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<document id="D52B372937BFD8DD10B812FE4609B5D2" ID-CLB-Dataset="55729" ID-DOI="10.1080/00222933.2020.1763491" ID-GBIF-Dataset="fb05d004-0f2d-44da-b91e-b6e8304ab2a5" ID-ISSN="1464-5262" ID-Zenodo-Dep="4609148" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1615924067624" checkinUser="carolina" docAuthor="Naruse, Tohru &amp; Ng, Peter K. L." docDate="2020" docId="B15D87DEFFF4BE426E5AFDDE0BD39A6D" docLanguage="en" docName="JNATHIST.54.7-8.445-532.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 54 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2020.1763491" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Scandarma Schubart, Liu and Cuesta 2003" docType="treatment" docVersion="9" lastPageNumber="476" masterDocId="4D64FFA6FFEABE626F33FF9E0F509F28" masterDocTitle="Revision of the sesarmid crab genera Labuanium Serène and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003 and Namlacium Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species" masterLastPageNumber="532" masterPageNumber="445" pageNumber="474" updateTime="1698933465109" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3D63BD972D60517A5048255D34C025A5">Revision of the sesarmid crab genera Labuanium Serène and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003 and Namlacium Serène and Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae), with descriptions of four new genera and two new species</mods:title>
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<mods:namePart id="E8AE0D298016CF8BFBA71D1D03A02B1C">Naruse, Tohru</mods:namePart>
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<mods:namePart id="9E439EEA1D16BCC4A81DCB27C13FC1F4">Ng, Peter K. L.</mods:namePart>
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<mods:date id="288C6411A0002AD365C6AB83F2092ECD">2020</mods:date>
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<treatment id="B15D87DEFFF4BE426E5AFDDE0BD39A6D" ID-DOI="http://doi.org/10.5281/zenodo.6518955" ID-GBIF-Taxon="195191647" ID-Zenodo-Dep="6518955" LSID="urn:lsid:plazi:treatment:B15D87DEFFF4BE426E5AFDDE0BD39A6D" httpUri="http://treatment.plazi.org/id/B15D87DEFFF4BE426E5AFDDE0BD39A6D" lastPageId="32" lastPageNumber="476" pageId="30" pageNumber="474">
<subSubSection id="71EE6543FFF4BE7C6E5AFDDE0CEA9D72" box="[361,954,576,602]" pageId="30" pageNumber="474" type="nomenclature">
<paragraph id="394B36C8FFF4BE7C6E5AFDDE0CEA9D72" blockId="30.[361,954,576,602]" box="[361,954,576,602]" pageId="30" pageNumber="474">
<heading id="620381A4FFF4BE7C6E5AFDDE0CEA9D72" box="[361,954,576,602]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="474" reason="2">
Genus
<taxonomicName id="FEF44D4BFFF4BE7C6E88FDDE0CEA9D72" ID-CoL="7PYWQ" authority="Schubart, Liu and Cuesta, 2003" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[443,954,576,602]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF4BE7C6E88FDDE0D159D72" bold="true" box="[443,581,576,602]" italics="true" pageId="30" pageNumber="474">Scandarma</emphasis>
<bibRefCitation id="5D654B39FFF4BE7C6D7EFDDE0CEA9D72" author="Schubart CD &amp; Liu HC &amp; Cuesta JA" box="[589,954,576,602]" pageId="30" pageNumber="474" pagination="49 - 59" refId="ref46378" refString="Schubart CD, Liu HC, Cuesta JA. 2003. A new genus and species of tree-climbing crab (Crustacea: Brachyura: Sesarmidae) from Taiwan with notes on its ecology and larval morphology. Raffles Bull Zool. 51 (1): 49 - 59." type="journal article" year="2003">Schubart, Liu and Cuesta, 2003</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="71EE6543FFF4BE7C6F93FDEA0E5C9D98" pageId="30" pageNumber="474" type="reference_group">
<paragraph id="394B36C8FFF4BE7C6F93FDEA0E5C9D98" blockId="30.[160,1156,628,688]" pageId="30" pageNumber="474">
<treatmentCitationGroup id="19E411E6FFF4BE7C6F93FDEA0E5C9D98" pageId="30" pageNumber="474">
<treatmentCitation id="B85510D9FFF4BE7C6F93FDEA0D999DA5" author="Schubart CD &amp; Liu HC &amp; Cuesta JA" box="[160,713,628,653]" page="50" pageId="30" pageNumber="474" year="2003">
<taxonomicName id="FEF44D4BFFF4BE7C6F93FDEA0D999DA5" ID-CoL="7PYWQ" authority="Schubart, Liu and Cuesta, 2003, p. 50" authorityName="Schubart, Liu and Cuesta" authorityPageNumber="50" authorityYear="2003" box="[160,713,628,653]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF4BE7C6F93FDEA0E4C9DA5" box="[160,284,628,653]" italics="true" pageId="30" pageNumber="474">Scandarma</emphasis>
<bibRefCitation id="5D654B39FFF4BE7C6E17FDEA0DD29DA5" author="Schubart CD &amp; Liu HC &amp; Cuesta JA" box="[292,642,628,653]" pageId="30" pageNumber="474" pagination="49 - 59" refId="ref46378" refString="Schubart CD, Liu HC, Cuesta JA. 2003. A new genus and species of tree-climbing crab (Crustacea: Brachyura: Sesarmidae) from Taiwan with notes on its ecology and larval morphology. Raffles Bull Zool. 51 (1): 49 - 59." type="journal article" year="2003">Schubart, Liu and Cuesta, 2003</bibRefCitation>
, p. 50
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="B85510D9FFF4BE7C6DE7FDEA0C849DA5" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[724,980,628,653]" page="223" pageId="30" pageNumber="474" year="2008">
<bibRefCitation id="5D654B39FFF4BE7C6DE7FDEA0C2F9DA5" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[724,895,628,653]" pageId="30" pageNumber="474" pagination="1 - 286" refId="ref44591" refString="Ng PKL, Guinot D, Davie PJF. 2008 a. Systema Brachyurorum: part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bull Zool Suppl. 17: 1 - 286." type="journal article" year="2008">Ng et al. 2008a</bibRefCitation>
, p. 223
</treatmentCitation>
; Ng et al. 2017, p. 102.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFF4BE7C6F93FD420D6E9CEE" pageId="30" pageNumber="474" type="type_taxon">
<paragraph id="394B36C8FFF4BE7C6F93FD420E639DDE" blockId="30.[160,1156,732,792]" box="[160,307,732,758]" pageId="30" pageNumber="474">
<emphasis id="0B80EADAFFF4BE7C6F93FD420E639DDE" bold="true" box="[160,307,732,758]" italics="true" pageId="30" pageNumber="474">
<typeStatus id="E64F886AFFF4BE7C6F93FD420F889DDE" box="[160,216,732,758]" pageId="30" pageNumber="474">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="394B36C8FFF4BE7C6F93FD610BD49C30" blockId="30.[160,1156,732,792]" box="[160,1156,767,792]" pageId="30" pageNumber="474">
<taxonomicName id="FEF44D4BFFF4BE7C6F93FD610D919C30" ID-CoL="6XS9X" authority="Schubart, Liu and Cuesta, 2003" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[160,705,767,792]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="lintou">
<emphasis id="0B80EADAFFF4BE7C6F93FD610E319C30" box="[160,353,767,792]" italics="true" pageId="30" pageNumber="474">Scandarma lintou</emphasis>
<bibRefCitation id="5D654B39FFF4BE7C6E5BFD610D919C30" author="Schubart CD &amp; Liu HC &amp; Cuesta JA" box="[360,705,767,792]" pageId="30" pageNumber="474" pagination="49 - 59" refId="ref46378" refString="Schubart CD, Liu HC, Cuesta JA. 2003. A new genus and species of tree-climbing crab (Crustacea: Brachyura: Sesarmidae) from Taiwan with notes on its ecology and larval morphology. Raffles Bull Zool. 51 (1): 49 - 59." type="journal article" year="2003">Schubart, Liu and Cuesta, 2003</bibRefCitation>
</taxonomicName>
, by original designation. Gender neuter.
