emphasis id="0EA4640BFFDDFF84D55DF889438B0E16" bold="true" box="[638,724,1874,1898]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
, cascade at Big Massaka, Cameroon; f)
<taxonomicName id="FBD0C39AFFDDFF84D3A6F88844240E16" authority="Barrage" authorityName="Barrage" box="[1157,1403,1875,1898]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDDFF84D3A6F88845CD0E16" box="[1157,1170,1875,1898]" italics="true" pageId="18" pageNumber="19">P</emphasis>
. aff.
<emphasis id="0EA4640BFFDDFF84D3E8F88844460E16" box="[1227,1305,1875,1898]" italics="true" pageId="18" pageNumber="19">parkeri</emphasis>
, Barrage
</taxonomicName>
de Tchimbélé, Gabon (ZFMK 73132; photo: V. Gossmann); g)
<taxonomicName id="FBD0C39AFFDDFF84D414F8A842BF0EF6" box="[823,992,1907,1930]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="johnstoni">
<emphasis id="0EA4640BFFDDFF84D414F8A842160EF6" box="[823,841,1907,1930]" italics="true" pageId="18" pageNumber="19">P.</emphasis>
cf.
<emphasis id="0EA4640BFFDDFF84D45BF8A842BF0EF6" box="[888,992,1907,1930]" italics="true" pageId="18" pageNumber="19">johnstoni</emphasis>
</taxonomicName>
near Langoué Bai, Gabon (no voucher; photo: J.P. Vande weghe):
</paragraph>
</caption>
<subSubSection id="74CAEB92FFDAFF83D7E7FF43428308CB" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="3C6FB819FFDAFF83D7E7FF43428308CB" blockId="21.[151,1437,152,1932]" pageId="21" pageNumber="22">
<taxonomicName id="FBD0C39AFFDAFF83D7E7FF4340E609CD" ID-CoL="776CY" box="[196,441,152,177]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D7E7FF4340E609CD" box="[196,441,152,177]" italics="true" pageId="21" pageNumber="22">Petropedetes parkeri</emphasis>
</taxonomicName>
shows, in contrast to other members of the genus, a reversed sexual size dimorphism (
<bibRefCitation id="5841C5E8FFDAFF83D7BCFF66406209AB" author="Amiet" box="[159,317,189,215]" pageId="21" pageNumber="45" refString="Amiet, J. - L. (1983) Une espece meconnue de Petropedetes du Cameroun: Petropedetes parkeri n. sp. (Amphibia Anura: Ranidae, Phrynobatrachinae). Revue Suisse de Zoologie, 90, 457 - 468." type="journal article" year="1983">Amiet 1983</bibRefCitation>
). The state that males grow larger than females is rare and occurs only in about 10% of amphibians (
<bibRefCitation id="5841C5E8FFDAFF83D611FF39409E0980" author="Shine" box="[306,449,226,252]" pageId="21" pageNumber="47" refString="Shine, R. (1979) Sexual selection and sexual dimorphism in the amphibia. Copeia, 1979, 297 - 306." type="journal article" year="1979">Shine 1979</bibRefCitation>
). Most often this is connected to some kind of territoriality or male parental care. However,
<bibRefCitation id="5841C5E8FFDAFF83D633FED34080085E" author="Sanderson" box="[272,479,264,290]" pageId="21" pageNumber="47" refString="Sanderson, I. T. (1936) The amphibians of the Mamfe Division, Cameroon. - II. Ecology of the frogs. Proceedings of the Zoological Society of London, 165 - 208." type="proceedings paper" year="1936">Sanderson (1936)</bibRefCitation>
notes the laceration of non-breeding males in the gular region and assumed relics of the previous breeding seasons. This could be regarded as an outcome of territorial behaviour. Further respective data are however lacking. This species appears to live in forests. Some specimens of
<taxonomicName id="FBD0C39AFFDAFF83D3D9FE8944C40817" ID-CoL="776CY" box="[1274,1435,338,364]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D3D9FE8944560817" box="[1274,1289,338,363]" italics="true" pageId="21" pageNumber="22">P</emphasis>
. aff.
