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<mods:title id="1C91F4015D2637D590563807E912804B">A revision of the genus Pachygrapsus Randall, 1840 (Crustacea: Decapoda: Brachyura, Grapsidae), with special reference to the Southwest Pacific species</mods:title>
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<bibRefCitation id="80054B73ED66FFD96F29FC3DFD34FC6D" author="De Man, J. G." box="[573,747,993,1019]" pageId="33" pageNumber="34" pagination="215 - 600" refId="ref31574" refString="De Man, J. G. (1888) Bericht uber die im indischen Archipel von Dr. J. Brock gesammelten, Decapoden und Stomatopoden. Archiv fur Naturgeschichte, 53, 215 - 600, pl. 7 - 22." type="journal article" year="1888">De Man, 1888</bibRefCitation>
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<figureCitation id="7CAF2A07ED66FFD96FEFFC3DFC77FC6D" box="[763,936,993,1019]" captionStart="FIGURE 10" captionStartId="35.[264,368,1718,1742]" captionTargetBox="[278,1311,690,1697]" captionTargetId="figure-144@35.[264,1323,673,1714]" captionTargetPageId="35" captionText="FIGURE 10. —Pachygrapsus planifrons de Man, 1888. Male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490): a) dorsal view; b) front; c) left chela, outer face; d) right P2; e) right P5; f) abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049252" httpUri="https://zenodo.org/record/5049252/files/figure.png" pageId="33" pageNumber="34">Figures 10a–f</figureCitation>
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<figureCitation id="7CAF2A07ED66FFD96EA1FC3DFC3BFC6D" box="[949,996,993,1019]" captionStart="FIGURE 14" captionStartId="47.[264,368,1403,1427]" captionTargetBox="[274,1309,325,1344]" captionTargetId="figure-211@47.[265,1322,294,1379]" captionTargetPageId="47" captionText="FIGURE 14. —Infra–orbital margin: a) Pachygrapsus fakaravensis Rathbun (1907), ov. female 15.0 x 16.5 mm, Taiaro, French Polynesia (MNHN B25782); b) Pachygrapsus laevimanus Stimpson, 1858, neotype male 18.0 x 21.7 mm, Sydney, Australia (AM P7478); c) Pachygrapsus minutus A. MilneEdwards, 1873, male 5.2 x 7.0 mm, Moruroa, French Polynesia (MNHN B29273); d) Pachygrapsus planifrons de Man, 1888, male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), male 12.1 x 15.3 mm, Taiaro, French Polynesia (MNHN B29279); f) Pachygrapsus corrugatus (von Martens, 1872) ov. female 13.6 x 15.2 mm, Virgin Is. (USNM 72340); g) Pachygrapsus crassipes Randall, 1840, male 31.9 x 36.6 mm, Sagami Bay, Japan (NSMT Cr7516); h) Pachygrapsus gracilis (de Saussure, 1858), ov. female 8.5 x 11 mm, Angola (MNHN B12903); i) Pachygrapsus loveridgei Chace, 1966, male 8.1 x 10 mm, St. Helena I. (MNHN B17836); j) Pachygrapsus marmoratus (Fabricius, 1787), female 12.7 x 14.3 mm, Préfailles, France (MNHN B22654); k) Pachygrapsus maurus (Lucas, 1846), female 12.7 x 14.3 mm, Banyuls, France (MNHN B12933); l) Pachygrapsus transversus (Gibbes, 1850), male 17.0 x 22.5 mm, French Guiana (MNHN B16013). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049262" httpUri="https://zenodo.org/record/5049262/files/figure.png" pageId="33" pageNumber="34">14d</figureCitation>
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<figureCitation id="7CAF2A07ED66FFD96EE6FC3DFBFFFC6D" box="[1010,1056,993,1019]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. MilneEdwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, g–l, from Chace, 1966, fig. 10–11; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="33" pageNumber="34">15d</figureCitation>
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, pl. 16, fig. 2 [
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<bibRefCitation id="80054B73ED66FFD96EE5FB9DFB1CFBCE" author="De Man, J. G." box="[1009,1219,1089,1112]" pageId="33" pageNumber="34" pagination="211 - 231" refId="ref31618" refString="De Man, J. G. (1908) The fauna of brackish ponds at Port Canning, Lower Bengal. Records of the Indian Museum, 2 (3), 211 - 231, pl. 18 - 19." type="journal article" year="1908">De Man, 1908: 218</bibRefCitation>
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status). —
<treatmentCitation id="65351093ED66FFD96F55FBBDFD37FBEE" author="Tesch, J. J." box="[577,744,1121,1144]" page="77" pageId="33" pageNumber="34" year="1918">
<bibRefCitation id="80054B73ED66FFD96F55FBBDFD37FBEE" author="Tesch, J. J." box="[577,744,1121,1144]" pageId="33" pageNumber="34" pagination="1 - 148" refId="ref34227" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga expedition. I. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae, and Gecarcinidae. Siboga - Expeditie, 39 c, 1 - 148, pl. 1 - 6." type="journal article" year="1918">Tesch, 1918: 77</bibRefCitation>
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(
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; synonymy). —
<treatmentCitation id="65351093ED66FFD96919FBBDFB6FFBEE" author="Ward, M." box="[1037,1200,1121,1144]" page="25" pageId="33" pageNumber="34" year="1934">
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(Christmas I.). —
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<bibRefCitation id="80054B73ED66FFD96C6DFB5DFDE0FB0E" author="Tweedie, M. W. F." box="[377,575,1153,1176]" pageId="33" pageNumber="34" pagination="44 - 70" refId="ref34274" refString="Tweedie, M. W. F. (1936) On the crabs of the family Grapsidae in the collections of the Raffles Museum. Bulletin of the Raffles Museum, 12, 44 - 70." type="journal article" year="1936">Tweedie, 1936: 45</bibRefCitation>
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(Christmas I.); 1950b: 130 (CocosKeeling Is.). —
<treatmentCitation id="65351093ED66FFD96972FB5DFAF2FB0E" author="Holthuis, L. B." box="[1126,1325,1153,1176]" page="31" pageId="33" pageNumber="34" year="1953">
<bibRefCitation id="80054B73ED66FFD96972FB5DFAF2FB0E" author="Holthuis, L. B." box="[1126,1325,1153,1176]" pageId="33" pageNumber="34" pagination="1 - 66" refId="ref30928" refString="Holthuis, L. B. (1953) Enumeration of the Decapod and Stomatopod Crustacea from Pacific coral islands. Atoll Research Bulletin, 24, 1 - 66." type="journal article" year="1953">Holthuis, 1953: 31</bibRefCitation>