</paragraph>
<paragraph id="394B36C8FFF4BE7C6F93FCDA0E0F9C76" blockId="30.[160,1156,836,966]" box="[160,351,836,862]" pageId="30" pageNumber="474">
<emphasis id="0B80EADAFFF4BE7C6F93FCDA0E0F9C76" bold="true" box="[160,351,836,862]" italics="true" pageId="30" pageNumber="474">Included species</emphasis>
</paragraph>
<paragraph id="394B36C8FFF4BE7C6F93FCF90D6E9CEE" blockId="30.[160,1156,836,966]" pageId="30" pageNumber="474">
<taxonomicName id="FEF44D4BFFF4BE7C6F93FCF90DEF9CA8" ID-CoL="6XS9X" authority="Schubart, Liu and Cuesta, 2003" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[160,703,871,896]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="lintou">
<emphasis id="0B80EADAFFF4BE7C6F93FCF90E319CA8" box="[160,353,871,896]" italics="true" pageId="30" pageNumber="474">Scandarma lintou</emphasis>
<bibRefCitation id="5D654B39FFF4BE7C6E54FCF90DEF9CA8" author="Schubart CD &amp; Liu HC &amp; Cuesta JA" box="[359,703,871,896]" pageId="30" pageNumber="474" pagination="49 - 59" refId="ref46378" refString="Schubart CD, Liu HC, Cuesta JA. 2003. A new genus and species of tree-climbing crab (Crustacea: Brachyura: Sesarmidae) from Taiwan with notes on its ecology and larval morphology. Raffles Bull Zool. 51 (1): 49 - 59." type="journal article" year="2003">Schubart, Liu and Cuesta, 2003</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="FEF44D4BFFF4BE7C6DF9FCF90C919CA8" ID-CoL="7Z29G" authority="(Alcock, 1900)" authorityName="Alcock" authorityYear="1900" baseAuthorityName="Alcock" baseAuthorityYear="1900" box="[714,961,871,896]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="finni">
<emphasis id="0B80EADAFFF4BE7C6DF9FCF90C4E9CA8" box="[714,798,871,896]" italics="true" pageId="30" pageNumber="474">Sc. finni</emphasis>
(
<bibRefCitation id="5D654B39FFF4BE7C6C19FCF90CE89CA8" author="Alcock A" box="[810,952,871,896]" pageId="30" pageNumber="474" pagination="279 - 486" refId="ref39915" refString="Alcock A. 1900. Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. J Asiat Soc Bengal. 69 (3): 279 - 486." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="FEF44D4BFFF4BE7C6CF8FCF90F879C8B" ID-CoL="6Y3Y9" authority="Ng, 2013" authorityName="Ng" authorityYear="2013" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="raymondi">
<emphasis id="0B80EADAFFF4BE7C6CF8FCF90B079CA8" box="[971,1111,871,896]" italics="true" pageId="30" pageNumber="474">Sc. raymondi</emphasis>
<bibRefCitation id="5D654B39FFF4BE7C6B6EFCF90F879C8B" author="Ng PKL" pageId="30" pageNumber="474" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng, 2013</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="FEF44D4BFFF4BE7C6FD4FC140DC09C8B" ID-CoL="6Y49X" authority="Naruse and Ng, 2007" authorityName="Naruse and Ng" authorityYear="2007" box="[231,656,906,931]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFF4BE7C6FD4FC140EDE9C8B" box="[231,398,906,931]" italics="true" pageId="30" pageNumber="474">Sc. splendidum</emphasis>
<bibRefCitation id="5D654B39FFF4BE7C6EAAFC140DC09C8B" author="Naruse T &amp; Ng PKL" box="[409,656,906,931]" pageId="30" pageNumber="474" pagination="337 - 341" refId="ref44118" refString="Naruse T, Ng PKL. 2007. Scandarma splendidum, a new species of tree-climbing crab (Decapoda: Brachyura: Sesarmidae) from Sarawak, Malaysia, Borneo. Raffles Bull Zool. 55 (2): 337 - 341." type="journal article" year="2007">Naruse and Ng, 2007</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="FEF44D4BFFF4BE7C6D93FC140C7F9C8B" ID-CoL="84H9N" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" box="[672,815,906,931]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="0B80EADAFFF4BE7C6D93FC140C7F9C8B" box="[672,815,906,931]" italics="true" pageId="30" pageNumber="474">Sc. gracilipes</emphasis>
</taxonomicName>
(H.