<emphasis id="0EA4640BFFDAFF83D266FE8944C40817" box="[1349,1435,338,363]" italics="true" pageId="21" pageNumber="22">parkeri</emphasis>
</taxonomicName>
from
<collectingCountry id="44C7F889FFDAFF83D7F4FEA3407A08EE" box="[215,293,376,402]" name="Gabon" pageId="21" pageNumber="22">Gabon</collectingCountry>
(see below,
<figureCitation id="A4EBA49CFFDAFF83D694FEA340A408EE" box="[439,507,376,402]" captionStart="FIGURE 7" captionStartId="18.[151,259,1746,1770]" captionTargetBox="[215,1371,183,1725]" captionTargetId="figure@18.[215,1372,183,1725]" captionTargetPageId="18" captionText="FIGURE 7. Habitat pictures and specimens of uncertain determination: a) Petropedetes parkeri, Mamfe region, Cameroon; b) P. vulpiae sp. nov. biotope, waterfall at Itsila, Gabon Republic of Congo border; c) P. vulpiae sp. nov., cascade 15 km north of Alen-Nkoma, right of Route L. 107, Kango Department, Estuaire Province, Gabon (photo: J. P. Vande weghe); d) type locality of P. juliawurstnerae sp. nov. and P. euskircheni sp. nov., habitat of P. vulpiae sp. nov., Nyasoso, Mt. Kupe, Cameroon; e) P. vulpiae sp. nov., cascade at Big Massaka, Cameroon; f) P. aff. parkeri, Barrage de Tchimbélé, Gabon (ZFMK 73132; photo: V. Gossmann); g) P. cf. johnstoni near Langoué Bai, Gabon (no voucher; photo: J. P. Vande weghe):" httpUri="https://zenodo.org/record/193295/files/figure.png" pageId="21" pageNumber="22">Fig. 7</figureCitation>
f) have been collected during the day in a gallery forest.
<taxonomicName id="FBD0C39AFFDAFF83D3B1FEA3446C08ED" ID-CoL="776CY" box="[1170,1331,376,402]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D3B1FEA345FE08ED" box="[1170,1185,376,401]" italics="true" pageId="21" pageNumber="22">P</emphasis>
. aff.
<emphasis id="0EA4640BFFDAFF83D3FEFEA3446C08ED" box="[1245,1331,376,401]" italics="true" pageId="21" pageNumber="22">parkeri</emphasis>
</taxonomicName>
is called “barking frog” by locals in western
<collectingCountry id="44C7F889FFDAFF83D519FE4643D708CB" box="[570,648,413,439]" name="Gabon" pageId="21" pageNumber="22">Gabon</collectingCountry>
(V. Gossmann
<emphasis id="0EA4640BFFDAFF83D41EFE46429408CA" box="[829,971,413,438]" italics="true" pageId="21" pageNumber="22">pers. comm.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="74CAEB92FFDAFF83D7E7FE1945110BEB" pageId="21" pageNumber="22" type="distribution">
<paragraph id="3C6FB819FFDAFF83D7E7FE1945110B7E" blockId="21.[151,1437,152,1932]" pageId="21" pageNumber="22">
<emphasis id="0EA4640BFFDAFF83D7E7FE19403908A0" bold="true" box="[196,358,450,476]" pageId="21" pageNumber="22">Distribution.</emphasis>
We can only assign populations from western
<collectingCountry id="44C7F889FFDAFF83D4B6FE19454F08A0" box="[917,1040,450,476]" name="Cameroon" pageId="21" pageNumber="22">Cameroon</collectingCountry>
and eastern
<collectingCountry id="44C7F889FFDAFF83D384FE19445F08A0" box="[1191,1280,450,476]" name="Nigeria" pageId="21" pageNumber="22">Nigeria</collectingCountry>
to
<taxonomicName id="FBD0C39AFFDAFF83D204FE1944C308A7" box="[1319,1436,450,475]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D204FE1944C308A7" box="[1319,1436,450,475]" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
with certainty (
<figureCitation id="A4EBA49CFFDAFF83D66AFE3340CD0B7E" box="[329,402,488,514]" captionStart="FIGURE 4" captionStartId="8.[151,255,1966,1990]" captionTargetBox="[212,1373,567,1944]" captionTargetId="figure@8.[212,1374,567,1944]" captionTargetPageId="8" captionText="FIGURE 4. Distribution of known Petropedetes species; compare with text. Specimens from localities marked with an asterisk (*) were not available and are added from literature." httpUri="https://zenodo.org/record/193292/files/figure.png" pageId="21" pageNumber="22">Fig. 4</figureCitation>
). At these locations species inhabit altitudes below
<quantity id="FB2815FCFFDAFF83D4CDFE3345150B7E" box="[1006,1098,488,514]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="m" value="1000.0">1000 m</quantity>
.