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(
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; Tuamotu Archip.). —Edmonson, 1959: 173, fig. 10b (Hawaiian Is.; synonymy). —
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<bibRefCitation id="80054B73ED66FFD96C2CFB1DFE39FB4E" author="Garth, J. S." box="[312,486,1217,1240]" pageId="33" pageNumber="34" pagination="1 - 46" refId="ref30463" refString="Garth, J. S. (1965) The brachyuran Decapod Crustaceans of Clipperton Island. Proceedings of the California Academy of Sciences, fourth series, 33 (1), 1 - 46." type="journal article" year="1965">Garth, 1965: 30</bibRefCitation>
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, fig. 15–16 (
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<treatmentCitation id="65351093ED66FFD96E0FFB1DFC04FB4E" author="Chace, F. A." box="[795,987,1217,1240]" page="644" pageId="33" pageNumber="34" year="1966">
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, fig. 11f (Hawaiian Is.; synonymy). —
<treatmentCitation id="65351093ED66FFD96C88FB3DFD8BFB6E" author="Chen, H." box="[412,596,1249,1272]" page="139" pageId="33" pageNumber="34" year="1980">
<bibRefCitation id="80054B73ED66FFD96C88FB3DFD8BFB6E" author="Chen, H." box="[412,596,1249,1272]" pageId="33" pageNumber="34" pagination="117 - 147" refId="ref29360" refString="Chen, H. (1980) Studies on the crabs of the Xisha Islands, Guangdong Province, China. II. Studia Marina Sinica, 17, 117 - 147, pl. 1 - 4 (in Chinese)." type="journal article" year="1980">Chen, 1980: 139</bibRefCitation>
</treatmentCitation>
: fig. 22, pl. IV3 (Paracel Is.). —
<treatmentCitation id="65351093ED66FFD96ED0FB3DFB18FB6E" author="Dai, A. &amp; Yang, S." box="[964,1223,1249,1272]" page="510" pageId="33" pageNumber="34" year="1991">
<bibRefCitation id="80054B73ED66FFD96ED0FB3DFB18FB6E" author="Dai, A. &amp; Yang, S." box="[964,1223,1249,1272]" pageId="33" pageNumber="34" refId="ref29508" refString="Dai, A. &amp; Yang, S. (1991) Crabs of the China seas. China Ocean Press Beijing; Springer-Verlag Berlin, Heidelberg, New York, Tokyo (English Edition), 682 pp." type="book" year="1991">Dai &amp; Yang, 1991: 510</bibRefCitation>
</treatmentCitation>
, fig. 261 (Hainan). —
<treatmentCitation id="65351093ED66FFD96CD4FADDFD61FA8E" author="Ng, P. K. L. &amp; Davie, P. J. F." box="[448,702,1281,1304]" page="379" pageId="33" pageNumber="34" year="2002">
<bibRefCitation id="80054B73ED66FFD96CD4FADDFD61FA8E" author="Ng, P. K. L. &amp; Davie, P. J. F." box="[448,702,1281,1304]" pageId="33" pageNumber="34" pagination="369 - 384" refId="ref32346" refString="Ng, P. K. L. &amp; Davie, P. J. F. (2002) A checklist of the brachyuran crabs of Phuket and western Thailand. Phuket Marine Biological Center Special Publication, 23 (2), 369 - 384." type="journal article" year="2002">Ng &amp; Davie, 2002: 379</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="9C837612ED66FFD96FD8FADDFCEEFA8E" box="[716,817,1281,1304]" name="Thailand" pageId="33" pageNumber="34">Thailand</collectingCountry>
). —
<treatmentCitation id="65351093ED66FFD96E4BFADDFB80FA8E" author="Paulay, G. &amp; Kropp, R. &amp; Ng, P. K. L. &amp; Eldredge, L. G." box="[863,1119,1281,1305]" page="495" pageId="33" pageNumber="34" year="2003">
<bibRefCitation id="80054B73ED66FFD96E4BFADDFB80FA8E" author="Paulay, G. &amp; Kropp, R. &amp; Ng, P. K. L. &amp; Eldredge, L. G." box="[863,1119,1281,1305]" pageId="33" pageNumber="34" pagination="456 - 513" refId="ref32663" refString="Paulay, G., Kropp, R., Ng, P. K. L. &amp; Eldredge, L. G. (2003) The crustaceans and pycnogonids of the Mariana Islands. Micronesica, 35 - 36, 456 - 513." type="journal article" year="2003">
Paulay
<emphasis id="D6E0EA90ED66FFD96EBAFADEFC02FA8F" box="[942,989,1282,1305]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
., 2003: 495
</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="9C837612ED66FFD9697AFADDFB6AFA8E" box="[1134,1205,1281,1304]" name="Guam" pageId="33" pageNumber="34">Guam</collectingCountry>
). —
<treatmentCitation id="65351093ED66FFD969F7FADDFE06FAAE" author="Komai, T. &amp; Nagai, T. &amp; Yogi, A. &amp; Naruse, T. &amp; Fujita, Y. &amp; Shokita, S" page="34" pageId="33" pageNumber="34" year="2004">
<bibRefCitation id="80054B73ED66FFD969F7FADDFE06FAAE" author="Komai, T. &amp; Nagai, T. &amp; Yogi, A. &amp; Naruse, T. &amp; Fujita, Y. &amp; Shokita, S" pageId="33" pageNumber="34" pagination="33 - 63" refId="ref31238" refString="Komai, T., Nagai, T., Yogi, A., Naruse, T., Fujita, Y., &amp; Shokita, S (2004) New records of four grapsoid crabs (Crustacea: Decapoda: Brachyura) from Japan, with notes on four rare species. Natural History Research, 8 (1), 33 - 63." type="journal article" year="2004">
Komai
<emphasis id="D6E0EA90ED66FFD96C2CFAFEFEB1FAAF" box="[312,366,1314,1337]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
, 2004: 34
</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="9C837612ED66FFD96CF3FAFDFDF6FAAE" box="[487,553,1313,1336]" name="Japan" pageId="33" pageNumber="34">Japan</collectingCountry>
). —
<treatmentCitation id="65351093ED66FFD96F41FAFDFCF9FAAE" author="Yeo, D. C. J. &amp; Rahayu, D. &amp; Ng, P. K. L." box="[597,806,1313,1337]" page="88" pageId="33" pageNumber="34" year="2004">
<bibRefCitation id="80054B73ED66FFD96F41FAFDFCF9FAAE" author="Yeo, D. C. J. &amp; Rahayu, D. &amp; Ng, P. K. L." box="[597,806,1313,1337]" pageId="33" pageNumber="34" pagination="79 - 88" refId="ref34775" refString="Yeo, D. C. J., Rahayu, D. L &amp; Ng, P. K. L. (2004) Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles Bulletin of Zoology, Supplement 11, 79 - 88." type="journal article" year="2004">
Yeo
<emphasis id="D6E0EA90ED66FFD96F92FAFEFD6BFAAF" box="[646,692,1314,1337]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
., 2004: 88
</bibRefCitation>
</treatmentCitation>
(Anambas Is., South
<collectingCountry id="9C837612ED66FFD9691AFAFDFB91FAAE" box="[1038,1102,1313,1336]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
Sea).