<bibRefCitation id="5D654B39FFF4BE7C6C50FC140B1E9C8B" author="Milne Edwards H" box="[867,1102,906,931]" pageId="30" pageNumber="474" pagination="163 - 228" refId="ref43918" refString="Milne Edwards H. 1853. Memoire sur la Famille des Ocypodiens. Ann Scie Nat Zool. 20 (3): 163 - 228, pls. 6 - 11." type="journal article" year="1853">Milne Edwards 1853</bibRefCitation>
);
<taxonomicName id="FEF44D4BFFF4BE7C6B55FC140E599CEE" ID-CoL="84H9P" authority="Naruse &amp; Ng, 2020" authorityName="Naruse &amp; Ng" authorityYear="2020" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="malagasy" status="sp. nov.">
<emphasis id="0B80EADAFFF4BE7C6B55FC140E599CEE" italics="true" pageId="30" pageNumber="474">Sc. malagasy</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10B357A1FFF4BE7C6E22FC330E3A9CEE" box="[273,362,941,966]" pageId="30" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="FEF44D4BFFF4BE7C6E47FC330E8F9CEE" ID-CoL="84H9Q" authority="Naruse &amp; Ng, 2020" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[372,479,941,966]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="papua" status="sp. nov.">
<emphasis id="0B80EADAFFF4BE7C6E47FC330E8F9CEE" box="[372,479,941,966]" italics="true" pageId="30" pageNumber="474">Sc. papua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10B357A1FFF4BE7C6ED5FC330D6E9CEE" box="[486,574,941,966]" pageId="30" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFF4BE7D6F93FC6F0BD49C75" lastPageId="31" lastPageNumber="475" pageId="30" pageNumber="474" type="diagnosis">
<paragraph id="394B36C8FFF4BE7C6F93FC6F0E449B23" blockId="30.[160,1156,1009,1728]" box="[160,276,1009,1035]" pageId="30" pageNumber="474">
<emphasis id="0B80EADAFFF4BE7C6F93FC6F0E449B23" bold="true" box="[160,276,1009,1035]" italics="true" pageId="30" pageNumber="474">Diagnosis</emphasis>
</paragraph>
<paragraph id="394B36C8FFF4BE7D6F93FB8B0BD49C75" blockId="30.[160,1156,1009,1728]" lastBlockId="31.[160,1156,142,861]" lastPageId="31" lastPageNumber="475" pageId="30" pageNumber="474">
Carapace squarish to subsquarish, wider than long; lateral margins parallel to weakly convex laterally, 0–2 rudimentary teeth behind external orbital angle; dorsal surface gently convex, regions weakly defined. Front deflexed at anterior margin of postfrontal lobes, distally recurved, directed anteriorly; frontal margin nearly straight to gradually concave medially, overhanging onto antennular septum and fossae. Two pairs of postfrontal lobes present (lateral lobes sometimes only feebly separated), lateral lobes slightly exceeding mesial lobes anteriorly (
<emphasis id="0B80EADAFFF4BE7C6D00FB7B0E4E9A08" italics="true" pageId="30" pageNumber="474">
<taxonomicName id="FEF44D4BFFF4BE7C6D00FB7B0D939BD6" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" box="[563,707,1253,1278]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="gracilipes">Sc. gracilipes</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6DE2FB7B0C699BD6" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[721,825,1253,1278]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="lintou">Sc. lintou</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6C7BFB7B0CE79BD6" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[840,951,1253,1278]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="papua">Sc. papua</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6CF5FB7B0B079BD6" authorityName="Ng" authorityYear="2013" box="[966,1111,1253,1278]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="raymondi">Sc. raymondi</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6B55FB7B0E4E9A08" authorityName="Naruse and Ng" authorityYear="2007" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="splendidum">Sc. splendidum</taxonomicName>
</emphasis>
) or all lobes aligned anteriorly (
<taxonomicName id="FEF44D4BFFF4BE7C6DB0FA990C409A08" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[643,784,1287,1312]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="malagasy">
<emphasis id="0B80EADAFFF4BE7C6DB0FA990C409A08" box="[643,784,1287,1312]" italics="true" pageId="30" pageNumber="474">Sc. malagasy</emphasis>
</taxonomicName>
); anterior margins of all lobes far from (
<emphasis id="0B80EADAFFF4BE7C6FD9FAB40C199A6B" box="[234,841,1322,1347]" italics="true" pageId="30" pageNumber="474">
<taxonomicName id="FEF44D4BFFF4BE7C6FD9FAB40E159A6B" authorityName="Alcock" authorityYear="1900" baseAuthorityName="Alcock" baseAuthorityYear="1900" box="[234,325,1322,1347]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="finni">Sc. finni</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6E65FAB40EBC9A6B" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[342,492,1322,1347]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="malagasy">Sc. malagasy</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6ECFFAB40DDF9A6B" authorityName="Ng" authorityYear="2013" box="[508,655,1322,1347]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="raymondi">Sc. raymondi</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6D92FAB40C199A6B" authorityName="Naruse and Ng" authorityYear="2007" box="[673,841,1322,1347]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="splendidum">Sc. splendidum</taxonomicName>
</emphasis>
) or relatively close to (
<emphasis id="0B80EADAFFF4BE7C6B55FAB40D5E9A4E" italics="true" pageId="30" pageNumber="474">
<taxonomicName id="FEF44D4BFFF4BE7C6B55FAB40E589A4E" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="gracilipes">Sc. gracilipes</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6E28FAD30ED79A4E" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[283,391,1357,1382]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="lintou">Sc. lintou</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6EA8FAD30D5E9A4E" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[411,526,1357,1382]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="papua">Sc. papua</taxonomicName>
</emphasis>
) but never reaching frontal margin in dorsal view. Antennular septum wide, short, dorsally covered by front. Orbit, in dorsal view, tilted J-shaped, median part of supraorbital margins oblique; inner orbital tooth triangular, short, directed dorsoanteriorly. No longitudinal ridge on ventral surface of external orbital angle. Epistome posterior margin with 3 triangular lobes, lateral lobes directed anteroventrally, median lobe directed ventrally (
<emphasis id="0B80EADAFFF4BE7C6D41FA640B2C993B" box="[626,1148,1530,1555]" italics="true" pageId="30" pageNumber="474">
<taxonomicName id="FEF44D4BFFF4BE7C6D41FA640DAE993B" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" box="[626,766,1530,1555]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="gracilipes">Sc. gracilipes</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6C3AFA640C3D993B" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[777,877,1530,1555]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="lintou">Sc. lintou</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6C4BFA640B57993B" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[888,1031,1530,1555]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="malagasy">Sc. malagasy</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6B21FA640B2C993B" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[1042,1148,1530,1555]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="papua">Sc. papua</taxonomicName>
</emphasis>
) or lateral lobes directed ventrally, median lobe directed ventroposteriorly (
<taxonomicName id="FEF44D4BFFF4BE7C6CDAF9830B26991E" authorityName="Ng" authorityYear="2013" box="[1001,1142,1565,1590]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="raymondi">
<emphasis id="0B80EADAFFF4BE7C6CDAF9830B26991E" box="[1001,1142,1565,1590]" italics="true" pageId="30" pageNumber="474">Sc. raymondi</emphasis>
</taxonomicName>
). Antenna entering orbit through wide gap between inner orbital tooth and front. Mxp3 exopod with distinct flagellum. Chela palm with 1 nearly complete row of granules bordering outer and inner surfaces on upper surface (
<emphasis id="0B80EADAFFF4BE7C6C35F91B0E8E99E8" italics="true" pageId="30" pageNumber="474">
<taxonomicName id="FEF44D4BFFF4BE7C6C35F91B0CC499B6" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" box="[774,916,1669,1694]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="gracilipes">Sc. gracilipes</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6CACF91B0B5599B6" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[927,1029,1669,1694]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="lintou">Sc. lintou</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6B21F91B0B2F99B6" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[1042,1151,1669,1694]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="papua">Sc. papua</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6F93F9390E7E99E8" authorityName="Ng" authorityYear="2013" box="[160,302,1703,1728]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="raymondi">Sc. raymondi</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF4BE7C6E08F9390E8E99E8" authorityName="Naruse and Ng" authorityYear="2007" box="[315,478,1703,1728]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="474" phylum="Arthropoda" rank="species" species="splendidum">Sc. splendidum</taxonomicName>
</emphasis>
) or a few incomplete rows of granules bordering outer and inner surfaces (
<taxonomicName id="FEF44D4BFFF5BE7D6E67FF100EB69F8F" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[340,486,142,167]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="malagasy">
<emphasis id="0B80EADAFFF5BE7D6E67FF100EB69F8F" box="[340,486,142,167]" italics="true" pageId="31" pageNumber="475">Sc. malagasy</emphasis>
</taxonomicName>