</paragraph>
<paragraph id="3C6FB819FFDAFF83D7E7FDD645110BEB" blockId="21.[151,1437,152,1932]" pageId="21" pageNumber="22">
<emphasis id="0EA4640BFFDAFF83D7E7FDD640620B5B" bold="true" box="[196,317,525,551]" pageId="21" pageNumber="22">Genetics.</emphasis>
The genetic comparison for the uncorrected p-distances of the mitochondrial 16S rRNAfragment (Table 4) gave the following results for
<taxonomicName id="FBD0C39AFFDAFF83D5C5FDE942810B37" box="[742,990,562,587]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D5C5FDE942810B37" box="[742,990,562,587]" italics="true" pageId="21" pageNumber="22">Petropedetes parkeri</emphasis>
</taxonomicName>
: interspecific comparison between
<taxonomicName id="FBD0C39AFFDAFF83D2A5FDE941A80B0D" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D2A5FDE941A80B0D" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
and all other Central African
<taxonomicName id="FBD0C39AFFDAFF83D5A4FD83426F0B0D" box="[647,816,600,625]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="0EA4640BFFDAFF83D5A4FD83426F0B0D" box="[647,816,600,625]" italics="true" pageId="21" pageNumber="22">Petropedetes</emphasis>
</taxonomicName>
taxa ranged between 5.94%-12.39%, while the intraspecific variation within
<emphasis id="0EA4640BFFDAFF83D6D2FDA6433F0BEA" box="[497,608,637,662]" italics="true" pageId="21" pageNumber="22">P. p a r k e r i</emphasis>
was much lower at 0.00%-0.22% (N= 4).
</paragraph>
</subSubSection>
<subSubSection id="74CAEB92FFDAFF83D7E7FD7943710E60" pageId="21" pageNumber="22" type="description">
<paragraph id="3C6FB819FFDAFF83D7E7FD79420F0D00" blockId="21.[151,1437,152,1932]" pageId="21" pageNumber="22">
<emphasis id="0EA4640BFFDAFF83D7E7FD79409A0BC0" bold="true" box="[196,453,674,700]" pageId="21" pageNumber="22">Systematic remarks.</emphasis>
<taxonomicName id="FBD0C39AFFDAFF83D6EEFD7943990BC7" box="[461,710,674,699]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D6EEFD7943990BC7" box="[461,710,674,699]" italics="true" pageId="21" pageNumber="22">Petropedetes parkeri</emphasis>
</taxonomicName>
has long been confused with “
<taxonomicName id="FBD0C39AFFDAFF83D315FD7945E60BC7" box="[1078,1209,674,699]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="newtonii">
<emphasis id="0EA4640BFFDAFF83D315FD7945E60BC7" box="[1078,1209,674,699]" italics="true" pageId="21" pageNumber="22">P. newtonii</emphasis>
</taxonomicName>
” (described herein as
<emphasis id="0EA4640BFFDAFF83D795FD1340790B9D" box="[182,294,712,737]" italics="true" pageId="21" pageNumber="22">P. v u l p i a e</emphasis>
<emphasis id="0EA4640BFFDAFF83D612FD1C40D10B9D" bold="true" box="[305,398,711,737]" pageId="21" pageNumber="22">sp. nov.</emphasis>
) and
<taxonomicName id="FBD0C39AFFDAFF83D6F2FD1342700B9E" authority="Gartshore 1994" authorityName="Gartshore" authorityYear="1994" box="[465,815,712,738]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="johnstoni">
<emphasis id="0EA4640BFFDAFF83D6F2FD1343040B9D" box="[465,603,712,737]" italics="true" pageId="21" pageNumber="22">P. johnstoni</emphasis>
(Gartshore 1994)
</taxonomicName>
.
<bibRefCitation id="5841C5E8FFDAFF83D419FD1342860B9E" author="Amiet" box="[826,985,712,738]" pageId="21" pageNumber="45" refString="Amiet, J. - L. (1983) Une espece meconnue de Petropedetes du Cameroun: Petropedetes parkeri n. sp. (Amphibia Anura: Ranidae, Phrynobatrachinae). Revue Suisse de Zoologie, 90, 457 - 468." type="journal article" year="1983">Amiet (1983)</bibRefCitation>
first recognizes that the description of the species habitat by
<bibRefCitation id="5841C5E8FFDAFF83D685FD3643180A7B" author="Parker" box="[422,583,749,775]" pageId="21" pageNumber="47" refString="Parker, H. W. (1936) The amphibians of the Mamfe Division, Cameroons. - I. Zoogeography and systematics. Proceedings of the Zoological Society of London, 135 - 163." type="proceedings paper" year="1936">Parker (1936)</bibRefCitation>
and
<bibRefCitation id="5841C5E8FFDAFF83D5A1FD36420D0A7B" author="Sanderson" box="[642,850,749,775]" pageId="21" pageNumber="47" refString="Sanderson, I. T. (1936) The amphibians of the Mamfe Division, Cameroon. - II. Ecology of the frogs. Proceedings of the Zoological Society of London, 165 - 208." type="proceedings paper" year="1936">Sanderson (1936)</bibRefCitation>
do not fit the circumstances of the description by
<bibRefCitation id="5841C5E8FFDAFF83D7B4FCC940340A50" author="Boulenger" box="[151,363,786,812]" pageId="21" pageNumber="45" refString="Boulenger, G. A. (1900) A list of the batrachians and reptiles of the Gaboon (French Congo), with descriptions of new genera and species. Proceedings of the Zoological Society of London, 1900, 433 - 456." type="proceedings paper" year="1900">Boulenger (1900)</bibRefCitation>
. He subsequently described this species using distinct morphological characters.