</treatmentCitationGroup>
</paragraph>
<paragraph id="E42B3682ED66FFD96C1CFA9EFBC3FAEE" blockId="33.[264,1325,1057,1464]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="C48411ACED66FFD96C1CFA9EFBC3FAEE" pageId="33" pageNumber="34">
<treatmentCitation id="65351093ED66FFD96C1CFA9EFC5BFACE" author="Rathbun, M. J." box="[264,900,1345,1369]" page="247" pageId="33" pageNumber="34" year="1893">
<taxonomicName id="23944D01ED66FFD96C1CFA9EFC5BFACE" ID-CoL="4BSC8" authority="Rathbun, 1893: 247 (Hawaiian Is.)" authorityName="Rathbun" authorityYear="1893" box="[264,900,1345,1369]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="D6E0EA90ED66FFD96C1CFA9EFDDEFACF" box="[264,513,1346,1369]" italics="true" pageId="33" pageNumber="34">Pachygrapsus longipes</emphasis>
<bibRefCitation id="80054B73ED66FFD96F1EFA9DFD3FFACE" author="Rathbun, M. J." box="[522,736,1345,1368]" pageId="33" pageNumber="34" pagination="223 - 260" refId="ref33152" refString="Rathbun, M. J. (1893) Scientific results of explorations by the U. S. Fish Commission steamer Albatross. XXIV, Description of new genera and species of crabs from the west coast of North America and the Sandwich Islands. Proceedings of the U. S. National Museum, 16, 223 - 260." type="journal article" year="1893">Rathbun, 1893: 247</bibRefCitation>
(Hawaiian Is.)
</taxonomicName>
</treatmentCitation>
. —
<treatmentCitation id="65351093ED66FFD96EB8FA9DFB5EFACE" author="Rathbun, M. J." box="[940,1153,1345,1368]" page="840" pageId="33" pageNumber="34" year="1906">
<bibRefCitation id="80054B73ED66FFD96EB8FA9DFB5EFACE" author="Rathbun, M. J." box="[940,1153,1345,1368]" pageId="33" pageNumber="34" pagination="827 - 930" refId="ref33214" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pl. 1 - 24." type="journal article" year="1906">Rathbun, 1906: 840</bibRefCitation>
</treatmentCitation>
(Hawaiian Is.); 1907: 30 (Fakarava Atoll, Tuamotu Archip.); 1911: 242 (
<collectingCountry id="9C837612ED66FFD96E8CFABDFBCFFAEE" box="[920,1040,1377,1400]" name="Seychelles" pageId="33" pageNumber="34">Seychelles</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="E42B3682ED66FFD96C1CFA5EFDF2FA2E" blockId="33.[264,1325,1057,1464]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="C48411ACED66FFD96C1CFA5EFDF2FA2E" pageId="33" pageNumber="34">
<treatmentCitation id="65351093ED66FFD96C1CFA5EFD07FA0E" author="Borradaile, L. A." box="[264,728,1409,1433]" page="592" pageId="33" pageNumber="34" year="1900">
<taxonomicName id="23944D01ED66FFD96C1CFA5EFD07FA0E" ID-CoL="4BSC7" authority="Borradaile, 1900: 592" authorityName="Borradaile" authorityPageNumber="592" authorityYear="1900" box="[264,728,1409,1433]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="D6E0EA90ED66FFD96C1CFA5EFE3BFA0F" box="[264,484,1410,1433]" italics="true" pageId="33" pageNumber="34">Pachygrapsus laevis</emphasis>
<bibRefCitation id="80054B73ED66FFD96CF8FA5DFD07FA0E" author="Borradaile, L. A." box="[492,728,1409,1432]" pageId="33" pageNumber="34" pagination="568 - 596" refId="ref28916" refString="Borradaile, L. A. (1900) On some Crustacea from the South Pacific. Part IV. The Crabs. Proceedings of the Zoological Society, London, 568 - 596." type="book chapter" year="1900">Borradaile, 1900: 592</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 42, fig. 7 (
<collectingCountry id="9C837612ED66FFD96E62FA5DFC1AFA0E" box="[886,965,1409,1432]" name="Tuvalu" pageId="33" pageNumber="34">Tuvalu</collectingCountry>
; new synonymy see remarks). —
<treatmentCitation id="65351093ED66FFD96C2CFA7DFE3FFA2E" author="Tesch, J. J." box="[312,480,1441,1464]" page="76" pageId="33" pageNumber="34" year="1918">
<bibRefCitation id="80054B73ED66FFD96C2CFA7DFE3FFA2E" author="Tesch, J. J." box="[312,480,1441,1464]" pageId="33" pageNumber="34" pagination="1 - 148" refId="ref34227" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga expedition. I. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae, and Gecarcinidae. Siboga - Expeditie, 39 c, 1 - 148, pl. 1 - 6." type="journal article" year="1918">Tesch, 1918: 76</bibRefCitation>
</treatmentCitation>
(Key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC8E6509ED66FFD96C1CFA03FC36F8CD" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="E42B3682ED66FFD96C1CFA03FE76FA6E" blockId="33.[264,1322,1503,1729]" box="[264,425,1503,1528]" pageId="33" pageNumber="34">
<emphasis id="D6E0EA90ED66FFD96C1CFA03FE76FA6E" box="[264,425,1503,1528]" italics="true" pageId="33" pageNumber="34">
<typeStatus id="3B2F8820ED66FFD96C1CFA03FEE0FA6E" box="[264,319,1503,1528]" pageId="33" pageNumber="34">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="E42B3682ED66FFD96C2CF9DBFAC6F957" blockId="33.[264,1322,1503,1729]" pageId="33" pageNumber="34">
Presumably lost. According to C. H. Fransen (pers. comm.) the
<typeStatus id="3B2F8820ED66FFD9693AF9DBFB4BF9B7" box="[1070,1172,1543,1569]" pageId="33" pageNumber="34" type="syntype">syntypes</typeStatus>
were probably deposited at Göttingen (
<collectionCode id="8285AE47ED66FFD96F47F9F3FD45F9DF" box="[595,666,1583,1609]" country="Germany" httpUri="http://grbio.org/cool/gs4h-1fee" name="Zoologischen Museums Greifswald" pageId="33" pageNumber="34">ZMG</collectionCode>
), but are not found there (G. Tröster, pers. comm.), nor are they part of the material that was transferred from Göttingen to Frankfurt (
<collectionCode id="8285AE47ED66FFD969B2F98BFB39F9E7" box="[1190,1254,1623,1649]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="33" pageNumber="34">SMF</collectionCode>
). The Göttingen collection was in poor condition before being transferred to Frankfurt, and a number of types of other species also appear to have been lost (M. Türkay, pers. comm.).