) in male and female; outer surface of chela with clearly delimited smooth slope towards bases of fingers, proximal to this slope with rounded to ridge-like protuberance (
<emphasis id="0B80EADAFFF5BE7D6EE5FF4A0B4A9FC5" box="[470,1050,212,237]" italics="true" pageId="31" pageNumber="475">
<taxonomicName id="FEF44D4BFFF5BE7D6EE5FF4A0D329FC5" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" box="[470,610,212,237]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="gracilipes">Sc. gracilipes</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6D5EFF4A0D829FC5" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[621,722,212,237]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="lintou">Sc. lintou</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6DEDFF4A0C3B9FC5" authorityName="Ng" authorityYear="2013" box="[734,875,212,237]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="raymondi">Sc. raymondi</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6C44FF4A0B4A9FC5" authorityName="Naruse and Ng" authorityYear="2007" box="[887,1050,212,237]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="splendidum">Sc. splendidum</taxonomicName>
</emphasis>
) or slope less clearly delimited with lower protuberance (
<emphasis id="0B80EADAFFF5BE7D6DFAFF680C889E27" box="[713,984,246,271]" italics="true" pageId="31" pageNumber="475">
<taxonomicName id="FEF44D4BFFF5BE7D6DFAFF680C0C9E27" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[713,860,246,271]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="malagasy">Sc. malagasy</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6C59FF680C889E27" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[874,984,246,271]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="papua">Sc. papua</taxonomicName>
</emphasis>
); inner surface without transverse crest (
<emphasis id="0B80EADAFFF5BE7D6E88FE870E4E9E7D" italics="true" pageId="31" pageNumber="475">
<taxonomicName id="FEF44D4BFFF5BE7D6E88FE870D169E1A" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" box="[443,582,281,306]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="gracilipes">Sc. gracilipes</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6D63FE870DE49E1A" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[592,692,281,306]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="lintou">Sc. lintou</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6D8CFE870C1E9E1A" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[703,846,281,306]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="malagasy">Sc. malagasy</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6C6BFE870C939E1A" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[856,963,281,306]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="papua">Sc. papua</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6CFDFE870B0B9E1A" authorityName="Ng" authorityYear="2013" box="[974,1115,281,306]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="raymondi">Sc. raymondi</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6B55FE870E4E9E7D" authorityName="Naruse and Ng" authorityYear="2007" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="splendidum">Sc. splendidum</taxonomicName>
</emphasis>
); thick rim extending along occlusal margin of immovable finger to dactylar articulation on both outer and inner surfaces, thick rims of both outer and inner surfaces not interrupted near dactylar articulation. Upper-inner margin of movable finger with a row of closely arranged small granules over proximal three-quarters (
<emphasis id="0B80EADAFFF5BE7D6CFFFE3A0E069EF7" italics="true" pageId="31" pageNumber="475">
<taxonomicName id="FEF44D4BFFF5BE7D6CFFFE3A0B0A9E95" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" box="[972,1114,420,445]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="gracilipes">Sc. gracilipes</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6B55FE3A0FB09EF7" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="lintou">Sc. lintou</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6FDFFE580E069EF7" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[236,342,454,479]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="papua">Sc. papua</taxonomicName>
</emphasis>
) or 1 row of sparsely aligned small granules (
<emphasis id="0B80EADAFFF5BE7D6C65FE580E129D2A" italics="true" pageId="31" pageNumber="475">
<taxonomicName id="FEF44D4BFFF5BE7D6C65FE580CB69EF7" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[854,998,454,479]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="malagasy">Sc. malagasy</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6CC2FE580B2F9EF7" authorityName="Ng" authorityYear="2013" box="[1009,1151,454,479]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="raymondi">Sc. raymondi</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6F93FE770E129D2A" authorityName="Naruse and Ng" authorityYear="2007" box="[160,322,489,514]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="splendidum">Sc. splendidum</taxonomicName>
</emphasis>
). Ambulatory legs (P2–5) relatively long, slender, dorso-ventrally flattened; distal anterior corner of each merus angled, not produced, followed proximally by subdistal tooth; carpi and propodi not distinctly narrower than respective meri; dactyli as long as or longer than half length of respective propodi. Male thoracic sternum transversely wide; sternite 8 clearly exposed. Male pleon wide (
<emphasis id="0B80EADAFFF5BE7D6DD9FDEA0CBC9DA5" box="[746,1004,628,653]" italics="true" pageId="31" pageNumber="475">
<taxonomicName id="FEF44D4BFFF5BE7D6DD9FDEA0C299DA5" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" box="[746,889,628,653]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="gracilipes">Sc. gracilipes</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6CB5FDEA0CBC9DA5" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[902,1004,628,653]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="lintou">Sc. lintou</taxonomicName>
</emphasis>
) or relatively narrow (
<emphasis id="0B80EADAFFF5BE7D6FCEFD080EAF9D87" box="[253,511,662,687]" italics="true" pageId="31" pageNumber="475">
<taxonomicName id="FEF44D4BFFF5BE7D6FCEFD080EDC9D87" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[253,396,662,687]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="malagasy">Sc. malagasy</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6EA6FD080EAF9D87" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[405,511,662,687]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="papua">Sc. papua</taxonomicName>
</emphasis>
,
<emphasis id="0B80EADAFFF5BE7D6D39FD080C139D87" box="[522,835,662,687]" italics="true" pageId="31" pageNumber="475">
<taxonomicName id="FEF44D4BFFF5BE7D6D39FD080DC79D87" authorityName="Ng" authorityYear="2013" box="[522,663,662,687]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="raymondi">Sc. raymondi</taxonomicName>
,
<taxonomicName id="FEF44D4BFFF5BE7D6D91FD080C139D87" authorityName="Naruse and Ng" authorityYear="2007" box="[674,835,662,687]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="splendidum">Sc. splendidum</taxonomicName>
</emphasis>
) bell shaped, lateral margins of somites 3–5 forming a concave margin, telson slightly exceeding midlength of bases of cheliped coxae. G1 stout, relatively short, reaching midlength to distal end of thoracic sternite
<quantity id="FE0C9B2DFFF5BE7D6FCCFD600E7D9C3F" box="[255,301,766,791]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="31" pageNumber="475" unit="in" value="5.0">5 in</quantity>
situ; beak-like and corneous process wide to narrow. Vulvae adjacent to thoracic sternal suture 5/6 on sternite 6, ellipsoidal, sternal vulval cover developed from anterolateral or posterolateral margin, convering almost entire to three-quarters of vulva.
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFF5BE426F93FC060E1B9E95" lastPageId="32" lastPageNumber="476" pageId="31" pageNumber="475" type="discussion">
<paragraph id="394B36C8FFF5BE7D6F93FC060E569C9A" blockId="31.[160,1156,920,1709]" box="[160,262,920,946]" pageId="31" pageNumber="475">
<emphasis id="0B80EADAFFF5BE7D6F93FC060E569C9A" bold="true" box="[160,262,920,946]" italics="true" pageId="31" pageNumber="475">Remarks</emphasis>
</paragraph>
<paragraph id="394B36C8FFF5BE426F93FC220E5C9FE2" blockId="31.[160,1156,920,1709]" lastBlockId="32.[160,1156,142,445]" lastPageId="32" lastPageNumber="476" pageId="31" pageNumber="475">
<taxonomicName id="FEF44D4BFFF5BE7D6F93FC220E4C9CFD" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[160,284,956,981]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6F93FC220E4C9CFD" box="[160,284,956,981]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
was established for
<taxonomicName id="FEF44D4BFFF5BE7D6D3FFC220CB09CFD" authority="Schubart, Liu and Cuesta, 2003" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[524,992,956,981]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="lintou">
<emphasis id="0B80EADAFFF5BE7D6D3FFC220D229CFD" box="[524,626,956,981]" italics="true" pageId="31" pageNumber="475">Sc. lintou</emphasis>
<bibRefCitation id="5D654B39FFF5BE7D6D48FC220CB09CFD" author="Schubart CD &amp; Liu HC &amp; Cuesta JA" box="[635,992,956,981]" pageId="31" pageNumber="475" pagination="49 - 59" refId="ref46378" refString="Schubart CD, Liu HC, Cuesta JA. 2003. A new genus and species of tree-climbing crab (Crustacea: Brachyura: Sesarmidae) from Taiwan with notes on its ecology and larval morphology. Raffles Bull Zool. 51 (1): 49 - 59." type="journal article" year="2003">Schubart, Liu and Cuesta, 2003</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="41E37658FFF5BE7D6B1DFC220BD09CFD" box="[1070,1152,956,981]" name="Taiwan" pageId="31" pageNumber="475">Taiwan</collectingCountry>
.