<taxonomicName id="FBD0C39AFFDAFF83D20BFCC944C30A57" box="[1320,1436,786,811]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D20BFCC944C30A57" box="[1320,1436,786,811]" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
and
<taxonomicName id="FBD0C39AFFDAFF83D7EBFCE340350A2D" box="[200,362,824,849]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="euskircheni" status="sp. nov.">
<emphasis id="0EA4640BFFDAFF83D7EBFCE340350A2D" box="[200,362,824,849]" italics="true" pageId="21" pageNumber="22">P. euskircheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDAFF83D651FCEC40900A2D" box="[370,463,823,849]" pageId="21" pageNumber="22" rank="species">
<emphasis id="0EA4640BFFDAFF83D651FCEC40900A2D" bold="true" box="[370,463,823,849]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
are the two largest members of the genus. The fleshy papilla in both species has an oval basis, while it is rounder in the remaining species.
<taxonomicName id="FBD0C39AFFDAFF83D40EFC8642FD0A0A" box="[813,930,861,886]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D40EFC8642FD0A0A" box="[813,930,861,886]" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
is only rudimentary-webbed and therefore easily distinguishable from more fully webbed species (
<taxonomicName id="FBD0C39AFFDAFF83D46BFC59428A0AE7" box="[840,981,898,923]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="palmipes">
<emphasis id="0EA4640BFFDAFF83D46BFC59428A0AE7" box="[840,981,898,923]" italics="true" pageId="21" pageNumber="22">P. palmipes</emphasis>
</taxonomicName>
and
<emphasis id="0EA4640BFFDAFF83D330FC5945DE0AE7" box="[1043,1153,898,923]" italics="true" pageId="21" pageNumber="22">P. p e r re t i</emphasis>
, both fully webbed;
<taxonomicName id="FBD0C39AFFDAFF83D2A5FC59401E0ABD" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="cameronensis">
<emphasis id="0EA4640BFFDAFF83D2A5FC59401E0ABD" italics="true" pageId="21" pageNumber="22">P. cameronensis</emphasis>
</taxonomicName>
and
<taxonomicName id="FBD0C39AFFDAFF83D65DFC73433C0ABD" box="[382,611,936,961]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="juliawurstnerae" status="sp. nov.">
<emphasis id="0EA4640BFFDAFF83D65DFC73433C0ABD" box="[382,611,936,961]" italics="true" pageId="21" pageNumber="22">P. juliawurstnerae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDAFF83D54FFC7C438F0ABD" box="[620,720,935,961]" pageId="21" pageNumber="22" rank="species">
<emphasis id="0EA4640BFFDAFF83D54FFC7C438F0ABD" bold="true" box="[620,720,935,961]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
, both half-webbed). Moreover, the presence of a distinct tympanum and a tympanal papilla seperates
<taxonomicName id="FBD0C39AFFDAFF83D595FC1642700A9A" box="[694,815,973,998]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D595FC1642700A9A" box="[694,815,973,998]" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
from
<taxonomicName id="FBD0C39AFFDAFF83D459FC1645610A9A" box="[890,1086,973,998]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="cameronensis">
<emphasis id="0EA4640BFFDAFF83D459FC1645610A9A" box="[890,1086,973,998]" italics="true" pageId="21" pageNumber="22">P. cameronensis</emphasis>
</taxonomicName>
and
<taxonomicName id="FBD0C39AFFDAFF83D358FC1644580A9A" box="[1147,1287,973,998]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="palmipes">
<emphasis id="0EA4640BFFDAFF83D358FC1644580A9A" box="[1147,1287,973,998]" italics="true" pageId="21" pageNumber="22">P. palmipes</emphasis>
</taxonomicName>
, which lack these characters. The size of the femoral gland distinguishes
<taxonomicName id="FBD0C39AFFDAFF83D4B6FC29454F0D77" box="[917,1040,1010,1035]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D4B6FC29454F0D77" box="[917,1040,1010,1035]" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
from