</paragraph>
<paragraph id="E42B3682ED66FFD96C1CF92DFE34F89C" blockId="33.[264,1322,1777,1883]" box="[264,491,1777,1802]" pageId="33" pageNumber="34">
<emphasis id="D6E0EA90ED66FFD96C1CF92DFE34F89C" box="[264,491,1777,1802]" italics="true" pageId="33" pageNumber="34">Material Examined</emphasis>
</paragraph>
<paragraph id="E42B3682ED66FFD96C2CF8C5FC36F8CD" blockId="33.[264,1322,1777,1883]" pageId="33" pageNumber="34">
<emphasis id="D6E0EA90ED66FFD96C2CF8C5FE11F8A5" bold="true" box="[312,462,1817,1843]" pageId="33" pageNumber="34">Mariana Is.</emphasis>
<materialsCitation id="54FC3CDFED66FFD96CCDF8C5FC3AF8CD" collectingDate="1998-03-29" collectionCode="UF" collectorName="G. Paulay" country="Guam" location="Bile Bay" pageId="33" pageNumber="34" specimenCode="UF 129" specimenCount="1">
<emphasis id="D6E0EA90ED66FFD96CCDF8C5FDF8F8A5" bold="true" box="[473,551,1817,1843]" pageId="33" pageNumber="34">
<collectingCountry id="9C837612ED66FFD96CCDF8C5FDF8F8A5" box="[473,551,1817,1843]" name="Guam" pageId="33" pageNumber="34">Guam</collectingCountry>
</emphasis>
:
<location id="E14B6059ED66FFD96F2EF8C5FD79F8A5" LSID="urn:lsid:plazi:treatment:6C3D8794ED66FFDC6C1CFC3DFE7CF9C8:E14B6059ED66FFD96F2EF8C5FD79F8A5" box="[570,678,1817,1843]" country="Guam" name="Bile Bay" pageId="33" pageNumber="34">Bile Bay</location>
, supratidal/intertidal, among rocks, common, coll.
<collectorName id="49615354ED66FFD96805F8C5FE86F8CD" pageId="33" pageNumber="34">G. Paulay</collectorName>
,
<date id="902A1042ED66FFD96C71F89DFDC5F8CD" box="[357,538,1857,1883]" pageId="33" pageNumber="34" value="1998-03-29">
<collectingDate id="806EE9AAED66FFD96C71F89DFDC5F8CD" box="[357,538,1857,1883]" pageId="33" pageNumber="34" value="1998-03-29">29 March 1998</collectingDate>
</date>
, 2 M 5.1 x 6.3, 5.6 x
<quantity id="236C9B67ED66FFD96E01F89DFCB0F8CD" box="[789,879,1857,1883]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="33" pageNumber="34" unit="mm" value="6.8">6.8 mm</quantity>
(
<specimenCode id="B4329EF9ED66FFD96E6AF89DFC03F8CD" box="[894,988,1857,1883]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="33" pageNumber="34" type="Museum">UF 129</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E6509ED65FFDA6C2CFEC7FBEDFEC8" pageId="34" pageNumber="35" type="description">
<paragraph id="E42B3682ED65FFDA6C2CFEC7FBEDFEC8" blockId="34.[264,1324,283,990]" pageId="34" pageNumber="35">
<emphasis id="D6E0EA90ED65FFDA6C2CFEC7FE10FEA3" bold="true" box="[312,463,283,309]" pageId="34" pageNumber="35">Caroline Is.</emphasis>
<materialsCitation id="54FC3CDFED65FFDA6CCFFEC7FBF1FEC8" collectingDate="2000-04-01" collectionCode="UF, GP" collectorName="Maap I. &amp; G. Paulay" location="Yap" pageId="34" pageNumber="35" specimenCode="UF 3849, GP805" specimenCount="1" stateProvince="Yap">
<emphasis id="D6E0EA90ED65FFDA6CCFFEC7FDD3FEA3" bold="true" box="[475,524,283,309]" pageId="34" pageNumber="35">
<collectingRegion id="2650F860ED65FFDA6CCFFEC7FDD3FEA3" box="[475,524,283,309]" country="Micronesia" name="Yap" pageId="34" pageNumber="35">Yap</collectingRegion>
</emphasis>
:
<collectorName id="49615354ED65FFDA6F34FEC0FD5DFEA0" box="[544,642,284,310]" pageId="34" pageNumber="35">Maap I.</collectorName>
, high intertidal, under rocks and plants debris, coll.
<collectorName id="49615354ED65FFDA6806FEC0FE86FEC8" pageId="34" pageNumber="35">G. Paulay</collectorName>
,
<date id="902A1042ED65FFDA6C71FE98FE23FEC8" box="[357,508,324,350]" pageId="34" pageNumber="35" value="2000-04-01">
<collectingDate id="806EE9AAED65FFDA6C71FE98FE23FEC8" box="[357,508,324,350]" pageId="34" pageNumber="35" value="2000-04-01">1 April 2000</collectingDate>
</date>
, 1 ov. F 5.2 x
<quantity id="236C9B67ED65FFDA6FB6FE98FD24FEC8" box="[674,763,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="34" pageNumber="35" unit="mm" value="6.7">6.7 mm</quantity>
(
<specimenCode id="B4329EF9ED65FFDA6E1FFE98FCA7FEC8" box="[779,888,324,350]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="34" pageNumber="35" type="Museum">UF 3849</specimenCode>
; Photo
<specimenCode id="B4329EF9ED65FFDA6EDBFE98FBF9FEC8" box="[975,1062,324,350]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="34" pageNumber="35">GP805</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E6509ED65FFDC6C2CFEB7FE7CF9C8" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="E42B3682ED65FFDA6C2CFEB7FCBAFE38" blockId="34.[264,1324,283,990]" pageId="34" pageNumber="35">
<materialsCitation id="54FC3CDFED65FFDA6C2CFEB7FCBEFE38" collectingDate="1897-11-01" collectionCode="UMZC" collectorName="J. S. Gardiner" country="Tuvalu" location="Funafuti" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Funafuti" typeStatus="holotype">
<emphasis id="D6E0EA90ED65FFDA6C2CFEB7FDD5FE13" bold="true" box="[312,522,363,389]" pageId="34" pageNumber="35">
<collectingCountry id="9C837612ED65FFDA6C2CFEB7FE4EFE13" box="[312,401,363,389]" name="Tuvalu" pageId="34" pageNumber="35">Tuvalu</collectingCountry>
.
<collectingRegion id="2650F860ED65FFDA6C89FEB7FDD5FE13" box="[413,522,363,389]" country="Tuvalu" name="Funafuti" pageId="34" pageNumber="35">Funafuti</collectingRegion>
</emphasis>
: coll.
<collectorName id="49615354ED65FFDA6F41FEB0FD26FE10" box="[597,761,364,390]" pageId="34" pageNumber="35">J. S. Gardiner</collectorName>
, 1 ov. F 5.1 x
<quantity id="236C9B67ED65FFDA6EB5FEB0FC23FE10" box="[929,1020,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="34" pageNumber="35" unit="mm" value="7.2">7.2 mm</quantity>
(
<collectionCode id="8285AE47ED65FFDA691FFEB0FBBDFE10" box="[1035,1122,364,390]" pageId="34" pageNumber="35">UMZC</collectionCode>
Reg.