<bibRefCitation id="5D654B39FFF5BE7D6F93FC400ECD9CDF" author="Naruse T &amp; Ng PKL" box="[160,413,990,1015]" pageId="31" pageNumber="475" pagination="337 - 341" refId="ref44118" refString="Naruse T, Ng PKL. 2007. Scandarma splendidum, a new species of tree-climbing crab (Decapoda: Brachyura: Sesarmidae) from Sarawak, Malaysia, Borneo. Raffles Bull Zool. 55 (2): 337 - 341." type="journal article" year="2007">Naruse and Ng (2007)</bibRefCitation>
;
<bibRefCitation id="5D654B39FFF5BE7D6E99FC400D4A9CDF" author="Ng PKL" box="[426,538,990,1015]" pageId="31" pageNumber="475" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng (2013)</bibRefCitation>
subsequently described
<taxonomicName id="FEF44D4BFFF5BE7D6C0AFC400C8E9CDF" authorityName="Naruse and Ng" authorityYear="2007" box="[825,990,990,1015]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFF5BE7D6C0AFC400C8E9CDF" box="[825,990,990,1015]" italics="true" pageId="31" pageNumber="475">Sc. splendidum</emphasis>
</taxonomicName>
from
<collectingCountry id="41E37658FFF5BE7D6B14FC400BD49CDF" box="[1063,1156,990,1015]" name="Malaysia" pageId="31" pageNumber="475">Sarawak</collectingCountry>
and
<taxonomicName id="FEF44D4BFFF5BE7D6FE0FB9F0E309B32" authorityName="Ng" authorityYear="2013" box="[211,352,1025,1050]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="species" species="raymondi">
<emphasis id="0B80EADAFFF5BE7D6FE0FB9F0E309B32" box="[211,352,1025,1050]" italics="true" pageId="31" pageNumber="475">Sc. raymondi</emphasis>
</taxonomicName>
from
<collectingRegion id="FB30F82AFFF5BE7D6E96FB9F0EBD9B32" box="[421,493,1025,1050]" country="Malaysia" name="Sabah" pageId="31" pageNumber="475">Sabah</collectingRegion>
, respectively. The present study adds four more species to the genus (see below).
<taxonomicName id="FEF44D4BFFF5BE7D6E87FBBA0D609B15" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[436,560,1060,1085]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6E87FBBA0D609B15" box="[436,560,1060,1085]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
is characterised by the almost parallel and usually entire (or with 2 rudimentary teeth) lateral margins of the carapace, proportionally wide front (
<figureCitation id="A1CF2A4DFFF5BE7D6FD7FBF70E0B9BAA" box="[228,347,1129,1154]" captionStart="Figure 15" captionStartId="33.[159,224,1193,1216]" captionTargetBox="[163,1153,142,1161]" captionTargetId="figure-160@33.[163,1153,142,1162]" captionTargetPageId="33" captionText="Figure 15. Scandarma lintou Schubart, Liu and Cuesta, 2003. Paratype, ZRC 2002.0168, male, 19.0 × 19.6 mm (Taiwan). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.4609184" httpUri="https://zenodo.org/record/4609184/files/figure.png" pageId="31" pageNumber="475">Figures 15</figureCitation>
(a), 18(a), 22(a), 24(a)), the presence of a distally sloping smooth area on the outer surface of the male cheliped palm towards the bases of the fingers (
<figureCitation id="A1CF2A4DFFF5BE7D6B3EFB120BD39B8D" box="[1037,1155,1164,1189]" captionStart="Figure 16" captionStartId="34.[160,225,729,752]" captionTargetBox="[163,1153,142,698]" captionTargetId="figure-342@34.[163,1153,142,698]" captionTargetPageId="34" captionText="Figure 16. Scandarma lintou Schubart, Liu and Cuesta, 2003. Paratype, ZRC 2002.0168, male, 19.0 × 19.6 mm (Taiwan). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.4609186" httpUri="https://zenodo.org/record/4609186/files/figure.png" pageId="31" pageNumber="475">Figures 16</figureCitation>
(a), 18(b), 22(b), 24(c)), and relatively long and slender ambulatory legs (
<figureCitation id="A1CF2A4DFFF5BE7D6CF8FB300B6F9BEF" box="[971,1087,1198,1223]" captionStart="Figure 15" captionStartId="33.[159,224,1193,1216]" captionTargetBox="[163,1153,142,1161]" captionTargetId="figure-160@33.[163,1153,142,1162]" captionTargetPageId="33" captionText="Figure 15. Scandarma lintou Schubart, Liu and Cuesta, 2003. Paratype, ZRC 2002.0168, male, 19.0 × 19.6 mm (Taiwan). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.4609184" httpUri="https://zenodo.org/record/4609184/files/figure.png" pageId="31" pageNumber="475">Figures 15</figureCitation>
(a), 18 (a), 22(a)).
<taxonomicName id="FEF44D4BFFF5BE7D6E26FB4F0EC19BC2" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[277,401,1233,1258]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6E26FB4F0EC19BC2" box="[277,401,1233,1258]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="FEF44D4BFFF5BE7D6D13FB4F0DD69BC2" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[544,646,1233,1258]" class="Malacostraca" family="Sesarmidae" genus="Circulium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6D13FB4F0DD69BC2" box="[544,646,1233,1258]" italics="true" pageId="31" pageNumber="475">Circulium</emphasis>
</taxonomicName>
in its moderately wide carapace and proportionally long legs.