<taxonomicName id="FBD0C39AFFDAFF83D37CFC2945850D77" box="[1119,1242,1010,1035]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="vulpiae" status="sp. nov.">
<emphasis id="0EA4640BFFDAFF83D37CFC2945850D77" box="[1119,1242,1010,1035]" italics="true" pageId="21" pageNumber="22">P. vulpiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDAFF83D3C0FC2944180D70" box="[1251,1351,1010,1036]" pageId="21" pageNumber="22" rank="species">
<emphasis id="0EA4640BFFDAFF83D3C0FC2944180D70" bold="true" box="[1251,1351,1010,1036]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="FBD0C39AFFDAFF83D2A5FC29405B0D4D" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="johnstoni">
<emphasis id="0EA4640BFFDAFF83D2A5FC29405B0D4D" italics="true" pageId="21" pageNumber="22">P. johnstoni</emphasis>
</taxonomicName>
, which possess much larger ones. Due to a similar appearance,
<emphasis id="0EA4640BFFDAFF83D4D5FBC345390D4D" box="[1014,1126,1048,1073]" italics="true" pageId="21" pageNumber="22">P. p a r k e r i</emphasis>
has been confused with a cryptic species in the past. Distinguishing characters between
<emphasis id="0EA4640BFFDAFF83D4BEFBE6454C0D2A" box="[925,1043,1085,1110]" italics="true" pageId="21" pageNumber="22">P. p a r k e r i</emphasis>
and
<taxonomicName id="FBD0C39AFFDAFF83D376FBE6445E0D2A" box="[1109,1281,1085,1110]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="euskircheni" status="sp. nov.">
<emphasis id="0EA4640BFFDAFF83D376FBE6445E0D2A" box="[1109,1281,1085,1110]" italics="true" pageId="21" pageNumber="22">P. euskircheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDAFF83D229FBE644320D2B" box="[1290,1389,1085,1111]" pageId="21" pageNumber="22" rank="species">
<emphasis id="0EA4640BFFDAFF83D229FBE644320D2B" bold="true" box="[1290,1389,1085,1111]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
are presented in the description of the new species (see below).
</paragraph>
<paragraph id="3C6FB819FFDAFF83D7E7FB5344CB0FFB" blockId="21.[151,1437,152,1932]" pageId="21" pageNumber="22">
As
<typeStatus id="E36B06BBFFDAFF83D7CEFB5340080DDE" box="[237,343,1160,1186]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
,
<bibRefCitation id="5841C5E8FFDAFF83D640FB53435B0DDE" author="Amiet" box="[355,516,1160,1186]" pageId="21" pageNumber="45" refString="Amiet, J. - L. (1983) Une espece meconnue de Petropedetes du Cameroun: Petropedetes parkeri n. sp. (Amphibia Anura: Ranidae, Phrynobatrachinae). Revue Suisse de Zoologie, 90, 457 - 468." type="journal article" year="1983">Amiet (1983)</bibRefCitation>
has chosen a
<taxonomicName id="FBD0C39AFFDAFF83D589FB5342400DDD" box="[682,799,1160,1185]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D589FB5342400DDD" box="[682,799,1160,1185]" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
from Atolo, Mamfe region. The
<typeStatus id="E36B06BBFFDAFF83D38BFB5344460DDE" box="[1192,1305,1160,1186]" pageId="21" pageNumber="22" type="paratype">paratypes</typeStatus>
were from localities in the Mamfe region, but also from the “Falaise de Mbos” and Bangwa. He further investigated material from Kala, Léna (Yaoundé region). He regards the species as vocally and morphologically homogenous throughout its discontinuous range, although he mentioned small deviations in the lengths of the femora between western and eastern populations (Mt. Kala, Yaoundé region;
<bibRefCitation id="5841C5E8FFDAFF83D30EFAC645E10C4B" author="Amiet" box="[1069,1214,1309,1335]" pageId="21" pageNumber="45" refString="Amiet, J. - L. (1983) Une espece meconnue de Petropedetes du Cameroun: Petropedetes parkeri n. sp. (Amphibia Anura: Ranidae, Phrynobatrachinae). Revue Suisse de Zoologie, 90, 457 - 468." type="journal article" year="1983">Amiet 1983</bibRefCitation>
). The
<typeStatus id="E36B06BBFFDAFF83D22AFAC644630C4B" box="[1289,1340,1309,1335]" pageId="21" pageNumber="22">type</typeStatus>
locality at Atolo is located at app.