<date id="902A1042ED65FFDA698AFEB0FAF8FE10" box="[1182,1319,364,390]" pageId="34" pageNumber="35" value="1897-11-01">
<collectingDate id="806EE9AAED65FFDA698AFEB0FAF8FE10" box="[1182,1319,364,390]" pageId="34" pageNumber="35" value="1897-11-01">11.01.1897</collectingDate>
</date>
;
<typeStatus id="3B2F8820ED65FFDA6C1CFE48FEB1FE38" box="[264,366,404,430]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
of
<taxonomicName id="23944D01ED65FFDA6C82FE48FC87FE38" authority="Borradaile, 1900" authorityName="Borradaile" authorityYear="1900" box="[406,856,404,430]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="D6E0EA90ED65FFDA6C82FE48FD58FE3B" box="[406,647,404,429]" italics="true" pageId="34" pageNumber="35">Pachygrapsus laevis</emphasis>
<bibRefCitation id="80054B73ED65FFDA6F9BFE48FC87FE38" author="Borradaile, L. A." box="[655,856,404,430]" pageId="34" pageNumber="35" pagination="568 - 596" refId="ref28916" refString="Borradaile, L. A. (1900) On some Crustacea from the South Pacific. Part IV. The Crabs. Proceedings of the Zoological Society, London, 568 - 596." type="book chapter" year="1900">Borradaile, 1900</bibRefCitation>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E42B3682ED65FFDA6C2CFE67FD9DFD08" blockId="34.[264,1324,283,990]" pageId="34" pageNumber="35">
<materialsCitation id="54FC3CDFED65FFDA6C2CFE67FD00FE68" collectingDate="1899-10-12" collectionCode="USNM" collectorName="RV Albatross" country="French Polynesia" location="Fakarava Atoll" pageId="34" pageNumber="35" specimenCode="USNM 33179" specimenCount="1" stateProvince="Tuamotu Archip.">
<emphasis id="D6E0EA90ED65FFDA6C2CFE67FD22FE43" bold="true" box="[312,765,443,469]" pageId="34" pageNumber="35">
<collectingCountry id="9C837612ED65FFDA6C2CFE67FDCEFE43" box="[312,529,443,469]" name="French Polynesia" pageId="34" pageNumber="35">French Polynesia</collectingCountry>
.
<collectingRegion id="2650F860ED65FFDA6F09FE67FD28FE43" box="[541,759,443,469]" pageId="34" pageNumber="35">Tuamotu Archip.</collectingRegion>
:
</emphasis>
<location id="E14B6059ED65FFDA6E11FE60FC69FE40" LSID="urn:lsid:plazi:treatment:6C3D8794ED66FFDC6C1CFC3DFE7CF9C8:E14B6059ED65FFDA6E11FE60FC69FE40" box="[773,950,444,470]" country="French Polynesia" name="Fakarava Atoll" pageId="34" pageNumber="35" stateProvince="Tuamotu Archip.">Fakarava Atoll</location>
, coll.
<collectorName id="49615354ED65FFDA6EE8FE60FB46FE43" box="[1020,1177,444,470]" pageId="34" pageNumber="35">
RV
<emphasis id="D6E0EA90ED65FFDA693EFE60FB46FE43" box="[1066,1177,444,469]" italics="true" pageId="34" pageNumber="35">Albatross</emphasis>
</collectorName>
,
<date id="902A1042ED65FFDA69B3FE60FE9AFE68" pageId="34" pageNumber="35" value="1899-10-12">
<collectingDate id="806EE9AAED65FFDA69B3FE60FE9AFE68" pageId="34" pageNumber="35" value="1899-10-12">12 October 1899</collectingDate>
</date>
, 1 F 3.6 x
<quantity id="236C9B67ED65FFDA6CD6FE38FDC4FE68" box="[450,539,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="34" pageNumber="35" unit="mm" value="4.9">4.9 mm</quantity>
(
<specimenCode id="B4329EF9ED65FFDA6F3FFE38FD08FE68" box="[555,727,484,510]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="34" pageNumber="35" type="Museum">USNM 33179</specimenCode>
)
</materialsCitation>
. —
<materialsCitation id="54FC3CDFED65FFDA6E1DFE38FBB6FDD8" collectingDate="2001-01-10" collectionCode="UF" collectorName="G. Paulay" country="French Polynesia" county="Rangiroa Atoll" elevation="5" location="Pension Henri" municipality="Avatoru Motu" pageId="34" pageNumber="35" specimenCode="UF 1490" specimenCount="1" stateProvince="Tuamotu Archip.">
<collectingCounty id="0D4A4E0EED65FFDA6E1DFE38FC65FE68" box="[777,954,484,510]" pageId="34" pageNumber="35">Rangiroa Atoll</collectingCounty>
,
<collectingMunicipality id="044FACF8ED65FFDA6ED2FE38FBB1FE68" box="[966,1134,484,510]" pageId="34" pageNumber="35">Avatoru Motu</collectingMunicipality>
, ca. at
<locationDeviation id="F5992FA0ED65FFDA69ABFE38FE71FDB0" location="Post Office" pageId="34" pageNumber="35">
<quantity id="236C9B67ED65FFDA69ABFE38FACEFE68" box="[1215,1297,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="km" value="1.5">1.5 km</quantity>
E of Post Office
</locationDeviation>
, at
<location id="E14B6059ED65FFDA6CC2FDD0FDA0FDB0" LSID="urn:lsid:plazi:treatment:6C3D8794ED66FFDC6C1CFC3DFE7CF9C8:E14B6059ED65FFDA6CC2FDD0FDA0FDB0" box="[470,639,524,550]" country="French Polynesia" county="Rangiroa Atoll" municipality="Avatoru Motu" name="Pension Henri" pageId="34" pageNumber="35" stateProvince="Tuamotu Archip.">Pension Henri</location>
, reef flat, under rocks,
<quantity id="236C9B67ED65FFDA6E9FFDD0FBDCFDB0" box="[907,1027,524,550]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="34" pageNumber="35" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">
<elevation id="6FB9D1B1ED65FFDA6E9FFDD0FBDCFDB0" box="[907,1027,524,550]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="34" pageNumber="35" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 meter</elevation>
</quantity>
, coll.
<collectorName id="49615354ED65FFDA695CFDD0FB66FDB0" box="[1096,1209,524,550]" pageId="34" pageNumber="35">G. Paulay</collectorName>
,
<date id="902A1042ED65FFDA69D1FDD0FEACFDD8" pageId="34" pageNumber="35" value="2001-01-10">
<collectingDate id="806EE9AAED65FFDA69D1FDD0FEACFDD8" pageId="34" pageNumber="35" value="2001-01-10">10 January 2001</collectingDate>
</date>
, 1 M 7.0 x 9.0 mm, 2 ov. F 5.4 x 7.0–5.6 x
<quantity id="236C9B67ED65FFDA6E91FDE8FC3EFDD8" box="[901,993,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="34" pageNumber="35" unit="mm" value="7.3">7.3 mm</quantity>
(
<specimenCode id="B4329EF9ED65FFDA6EE5FDE8FBBEFDD8" box="[1009,1121,564,590]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="34" pageNumber="35" type="Museum">UF 1490</specimenCode>
)
</materialsCitation>
. —
<materialsCitation id="54FC3CDFED65FFDA6987FDE8FDE2FD08" accessionNumber="B29264" collectingDate="1994-02-12" collectingDateMax="1994-02-20" collectingDateMin="1994-02-12" collectionCode="MNHN" collectorName="J. Poupin" country="French Polynesia" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Tuamotu Archip.">
Taiaro Atoll, Taiaro Expedition, intertidal, outer reef,
<date id="902A1042ED65FFDA6FC9FD80FC04FDE0" box="[733,987,604,630]" pageId="34" pageNumber="35" value="1994-02-12" valueMax="1994-02-20" valueMin="1994-02-12">
<collectingDate id="806EE9AAED65FFDA6FC9FD80FC04FDE0" box="[733,987,604,630]" pageId="34" pageNumber="35" value="1994-02-12" valueMax="1994-02-20" valueMin="1994-02-12">12–20 February 1994</collectingDate>
</date>
, coll.