<taxonomicName id="FEF44D4BFFF5BE7D6E5AFB6A0EB59A25" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[361,485,1268,1293]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6E5AFB6A0EB59A25" box="[361,485,1268,1293]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
, however, can be distinguished from
<taxonomicName id="FEF44D4BFFF5BE7D6CB0FB6A0CB99A25" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[899,1001,1268,1293]" class="Malacostraca" family="Sesarmidae" genus="Circulium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6CB0FB6A0CB99A25" box="[899,1001,1268,1293]" italics="true" pageId="31" pageNumber="475">Circulium</emphasis>
</taxonomicName>
by characters of the carapace, male chela, ambulatory legs and male pleon (
<tableCitation id="74760373FFF5BE7D6C4DFA880C849A07" box="[894,980,1302,1327]" captionStart="Table 1" captionStartId="5.[133,189,438,461]" captionTargetPageId="5" captionText="Table 1. Diagnostic characters of Labuanium Serène and Soh, 1970, Circulium gen. nov., Shinobium gen. nov. and Scandarma Schubart, Liu and Cuesta, 2003." httpUri="http://table.plazi.org/id/6D8B6640FFEFBE676FB6FE2809C99EE5" pageId="31" pageNumber="475" tableUuid="6D8B6640FFEFBE676FB6FE2809C99EE5">Table 1</tableCitation>
). The carapace contour of
<taxonomicName id="FEF44D4BFFF5BE7D6E2AFAA70EC59A7A" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[281,405,1337,1362]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6E2AFAA70EC59A7A" box="[281,405,1337,1362]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
is distinctly squarish, with almost parallel and usually entire (or with 1 or 2 rudimentary teeth) lateral margins (
<figureCitation id="A1CF2A4DFFF5BE7D6DB1FAC20DA89A5D" box="[642,760,1372,1397]" captionStart="Figure 15" captionStartId="33.[159,224,1193,1216]" captionTargetBox="[163,1153,142,1161]" captionTargetId="figure-160@33.[163,1153,142,1162]" captionTargetPageId="33" captionText="Figure 15. Scandarma lintou Schubart, Liu and Cuesta, 2003. Paratype, ZRC 2002.0168, male, 19.0 × 19.6 mm (Taiwan). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.4609184" httpUri="https://zenodo.org/record/4609184/files/figure.png" pageId="31" pageNumber="475">Figures 15</figureCitation>
(a), 18(a), 22(a), 24(a)). In
<taxonomicName id="FEF44D4BFFF5BE7D6B24FAC20B2D9A5D" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[1047,1149,1372,1397]" class="Malacostraca" family="Sesarmidae" genus="Circulium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6B24FAC20B2D9A5D" box="[1047,1149,1372,1397]" italics="true" pageId="31" pageNumber="475">Circulium</emphasis>
</taxonomicName>
, the lateral margin of the carapace is inflated laterally and dorsally around its branchial regions, appearing rounded, with usually 2 distinct teeth (
<figureCitation id="A1CF2A4DFFF5BE7D6C1DFA3F0CDB9A92" box="[814,907,1441,1466]" captionStart="Figure 5" captionStartId="15.[160,225,1123,1146]" captionTargetBox="[181,1133,142,1091]" captionTargetId="figure-132@15.[181,1133,142,1091]" captionTargetPageId="15" captionText="Figure 5. Circulium rotundatum (Hess, 1865). Neotype, USNM 1150304, male, 36.3 × 36.3 mm (Tonga). a, habitus, dorsal view; b, anterior part of cephalothorax, anteroventral view." figureDoi="http://doi.org/10.5281/zenodo.4609164" httpUri="https://zenodo.org/record/4609164/files/figure.png" pageId="31" pageNumber="475">Figure 5</figureCitation>
(a)). The male chela of
<taxonomicName id="FEF44D4BFFF5BE7D6F93FA5A0E4C9AF5" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[160,284,1476,1501]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6F93FA5A0E4C9AF5" box="[160,284,1476,1501]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
is unique in the presence of a distally sloping smooth area on the outer surface between the bases of the fingers (
<figureCitation id="A1CF2A4DFFF5BE7D6D00FA780DFB9AD7" box="[563,683,1510,1535]" captionStart="Figure 16" captionStartId="34.[160,225,729,752]" captionTargetBox="[163,1153,142,698]" captionTargetId="figure-342@34.[163,1153,142,698]" captionTargetPageId="34" captionText="Figure 16. Scandarma lintou Schubart, Liu and Cuesta, 2003. Paratype, ZRC 2002.0168, male, 19.0 × 19.6 mm (Taiwan). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.4609186" httpUri="https://zenodo.org/record/4609186/files/figure.png" pageId="31" pageNumber="475">Figures 16</figureCitation>
(a), 18(b), 22(b), 24(c)), proximally having a rounded or ridge-like protuberance in most species (
<figureCitation id="A1CF2A4DFFF5BE7D6C19F9970CF4990A" box="[810,932,1545,1570]" captionStart="Figure 16" captionStartId="34.[160,225,729,752]" captionTargetBox="[163,1153,142,698]" captionTargetId="figure-342@34.[163,1153,142,698]" captionTargetPageId="34" captionText="Figure 16. Scandarma lintou Schubart, Liu and Cuesta, 2003. Paratype, ZRC 2002.0168, male, 19.0 × 19.6 mm (Taiwan). a–c, left chela; a, outer view; b, inner view; c, upper view; d, meri of right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.4609186" httpUri="https://zenodo.org/record/4609186/files/figure.png" pageId="31" pageNumber="475">Figures 16</figureCitation>
(a), 18(b)), which is absent in
<taxonomicName id="FEF44D4BFFF5BE7D6E26F9B20E2B996D" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[277,379,1580,1605]" class="Malacostraca" family="Sesarmidae" genus="Circulium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6E26F9B20E2B996D" box="[277,379,1580,1605]" italics="true" pageId="31" pageNumber="475">Circulium</emphasis>
</taxonomicName>
(
<figureCitation id="A1CF2A4DFFF5BE7D6EBEF9B20EBC996D" box="[397,492,1580,1605]" captionStart="Figure 6" captionStartId="16.[159,224,776,799]" captionTargetBox="[181,1133,142,744]" captionTargetId="figure-251@16.[181,1133,142,744]" captionTargetPageId="16" captionText="Figure 6. Circulium rotundatum (Hess, 1865). Neotype, USNM 1150304, male, 36.3 × 36.3 mm (Tonga). a, right chela, outer view; bright chela, upper view; c, cephalothorax, ventral view; d, meri of right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.4609166" httpUri="https://zenodo.org/record/4609166/files/figure.png" pageId="31" pageNumber="475">Figure 6</figureCitation>
(a)).