<quantity id="FB2815FCFFDAFF83D6F2FA9943430C20" box="[465,540,1346,1372]" metricMagnitude="2" metricUnit="m" metricValue="4.25" pageId="21" pageNumber="22" unit="m" value="425.0">425 m</quantity>
a.s.l. The overall altitudinal distribution ranges between 400 and
<quantity id="FB2815FCFFDAFF83D20FFA9944C30C20" box="[1324,1436,1346,1372]" metricMagnitude="2" metricUnit="m" metricValue="7.0104000000000015" pageId="21" pageNumber="22" unit="ft" value="2300.0">2300 feet</quantity>
(approximately
<quantity id="FB2815FCFFDAFF83D678FAB340BB0CFE" box="[347,484,1384,1410]" metricMagnitude="2" metricUnit="m" metricValue="4.1" metricValueMax="7.0" metricValueMin="1.2" pageId="21" pageNumber="22" unit="m" value="410.0" valueMax="700.0" valueMin="120.0">120-700 m</quantity>
;
<bibRefCitation id="5841C5E8FFDAFF83D6D1FAB343D40CFE" author="Parker" box="[498,651,1384,1410]" pageId="21" pageNumber="47" refString="Parker, H. W. (1936) The amphibians of the Mamfe Division, Cameroons. - I. Zoogeography and systematics. Proceedings of the Zoological Society of London, 135 - 163." type="proceedings paper" year="1936">Parker 1936</bibRefCitation>
;
<bibRefCitation id="5841C5E8FFDAFF83D5BAFAB3423C0CFE" author="Sanderson" box="[665,867,1384,1410]" pageId="21" pageNumber="47" refString="Sanderson, I. T. (1936) The amphibians of the Mamfe Division, Cameroon. - II. Ecology of the frogs. Proceedings of the Zoological Society of London, 165 - 208." type="proceedings paper" year="1936">Sanderson 1936</bibRefCitation>
). Own and unpublished findings confirm sites below
<quantity id="FB2815FCFFDAFF83D7C5FA56401D0CDB" box="[230,322,1421,1447]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="m" value="1000.0">1000 m</quantity>
. The sites are a hill close to Mukwecha (
<quantity id="FB2815FCFFDAFF83D43EFA5642330CDB" box="[797,876,1421,1447]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="21" pageNumber="22" unit="m" value="720.0">720 m</quantity>
), Okwangwo (
<quantity id="FB2815FCFFDAFF83D33BFA56453A0CDB" box="[1048,1125,1421,1447]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="22" unit="m" value="900.0">900 m</quantity>
) and material collected by Les Minter from Nkwende Hills near Nguti (
<quantity id="FB2815FCFFDAFF83D585FA6943A80CB0" box="[678,759,1458,1484]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="22" unit="m" value="400.0">400 m</quantity>
).
<bibRefCitation id="5841C5E8FFDAFF83D428FA6942FC0CB0" author="Amiet" box="[779,931,1458,1484]" pageId="21" pageNumber="44" refString="Amiet, J. - L. (1978) Les amphibiens anoures de la region de Mamfe (Cameroun). Annales de la Faculte de Sciences du Cameroun, 25, 189 - 219." type="journal article" year="1978">Amiet (1978</bibRefCitation>
,
<bibRefCitation id="5841C5E8FFDAFF83D48CFA6942B40CB0" author="Amiet" box="[943,1003,1458,1484]" pageId="21" pageNumber="45" refString="Amiet, J. - L. (1983) Une espece meconnue de Petropedetes du Cameroun: Petropedetes parkeri n. sp. (Amphibia Anura: Ranidae, Phrynobatrachinae). Revue Suisse de Zoologie, 90, 457 - 468." type="journal article" year="1983">1983</bibRefCitation>
) instead lists altitudes between
<quantity id="FB2815FCFFDAFF83D242FA6941AC0C8E" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.4" metricValueMin="0.75" pageId="21" pageNumber="22" unit="m" value="1075.0" valueMax="1400.0" valueMin="750.0">750 1400 m</quantity>
from Mbakang and Foto-Ndonchwet and doubts localities in very low altitudes from
<bibRefCitation id="5841C5E8FFDAFF83D23EFA0341B70F6B" author="Sanderson" pageId="21" pageNumber="47" refString="Sanderson, I. T. (1936) The amphibians of the Mamfe Division, Cameroon. - II. Ecology of the frogs. Proceedings of the Zoological Society of London, 165 - 208." type="proceedings paper" year="1936">Sanderson (1936)</bibRefCitation>
and