<collectorName id="49615354ED65FFDA6934FD80FB50FDE0" box="[1056,1167,604,630]" pageId="34" pageNumber="35">J. Poupin</collectorName>
, 1 ov. F 4.6 x 6.0 mm (
<collectionCode id="8285AE47ED65FFDA6C64FD58FE14FD08" box="[368,459,644,670]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="35">MNHN</collectionCode>
B29264)
</materialsCitation>
.
</paragraph>
<paragraph id="E42B3682ED65FFDA6C2CFD70FD6FFC80" blockId="34.[264,1324,283,990]" pageId="34" pageNumber="35">
<emphasis id="D6E0EA90ED65FFDA6C2CFD70FE05FD50" bold="true" box="[312,474,684,710]" pageId="34" pageNumber="35">Hawaiian Is.</emphasis>
:
<materialsCitation id="54FC3CDFED65FFDA6CFAFD70FD73FC80" collectingDate="1902-06-06" collectionCode="USNM, UNSM" collectorName="Honolulu &amp; RV Albatross" pageId="34" pageNumber="35" specimenCode="USNM 17320, UNSM 29350" specimenCount="1">
<collectorName id="49615354ED65FFDA6CFAFD70FDBFFD50" box="[494,608,684,710]" pageId="34" pageNumber="35">Honolulu</collectorName>
, coll.
<collectorName id="49615354ED65FFDA6FBBFD70FC8FFD53" box="[687,848,684,710]" pageId="34" pageNumber="35">
RV
<emphasis id="D6E0EA90ED65FFDA6FF5FD70FC8FFD53" box="[737,848,684,709]" italics="true" pageId="34" pageNumber="35">Albatross</emphasis>
</collectorName>
, 1891, 1 ov. F 6.2 x
<quantity id="236C9B67ED65FFDA6976FD70FB1EFD50" box="[1122,1217,684,710]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="34" pageNumber="35" unit="mm" value="8.2">8.2 mm</quantity>
(
<specimenCode id="B4329EF9ED65FFDA69C0FD70FE8BFD78" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="34" pageNumber="35" type="Museum">USNM 17320</specimenCode>
,
<typeStatus id="3B2F8820ED65FFDA6C4BFD08FE4EFD78" box="[351,401,724,750]" pageId="34" pageNumber="35">type</typeStatus>
of
<taxonomicName id="23944D01ED65FFDA6CA3FD08FD32FD78" authority="Rathbun, 1893" authorityName="Rathbun" authorityYear="1893" box="[439,749,724,750]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="D6E0EA90ED65FFDA6CA3FD08FDE9FD7B" box="[439,566,724,749]" italics="true" pageId="34" pageNumber="35">P. longipes</emphasis>
<bibRefCitation id="80054B73ED65FFDA6F29FD08FD32FD78" author="Rathbun, M. J." box="[573,749,724,750]" pageId="34" pageNumber="35" pagination="223 - 260" refId="ref33152" refString="Rathbun, M. J. (1893) Scientific results of explorations by the U. S. Fish Commission steamer Albatross. XXIV, Description of new genera and species of crabs from the west coast of North America and the Sandwich Islands. Proceedings of the U. S. National Museum, 16, 223 - 260." type="journal article" year="1893">Rathbun, 1893</bibRefCitation>
</taxonomicName>
);
<collectorName id="49615354ED65FFDA6E15FD08FCAFFD78" box="[769,880,724,750]" pageId="34" pageNumber="35">Honolulu</collectorName>
reef, coll.
<collectorName id="49615354ED65FFDA6EF8FD08FB56FD7B" box="[1004,1161,724,750]" pageId="34" pageNumber="35">
RV
<emphasis id="D6E0EA90ED65FFDA690EFD08FB56FD7B" box="[1050,1161,724,749]" italics="true" pageId="34" pageNumber="35">Albatross</emphasis>
</collectorName>
,
<date id="902A1042ED65FFDA6983FD08FAF9FD78" box="[1175,1318,724,750]" pageId="34" pageNumber="35" value="1902-06-06">
<collectingDate id="806EE9AAED65FFDA6983FD08FAF9FD78" box="[1175,1318,724,750]" pageId="34" pageNumber="35" value="1902-06-06">6 June 1902</collectingDate>
</date>
, 1 M 9.5 x
<quantity id="236C9B67ED65FFDA6C95FD20FE36FC80" box="[385,489,764,790]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="34" pageNumber="35" unit="mm" value="11.8">11.8 mm</quantity>
(
<specimenCode id="B4329EF9ED65FFDA6CECFD20FD7CFC80" box="[504,675,764,790]" collectionCode="UNSM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="34" pageNumber="35">UNSM 29350</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E42B3682ED65FFDA6C2CFCF8FD5CFC48" blockId="34.[264,1324,283,990]" pageId="34" pageNumber="35">
<materialsCitation id="54FC3CDFED65FFDA6C2CFCF8FDA0FC48" accessionNumber="B29859, B29858" collectingDate="2005-01-25" collectingDateMax="2005-02" collectingDateMin="2005-01-25" collectionCode="MNHN" collectorName="J. Poupin &amp; L. Albenga &amp; Albenga, J. M" country="Clipperton Island" location="Clipperton" municipality="Jean-Louis Etienne Expedition" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Atoll.">
<emphasis id="D6E0EA90ED65FFDA6C2CFCF8FDD0FCA8" bold="true" box="[312,527,804,830]" pageId="34" pageNumber="35">
<collectingCountry id="9C837612ED65FFDA6C2CFCF8FE1FFCA8" box="[312,448,804,830]" name="Clipperton Island" pageId="34" pageNumber="35">Clipperton</collectingCountry>
<collectingRegion id="2650F860ED65FFDA6CDEFCF8FDD0FCA8" box="[458,527,804,830]" pageId="34" pageNumber="35">Atoll.</collectingRegion>
</emphasis>
Among rocks at low tide,
<collectingMunicipality id="044FACF8ED65FFDA6E41FCF8FB1DFCA8" box="[853,1218,804,830]" pageId="34" pageNumber="35">JeanLouis Etienne Expedition</collectingMunicipality>
to
<location id="E14B6059ED65FFDA69F9FCF8FE89FCF0" LSID="urn:lsid:plazi:treatment:6C3D8794ED66FFDC6C1CFC3DFE7CF9C8:E14B6059ED65FFDA69F9FCF8FE89FCF0" country="Clipperton Island" municipality="Jean-Louis Etienne Expedition" name="Clipperton" pageId="34" pageNumber="35" stateProvince="Atoll.">Clipperton</location>
, coll.