<taxonomicName id="FEF44D4BFFF5BE7D6D2CF9B20DCB996D" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[543,667,1580,1605]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6D2CF9B20DCB996D" box="[543,667,1580,1605]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
and
<taxonomicName id="FEF44D4BFFF5BE7D6DE9F9B20C10996D" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[730,832,1580,1605]" class="Malacostraca" family="Sesarmidae" genus="Circulium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6DE9F9B20C10996D" box="[730,832,1580,1605]" italics="true" pageId="31" pageNumber="475">Circulium</emphasis>
</taxonomicName>
share proportionally longer legs, but those of
<taxonomicName id="FEF44D4BFFF5BE7D6E58F9D00EB7994F" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[363,487,1614,1639]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6E58F9D00EB7994F" box="[363,487,1614,1639]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
are relatively more slender than in
<taxonomicName id="FEF44D4BFFF5BE7D6C4BF9D00C8E994F" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[888,990,1614,1639]" class="Malacostraca" family="Sesarmidae" genus="Circulium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6C4BF9D00C8E994F" box="[888,990,1614,1639]" italics="true" pageId="31" pageNumber="475">Circulium</emphasis>
</taxonomicName>
. The dactyli of the ambulatory legs are proportionally longer in
<taxonomicName id="FEF44D4BFFF5BE7D6DF0F9EF0C6F99A2" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[707,831,1649,1674]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6DF0F9EF0C6F99A2" box="[707,831,1649,1674]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
than in
<taxonomicName id="FEF44D4BFFF5BE7D6CA8F9EF0B5199A2" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[923,1025,1649,1674]" class="Malacostraca" family="Sesarmidae" genus="Circulium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6CA8F9EF0B5199A2" box="[923,1025,1649,1674]" italics="true" pageId="31" pageNumber="475">Circulium</emphasis>
</taxonomicName>
: the dactyli of
<taxonomicName id="FEF44D4BFFF5BE7D6F8FF90A0E689985" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[188,312,1684,1709]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="475" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFF5BE7D6F8FF90A0E689985" box="[188,312,1684,1709]" italics="true" pageId="31" pageNumber="475">Scandarma</emphasis>
</taxonomicName>
are as long as or longer than half the length of the respective propodi in P2– 5, rather than being shorter than half the length of the respective propodi in P
<quantity id="FE0C9B2DFFCABE426B0EFF100BD39F8F" box="[1085,1155,142,167]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="12.7" metricValueMin="5.08" pageId="32" pageNumber="476" unit="in" value="3.5" valueMax="5.0" valueMin="2.0">2–5 in</quantity>
<taxonomicName id="FEF44D4BFFCABE426F93FF2F0E569FE2" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[160,262,177,202]" class="Malacostraca" family="Sesarmidae" genus="Circulium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFCABE426F93FF2F0E569FE2" box="[160,262,177,202]" italics="true" pageId="32" pageNumber="476">Circulium</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="394B36C8FFCABE426FF3FF4A0E1B9E95" blockId="32.[160,1156,142,445]" pageId="32" pageNumber="476">
In the present study, we recognise seven species, including two new species, of
<taxonomicName id="FEF44D4BFFCABE426F93FF680E4C9E27" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[160,284,246,271]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFCABE426F93FF680E4C9E27" box="[160,284,246,271]" italics="true" pageId="32" pageNumber="476">Scandarma</emphasis>
</taxonomicName>
. Of them,
<taxonomicName id="FEF44D4BFFCABE426EA4FF680EBE9E27" authorityName="Alcock" authorityYear="1900" baseAuthorityName="Alcock" baseAuthorityYear="1900" box="[407,494,246,271]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="species" species="finni">
<emphasis id="0B80EADAFFCABE426EA4FF680EBE9E27" box="[407,494,246,271]" italics="true" pageId="32" pageNumber="476">Sc. finni</emphasis>
</taxonomicName>
is certainly different from all other species of
<taxonomicName id="FEF44D4BFFCABE426B31FF680B2E9E27" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[1026,1150,246,271]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFCABE426B31FF680B2E9E27" box="[1026,1150,246,271]" italics="true" pageId="32" pageNumber="476">Scandarma</emphasis>
</taxonomicName>
, but its generic identity is not fully established, because the
<typeStatus id="E64F886AFFCABE426C1FFE870CC39E1A" box="[812,915,281,306]" pageId="32" pageNumber="476" type="holotype">holotype</typeStatus>
, the only specimen of the taxon available for this study, is an immature female (see below). The taxon is treated as a species of
<taxonomicName id="FEF44D4BFFCABE426E60FEC00E9F9E5F" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[339,463,350,375]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFCABE426E60FEC00E9F9E5F" box="[339,463,350,375]" italics="true" pageId="32" pageNumber="476">Scandarma</emphasis>
</taxonomicName>
for the time being, as its characters best fit the genus. The following identification key includes six species, excluding
<taxonomicName id="FEF44D4BFFCABE426C05FE1F0CDB9EB2" authorityName="Alcock" authorityYear="1900" baseAuthorityName="Alcock" baseAuthorityYear="1900" box="[822,907,385,410]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="species" species="finni">
<emphasis id="0B80EADAFFCABE426C05FE1F0CDB9EB2" box="[822,907,385,410]" italics="true" pageId="32" pageNumber="476">Sc. finni</emphasis>
</taxonomicName>
as its male characters are not known.
</paragraph>
</subSubSection>
<subSubSection id="71EE6543FFCABE426F93FE770BD39A6D" pageId="32" pageNumber="476" type="key">
<paragraph id="394B36C8FFCABE426F93FE770CCC9D2A" blockId="32.[160,924,489,514]" box="[160,924,489,514]" pageId="32" pageNumber="476">
<emphasis id="0B80EADAFFCABE426F93FE770CCC9D2A" bold="true" box="[160,924,489,514]" pageId="32" pageNumber="476">
Key to species of
<taxonomicName id="FEF44D4BFFCABE426E40FE770EA69D2A" ID-CoL="7PYWQ" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[371,502,489,514]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="0B80EADAFFCABE426E40FE770EA69D2A" bold="true" box="[371,502,489,514]" italics="true" pageId="32" pageNumber="476">Scandarma</emphasis>
</taxonomicName>
(except for
<taxonomicName id="FEF44D4BFFCABE426DB4FE770CCC9D2A" ID-CoL="7Z29G" authority="(Alcock, 1900))" authorityName="Alcock" authorityYear="1900" baseAuthorityName="Alcock" baseAuthorityYear="1900" box="[647,924,489,514]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="species" species="finni">
<emphasis id="0B80EADAFFCABE426DB4FE770DB39D2A" bold="true" box="[647,739,489,514]" italics="true" pageId="32" pageNumber="476">Sc. finni</emphasis>
(
<bibRefCitation id="5D654B39FFCABE426DC2FE770CDB9D2A" author="Alcock A" box="[753,907,489,514]" pageId="32" pageNumber="476" pagination="279 - 486" refId="ref39915" refString="Alcock A. 1900. Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. J Asiat Soc Bengal. 69 (3): 279 - 486." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
))
</taxonomicName>
</emphasis>
</paragraph>
<key id="4D6561F9FFCABE426F93FDB20BD39A6D" pageId="32" pageNumber="476">
<keyStep id="82002E6DFFCABE426F93FDB20BD39DA2" pageId="32" pageNumber="476">
<paragraph id="394B36C8FFCABE426F93FDB20BD39D4F" blockId="32.[160,1155,556,650]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FDB20BD39D4F" pageId="32" pageNumber="476">1. Anterior margins of postfrontal lobes relatively close to but never reaching frontal margin in dorsal view... ............................................................................................................................ 2</keyLead>
</paragraph>
<paragraph id="394B36C8FFCABE426F93FDEF0BD39DA2" blockId="32.[160,1155,556,650]" box="[160,1155,625,650]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FDEF0BD39DA2" box="[160,1155,625,650]" pageId="32" pageNumber="476">– Anterior margins of postfrontal lobes far from frontal margin in dorsal view... ...... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="82002E6DFFCABE426F93FD380BD39C0F" pageId="32" pageNumber="476">
<paragraph id="394B36C8FFCABE426F93FD380BD49DCA" blockId="32.[160,1156,678,807]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FD380BD49DCA" pageId="32" pageNumber="476">
2. Outer surface of male chela with a weak protuberance proximal to smooth slope area around bases of fingers.. .........................................................................................