<bibRefCitation id="5841C5E8FFDAFF83D606FA26408E0F6B" author="Parker" box="[293,465,1533,1559]" pageId="21" pageNumber="47" refString="Parker, H. W. (1936) The amphibians of the Mamfe Division, Cameroons. - I. Zoogeography and systematics. Proceedings of the Zoological Society of London, 135 - 163." type="proceedings paper" year="1936">Parker (1936)</bibRefCitation>
. The altitudinal range of
<taxonomicName id="FBD0C39AFFDAFF83D424FA2642DF0F6A" box="[775,896,1533,1558]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D424FA2642DF0F6A" box="[775,896,1533,1558]" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
may overlap with the altitudinal range of
<taxonomicName id="FBD0C39AFFDAFF83D2A5FA26407D0F40" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="euskircheni" status="sp. nov.">
<emphasis id="0EA4640BFFDAFF83D2A5FA26407D0F40" italics="true" pageId="21" pageNumber="22">P. euskircheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDAFF83D609F9F940D40F40" box="[298,395,1570,1596]" pageId="21" pageNumber="22" rank="species">
<emphasis id="0EA4640BFFDAFF83D609F9F940D40F40" bold="true" box="[298,395,1570,1596]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<typeStatus id="E36B06BBFFDAFF83D6BFF9F9408D0F40" box="[412,466,1570,1596]" pageId="21" pageNumber="22">type</typeStatus>
locality at about
<quantity id="FB2815FCFFDAFF83D584F9F943A70F40" box="[679,760,1570,1596]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="22" unit="m" value="900.0">900 m</quantity>
; see below). Without genetic comparisons, so far the
<taxonomicName id="FBD0C39AFFDAFF83D2A5F9F841AF0F1D" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D2A5F9F841AF0F1D" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
-like populations in Central
<collectingCountry id="44C7F889FFDAFF83D564F993439B0F1E" box="[583,708,1608,1634]" name="Cameroon" pageId="21" pageNumber="22">Cameroon</collectingCountry>
region around Yaoundé and the northern populations of the Manengouba Mountains cannot be assigned with certainty, neither to
<taxonomicName id="FBD0C39AFFDAFF83D4E5F9B645660FFA" box="[966,1081,1645,1670]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D4E5F9B645660FFA" box="[966,1081,1645,1670]" italics="true" pageId="21" pageNumber="22">P. parkeri</emphasis>
</taxonomicName>
nor to
<taxonomicName id="FBD0C39AFFDAFF83D3AEF9B6446F0FFA" box="[1165,1328,1645,1670]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="euskircheni" status="sp. nov.">
<emphasis id="0EA4640BFFDAFF83D3AEF9B6446F0FFA" box="[1165,1328,1645,1670]" italics="true" pageId="21" pageNumber="22">P. euskircheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDAFF83D214F9B644CB0FFB" box="[1335,1428,1645,1671]" pageId="21" pageNumber="22" rank="species">
<emphasis id="0EA4640BFFDAFF83D214F9B644CB0FFB" bold="true" box="[1335,1428,1645,1671]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="3C6FB819FFDAFF83D7E7F94943710E60" blockId="21.[151,1437,152,1932]" pageId="21" pageNumber="22">
Due to the finding of
<taxonomicName id="FBD0C39AFFDAFF83D6F6F94843DB0FD0" box="[469,644,1683,1708]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="euskircheni" status="sp. nov.">
<emphasis id="0EA4640BFFDAFF83D6F6F94843DB0FD0" box="[469,644,1683,1708]" italics="true" pageId="21" pageNumber="22">P. euskircheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1597D970FFDAFF83D5AFF94943AE0FD0" box="[652,753,1682,1708]" pageId="21" pageNumber="22" rank="species">
<emphasis id="0EA4640BFFDAFF83D5AFF94943AE0FD0" bold="true" box="[652,753,1682,1708]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
not all populations can be irrespectively assigned to a particular taxon. We therefore assign unsure localities within
<collectingCountry id="44C7F889FFDAFF83D455F96342AE0FAE" box="[886,1009,1720,1746]" name="Cameroon" pageId="21" pageNumber="22">Cameroon</collectingCountry>
(e.g. Mt. Kala), but also specimens from
<collectingCountry id="44C7F889FFDAFF83D7F4F906407A0F8B" box="[215,293,1757,1783]" name="Gabon" pageId="21" pageNumber="22">Gabon</collectingCountry>
(e.g.