<collectorName id="49615354ED65FFDA6C8FFC90FDD5FCF0" box="[411,522,844,870]" pageId="34" pageNumber="35">J. Poupin</collectorName>
,
<date id="902A1042ED65FFDA6F01FC90FD1DFCF0" box="[533,706,844,870]" pageId="34" pageNumber="35" value="2005-02">
<collectingDate id="806EE9AAED65FFDA6F01FC90FD1DFCF0" box="[533,706,844,870]" pageId="34" pageNumber="35" value="2005-02">February 2005</collectingDate>
</date>
, 6 M 5.5 x 7.0–9.7 x
<quantity id="236C9B67ED65FFDA6EACFC90FBFDFCF0" box="[952,1058,844,870]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="34" pageNumber="35" unit="mm" value="11.5">11.5 mm</quantity>
, 10 ov. F 4.0 x 6.5–9.5 x 12.0 mm, 1 F 4.5 x
<quantity id="236C9B67ED65FFDA6F1CFCA8FDBDFC18" box="[520,610,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="34" pageNumber="35" unit="mm" value="5.5">5.5 mm</quantity>
(
<collectionCode id="8285AE47ED65FFDA6F66FCA8FD12FC18" box="[626,717,884,910]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="35">MNHN</collectionCode>
B29859); reef at low tide, stn 33, coll.
<collectorName id="49615354ED65FFDA698BFCA8FAF9FC18" box="[1183,1318,884,910]" pageId="34" pageNumber="35">L. Albenga</collectorName>
, J. M. Bouchard,
<date id="902A1042ED65FFDA6CDAFC40FD4CFC20" box="[462,659,924,950]" pageId="34" pageNumber="35" value="2005-01-25">
<collectingDate id="806EE9AAED65FFDA6CDAFC40FD4CFC20" box="[462,659,924,950]" pageId="34" pageNumber="35" value="2005-01-25">25 January 2005</collectingDate>
</date>
, 5 M 6.0 x 7.0–7.0 x
<quantity id="236C9B67ED65FFDA6E87FC40FC2FFC20" box="[915,1008,924,950]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="34" pageNumber="35" unit="mm" value="8.5">8.5 mm</quantity>
, 1 ov. F 6.5 x 8.0 mm, 1 F 4.5 x
<quantity id="236C9B67ED65FFDA6C5EFC18FE7CFC48" box="[330,419,964,990]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="34" pageNumber="35" unit="mm" value="5.5">5.5 mm</quantity>
(
<collectionCode id="8285AE47ED65FFDA6CA6FC18FDD2FC48" box="[434,525,964,990]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="35">MNHN</collectionCode>
B29858)
</materialsCitation>
.
</paragraph>
<paragraph id="E42B3682ED65FFDA6C1CFBDDFEA0FB8C" blockId="34.[264,1323,1025,1891]" box="[264,383,1025,1050]" pageId="34" pageNumber="35">
<emphasis id="D6E0EA90ED65FFDA6C1CFBDDFEA0FB8C" box="[264,383,1025,1050]" italics="true" pageId="34" pageNumber="35">Diagnosis</emphasis>
</paragraph>
<paragraph id="E42B3682ED65FFDA6C2CFBF5FE17FACD" blockId="34.[264,1323,1025,1891]" pageId="34" pageNumber="35">
Smallsize species. Carapace with concave lateral margins, markedly convergent posteriorly, without tooth behind exorbital angle (
<figureCitation id="7CAF2A07ED65FFDA6E2BFB8DFC77FBFD" box="[831,936,1105,1131]" captionStart="FIGURE 10" captionStartId="35.[264,368,1718,1742]" captionTargetBox="[278,1311,690,1697]" captionTargetId="figure-144@35.[264,1323,673,1714]" captionTargetPageId="35" captionText="FIGURE 10. —Pachygrapsus planifrons de Man, 1888. Male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490): a) dorsal view; b) front; c) left chela, outer face; d) right P2; e) right P5; f) abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049252" httpUri="https://zenodo.org/record/5049252/files/figure.png" pageId="34" pageNumber="35">Fig. 10a</figureCitation>
). Dorsal surface flattened, only weakly convex; protogastric, hepatic regions with thin transverse striae, branchial region with oblique striae; all striae glabrous; mesogastric, cardiac, intestinal regions almost totally smooth. Front 0.6 times width at exorbital angles, anterior margin almost straight (
<figureCitation id="7CAF2A07ED65FFDA6C05FB2DFEA7FA9D" box="[273,376,1265,1291]" captionStart="FIGURE 10" captionStartId="35.[264,368,1718,1742]" captionTargetBox="[278,1311,690,1697]" captionTargetId="figure-144@35.[264,1323,673,1714]" captionTargetPageId="35" captionText="FIGURE 10. —Pachygrapsus planifrons de Man, 1888. Male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490): a) dorsal view; b) front; c) left chela, outer face; d) right P2; e) right P5; f) abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049252" httpUri="https://zenodo.org/record/5049252/files/figure.png" pageId="34" pageNumber="35">Fig. 10b</figureCitation>
). Infraorbital margin nearly entire, composed of low granules, outer indentation very low; infraorbital margin forming very obtuse angle with inner margin of exorbital tooth (
<figureCitation id="7CAF2A07ED65FFDA6C40FA9DFE64FACD" box="[340,443,1345,1371]" captionStart="FIGURE 14" captionStartId="47.[264,368,1403,1427]" captionTargetBox="[274,1309,325,1344]" captionTargetId="figure-211@47.[265,1322,294,1379]" captionTargetPageId="47" captionText="FIGURE 14. —Infra–orbital margin: a) Pachygrapsus fakaravensis Rathbun (1907), ov. female 15.0 x 16.5 mm, Taiaro, French Polynesia (MNHN B25782); b) Pachygrapsus laevimanus Stimpson, 1858, neotype male 18.0 x 21.7 mm, Sydney, Australia (AM P7478); c) Pachygrapsus minutus A. MilneEdwards, 1873, male 5.2 x 7.0 mm, Moruroa, French Polynesia (MNHN B29273); d) Pachygrapsus planifrons de Man, 1888, male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), male 12.1 x 15.3 mm, Taiaro, French Polynesia (MNHN B29279); f) Pachygrapsus corrugatus (von Martens, 1872) ov. female 13.6 x 15.2 mm, Virgin Is. (USNM 72340); g) Pachygrapsus crassipes Randall, 1840, male 31.9 x 36.6 mm, Sagami Bay, Japan (NSMT Cr7516); h) Pachygrapsus gracilis (de Saussure, 1858), ov. female 8.5 x 11 mm, Angola (MNHN B12903); i) Pachygrapsus loveridgei Chace, 1966, male 8.1 x 10 mm, St. Helena I. (MNHN B17836); j) Pachygrapsus marmoratus (Fabricius, 1787), female 12.7 x 14.3 mm, Préfailles, France (MNHN B22654); k) Pachygrapsus maurus (Lucas, 1846), female 12.7 x 14.3 mm, Banyuls, France (MNHN B12933); l) Pachygrapsus transversus (Gibbes, 1850), male 17.0 x 22.5 mm, French Guiana (MNHN B16013). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049262" httpUri="https://zenodo.org/record/5049262/files/figure.png" pageId="34" pageNumber="35">Fig. 14d</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED65FFDA6C2CFAB5FE00F9DD" blockId="34.[264,1323,1025,1891]" pageId="34" pageNumber="35">