<taxonomicName id="FEF44D4BFFCABE426C88FD570B759DCA" ID-CoL="84H9Q" authority="Naruse &amp; Ng, 2020" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[955,1061,713,738]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="species" species="papua" status="sp. nov.">
<emphasis id="0B80EADAFFCABE426C88FD570B759DCA" box="[955,1061,713,738]" italics="true" pageId="32" pageNumber="476">Sc. papua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10B357A1FFCABE426B1EFD570BD49DCA" box="[1069,1156,713,738]" pageId="32" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="394B36C8FFCABE426F93FD720BD39C0F" blockId="32.[160,1156,678,807]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FD720BD39C0F" pageId="32" pageNumber="476">– Outer surface of male chela with a strong protuberance proximal to smooth slope area around base of fingers.. ......................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="82002E6DFFCABE426F93FCDA0BD49CED" pageId="32" pageNumber="476">
<paragraph id="394B36C8FFCABE426F93FCDA0BD39C57" blockId="32.[160,1156,836,965]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FCDA0BD39C57" pageId="32" pageNumber="476">
3. Ambulatory legs relatively longer (P4 merus 0.79–0.89 times carapace length).. ............ ...............................................................................................
<taxonomicName id="FEF44D4BFFCABE426D8FFCF80BD39C57" ID-CoL="6XS9X" authority="Schubart, Liu and Cuesta, 2003" authorityName="Schubart, Liu and Cuesta" authorityYear="2003" box="[700,1155,870,895]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="species" species="lintou">
<emphasis id="0B80EADAFFCABE426D8FFCF80C709C57" box="[700,800,870,895]" italics="true" pageId="32" pageNumber="476">Sc. lintou</emphasis>
<bibRefCitation id="5D654B39FFCABE426C14FCF80BD39C57" author="Schubart CD &amp; Liu HC &amp; Cuesta JA" box="[807,1155,870,895]" pageId="32" pageNumber="476" pagination="49 - 59" refId="ref46378" refString="Schubart CD, Liu HC, Cuesta JA. 2003. A new genus and species of tree-climbing crab (Crustacea: Brachyura: Sesarmidae) from Taiwan with notes on its ecology and larval morphology. Raffles Bull Zool. 51 (1): 49 - 59." type="journal article" year="2003">Schubart, Liu and Cuesta, 2003</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="394B36C8FFCABE426F93FC170BD49CED" blockId="32.[160,1156,836,965]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FC170BD49CED" pageId="32" pageNumber="476">
– Ambulatory legs relatively shorter (P4 merus 0.76–0.78 times carapace length).. ....... ............................................................................................
<taxonomicName id="FEF44D4BFFCABE426DE8FC320C369CED" ID-CoL="84H9N" baseAuthorityName="H.Milne Edwards" baseAuthorityYear="1853" box="[731,870,940,965]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="0B80EADAFFCABE426DE8FC320C369CED" box="[731,870,940,965]" italics="true" pageId="32" pageNumber="476">Sc. gracilipes</emphasis>
</taxonomicName>
(H.
<bibRefCitation id="5D654B39FFCABE426CA0FC320B2C9CED" author="Milne Edwards H" box="[915,1148,940,965]" pageId="32" pageNumber="476" pagination="163 - 228" refId="ref43918" refString="Milne Edwards H. 1853. Memoire sur la Famille des Ocypodiens. Ann Scie Nat Zool. 20 (3): 163 - 228, pls. 6 - 11." type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="82002E6DFFCABE426F93FC7F0BD39B4A" pageId="32" pageNumber="476">
<paragraph id="394B36C8FFCABE426F93FC7F0BD49B35" blockId="32.[160,1156,993,1122]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FC7F0BD49B35" pageId="32" pageNumber="476">
4. Upper surface of male cheliped palm without longitudinally traversing row of granules; G1 distal corneous process wide.. ................................ ...............................
<taxonomicName id="FEF44D4BFFCABE426CA4FB9A0B759B35" ID-CoL="84H9P" authority="Naruse &amp; Ng, 2020" authorityName="Naruse &amp; Ng" authorityYear="2020" box="[919,1061,1028,1053]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="species" species="malagasy" status="sp. nov.">
<emphasis id="0B80EADAFFCABE426CA4FB9A0B759B35" box="[919,1061,1028,1053]" italics="true" pageId="32" pageNumber="476">Sc. malagasy</emphasis>
</taxonomicName>
<taxonomicNameLabel id="10B357A1FFCABE426B1EFB9A0BD49B35" box="[1069,1156,1028,1053]" pageId="32" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="394B36C8FFCABE426F93FBB80BD39B4A" blockId="32.[160,1156,993,1122]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FBB80BD39B4A" pageId="32" pageNumber="476">– Upper surface of male cheliped palm with longitudinally traversing row of granules; G1 distal corneous process narrow.. ........................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="82002E6DFFCABE426F93FBE00BD39A6D" pageId="32" pageNumber="476">
<paragraph id="394B36C8FFCABE426F93FBE00BD39BF5" blockId="32.[160,1156,1150,1349]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FBE00BD39BF5" pageId="32" pageNumber="476">
5. Upper surface of male cheliped palm with more widely spaced row of granules; granules on inner side of upper margin of male cheliped movable finger acute.. .............. .....................................................................................................................................
<taxonomicName id="FEF44D4BFFCABE426CB8FB5A0BD39BF5" ID-CoL="6Y3Y9" authority="Ng, 2013" authorityName="Ng" authorityYear="2013" box="[907,1155,1220,1245]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="species" species="raymondi">
<emphasis id="0B80EADAFFCABE426CB8FB5A0B489BF5" box="[907,1048,1220,1245]" italics="true" pageId="32" pageNumber="476">Sc. raymondi</emphasis>
<bibRefCitation id="5D654B39FFCABE426B2CFB5A0BD39BF5" author="Ng PKL" box="[1055,1155,1220,1245]" pageId="32" pageNumber="476" pagination="289 - 294" refId="ref44391" refString="Ng PKL. 2013. A new species of Scandarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Borneo, Malaysia. Zootaxa. 3652 (2): 289 - 294. doi: 10.11646 / zootaxa. 3652.2.6" type="journal article" year="2013">Ng, 2013</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="394B36C8FFCABE426F93FB780BD39A6D" blockId="32.[160,1156,1150,1349]" pageId="32" pageNumber="476">
<keyLead id="820595FDFFCABE426F93FB780BD39A6D" pageId="32" pageNumber="476">
– Upper surface of male cheliped palm with more closely spaced row of granules; granules on inner side of upper margin of male cheliped movable finger rounded... ................................................................................................
<taxonomicName id="FEF44D4BFFCABE426DDEFAB20BD39A6D" ID-CoL="6Y49X" authority="Naruse and Ng, 2007" authorityName="Naruse and Ng" authorityYear="2007" box="[749,1155,1324,1349]" class="Malacostraca" family="Sesarmidae" genus="Scandarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="476" phylum="Arthropoda" rank="species" species="splendidum">
<emphasis id="0B80EADAFFCABE426DDEFAB20CC09A6D" box="[749,912,1324,1349]" italics="true" pageId="32" pageNumber="476">Sc. splendidum</emphasis>
<bibRefCitation id="5D654B39FFCABE426CABFAB20BD39A6D" author="Naruse T &amp; Ng PKL" box="[920,1155,1324,1349]" pageId="32" pageNumber="476" pagination="337 - 341" refId="ref44118" refString="Naruse T, Ng PKL. 2007. Scandarma splendidum, a new species of tree-climbing crab (Decapoda: Brachyura: Sesarmidae) from Sarawak, Malaysia, Borneo. Raffles Bull Zool. 55 (2): 337 - 341." type="journal article" year="2007">Naruse and Ng, 2007</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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