<bibRefCitation id="5841C5E8FFDAFF83D64AF906433C0F8B" author="Fretey" box="[361,611,1757,1783]" pageId="21" pageNumber="46" refString="Fretey, T. &amp; Blanc, C. P. (2001) Inventaire systematique des amphibiens anoures du centre du Gabon. Bulletin de la Societe Zoologique de France, 126, 375 - 390." type="journal article" year="2001">Frétey &amp; Blanc 2001</bibRefCitation>
;
<bibRefCitation id="5841C5E8FFDAFF83D54CF90642160F8B" author="Lotters" box="[623,841,1757,1783]" pageId="21" pageNumber="46" refString="Lotters, S., Gossmann, V., Obame, F. &amp; Bohme, W. (2001) Zur Herpetofauna Gabuns Teil I: Einleitung, Untersuchungsgebiet und Methodik, kommentierte Artenliste der gefundenen Froschlurche. Herpetofauna, 23, 19 - 34." type="journal article" year="2001">
Lötters
<emphasis id="0EA4640BFFDAFF83D5E9F90643A10F8A" box="[714,766,1757,1782]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
. 2001
</bibRefCitation>
) and
<collectingCountry id="44C7F889FFDAFF83D4A9F906453E0F8B" box="[906,1121,1757,1783]" name="Equatorial Guinea" pageId="21" pageNumber="22">Equatorial Guinea</collectingCountry>
(de la
<bibRefCitation id="5841C5E8FFDAFF83D391F90644700F8B" author="Riva" box="[1202,1327,1757,1783]" pageId="21" pageNumber="45" refString="de la Riva, I. (1994) Anfibios anuros del Parque Nacional de Monte Alen, Rio Muni, Guinea Equatorial. Revista Espanola de Herpetologia, 8, 123 - 139." type="journal article" year="1994">Riva 1994</bibRefCitation>
;
<bibRefCitation id="5841C5E8FFDAFF83D218F90641A70E60" author="Lasso" pageId="21" pageNumber="46" refString="Lasso, C. A., Rial, A. I., Castroviejo, J. &amp; de la Riva, I. (2002) Herpetofauna del Parque Nacionale de Monte Alen (Rio Muni, Guinea Equatorial). Graellsia, 58, 21 - 34." type="journal article" year="2002">
Lasso
<emphasis id="0EA4640BFFDAFF83D2A4F90641F10E60" italics="true" pageId="21" pageNumber="22">et al</emphasis>
. 2002
</bibRefCitation>
) to
<taxonomicName id="FBD0C39AFFDAFF83D605F8D840980E60" box="[294,455,1794,1820]" class="Amphibia" family="Petropedetidae" genus="Petropedetes" kingdom="Animalia" order="Anura" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="parkeri">
<emphasis id="0EA4640BFFDAFF83D605F8D840660E60" box="[294,313,1795,1820]" italics="true" pageId="21" pageNumber="22">P.</emphasis>
aff.
<emphasis id="0EA4640BFFDAFF83D652F8D840980E60" box="[369,455,1795,1820]" italics="true" pageId="21" pageNumber="22">parkeri</emphasis>
</taxonomicName>
(
<figureCitation id="A4EBA49CFFDAFF83D6FBF8D9437F0E60" box="[472,544,1794,1820]" captionStart="FIGURE 4" captionStartId="8.[151,255,1966,1990]" captionTargetBox="[212,1373,567,1944]" captionTargetId="figure@8.[212,1374,567,1944]" captionTargetPageId="8" captionText="FIGURE 4. Distribution of known Petropedetes species; compare with text. Specimens from localities marked with an asterisk (*) were not available and are added from literature." httpUri="https://zenodo.org/record/193292/files/figure.png" pageId="21" pageNumber="22">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="74CAEB92FFDAFF83D7E7F8FC43DF0EF0" pageId="21" pageNumber="22" type="etymology">
<paragraph id="3C6FB819FFDAFF83D7E7F8FC43DF0EF0" blockId="21.[151,1437,152,1932]" pageId="21" pageNumber="22">
<emphasis id="0EA4640BFFDAFF83D7E7F8FC400C0E3D" bold="true" box="[196,339,1831,1857]" pageId="21" pageNumber="22">Etymology.</emphasis>
The species has been named by
<bibRefCitation id="5841C5E8FFDAFF83D5FCF8F342DD0E3E" author="Amiet" box="[735,898,1832,1858]" pageId="21" pageNumber="45" refString="Amiet, J. - L. (1983) Une espece meconnue de Petropedetes du Cameroun: Petropedetes parkeri n. sp. (Amphibia Anura: Ranidae, Phrynobatrachinae). Revue Suisse de Zoologie, 90, 457 - 468." type="journal article" year="1983">Amiet (1983)</bibRefCitation>
in honour of Mr. Hampton Wildman Parker, former curator of the Natural History Museum, who conducted a meticulous morphological study on specimens from the Sanderson collection.
</paragraph>
</subSubSection>
</treatment>
</document>