Cheliped merus with 4–5 acute spines at inner distoventral angle. Carpus with sharp inner dorsodistal spine. Outer face of chelae regularly convex; fixed finger with a longitudinal stria extending slightly on outer face of chela; cutting edge with 2–3 large teeth medially and few smaller teeth proximally. Dactyl with 6–8 small teeth over whole length of cutting edge, sometimes indistinct. Tip of fingers spoonlike, with distinctive rows of bristles (
<figureCitation id="7CAF2A07ED65FFDA6C79F9EDFE0EF9DD" box="[365,465,1585,1611]" captionStart="FIGURE 10" captionStartId="35.[264,368,1718,1742]" captionTargetBox="[278,1311,690,1697]" captionTargetId="figure-144@35.[264,1323,673,1714]" captionTargetPageId="35" captionText="FIGURE 10. —Pachygrapsus planifrons de Man, 1888. Male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490): a) dorsal view; b) front; c) left chela, outer face; d) right P2; e) right P5; f) abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049252" httpUri="https://zenodo.org/record/5049252/files/figure.png" pageId="34" pageNumber="35">Fig. 10c</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED65FFDA6C2CF985FD31F97D" blockId="34.[264,1323,1025,1891]" pageId="34" pageNumber="35">
Ambulatory legs with scattered long bristles on carpi, propodi, dactyli, but without brush of setae. Upper margins of meri armed with several long mobile spines, lower margins with 1–2 spines (usually 2) at distoventral angles; lower margin of P5 merus smooth, without submedian tubercle (
<figureCitation id="7CAF2A07ED65FFDA6F4AF90DFD3DF97D" box="[606,738,1745,1771]" captionStart="FIGURE 10" captionStartId="35.[264,368,1718,1742]" captionTargetBox="[278,1311,690,1697]" captionTargetId="figure-144@35.[264,1323,673,1714]" captionTargetPageId="35" captionText="FIGURE 10. —Pachygrapsus planifrons de Man, 1888. Male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490): a) dorsal view; b) front; c) left chela, outer face; d) right P2; e) right P5; f) abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049252" httpUri="https://zenodo.org/record/5049252/files/figure.png" pageId="34" pageNumber="35">Fig. 10d–e</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED65FFDA6C2CF925FCB9F8F5" blockId="34.[264,1323,1025,1891]" pageId="34" pageNumber="35">
Abdominal tergites smooth. Telson of male rounded at tip, longer than sixth somite; telson and sixth somite forming triangle with lateral margins slightly concave (
<figureCitation id="7CAF2A07ED65FFDA69AEF8FDFAC1F8AD" box="[1210,1310,1825,1851]" captionStart="FIGURE 10" captionStartId="35.[264,368,1718,1742]" captionTargetBox="[278,1311,690,1697]" captionTargetId="figure-144@35.[264,1323,673,1714]" captionTargetPageId="35" captionText="FIGURE 10. —Pachygrapsus planifrons de Man, 1888. Male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490): a) dorsal view; b) front; c) left chela, outer face; d) right P2; e) right P5; f) abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049252" httpUri="https://zenodo.org/record/5049252/files/figure.png" pageId="34" pageNumber="35">Fig. 10f</figureCitation>
). G1 slender, petallike at tip (
<figureCitation id="7CAF2A07ED65FFDA6F43F895FD62F8F5" box="[599,701,1865,1891]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. MilneEdwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, g–l, from Chace, 1966, fig. 10–11; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="34" pageNumber="35">Fig. 15d</figureCitation>
; see remarks).
</paragraph>
<paragraph id="E42B3682ED64FFDB6C2CFEC0FDF2FE38" blockId="35.[264,1325,284,590]" pageId="35" pageNumber="36">
Colour (from
<collectingCountry id="9C837612ED64FFDB6CCAFEC0FD84FEA0" box="[478,603,284,310]" name="Clipperton Island" pageId="35" pageNumber="36">Clipperton</collectingCountry>
specimens): brown overall, with darker transverse bands on the ambulatory legs: proximal, median and distal bands on meri (sometimes unclear); proximal and distal bands on carpi and propodi; and median band of variable width on dactyli. Eggs orange red.
</paragraph>
<paragraph id="E42B3682ED64FFDB6C2CFE60FE74FDD8" blockId="35.[264,1325,284,590]" pageId="35" pageNumber="36">
Measurements: small size; usual carapace size 5 x
<quantity id="236C9B67ED64FFDB6E9FFE60FC0DFE40" box="[907,978,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="35" pageNumber="36" unit="mm" value="7.0">7 mm</quantity>
; greatest size indicated in the literature 9 x
<quantity id="236C9B67ED64FFDB6CB2FE38FE26FE68" box="[422,505,484,510]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="35" pageNumber="36" unit="mm" value="11.0">11 mm</quantity>
(
<bibRefCitation id="80054B73ED64FFDB6F1CFE38FD3CFE68" author="Edmondson, C. H." box="[520,739,484,510]" pageId="35" pageNumber="36" pagination="153 - 202" refId="ref30023" refString="Edmondson, C. H. (1959) Hawaiian Grapsidae. Occasional Papers of Bernice Pauahi Bishop Museum, 22 (10), 153 - 202." type="journal article" year="1959">Edmondson, 1959</bibRefCitation>
;
<bibRefCitation id="80054B73ED64FFDB6FE4FE38FC5EFE68" author="Garth, J. S." box="[752,897,484,510]" pageId="35" pageNumber="36" pagination="1 - 46" refId="ref30463" refString="Garth, J. S. (1965) The brachyuran Decapod Crustaceans of Clipperton Island. Proceedings of the California Academy of Sciences, fourth series, 33 (1), 1 - 46." type="journal article" year="1965">Garth, 1965</bibRefCitation>
); specimens examined ranged from: males 5.1 x 6.3–5.6 x
<quantity id="236C9B67ED64FFDB6F18FDD0FDB5FDB0" box="[524,618,524,550]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="35" pageNumber="36" unit="mm" value="6.8">6.8 mm</quantity>
, females 4.6 x 6.0–5.6 x
<quantity id="236C9B67ED64FFDB6E85FDD0FC31FDB0" box="[913,1006,524,550]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="35" pageNumber="36" unit="mm" value="7.3">7.3 mm</quantity>
; smallest ovigerous female 4.6 x 6.0 mm.
</paragraph>
<caption id="B0EB660AED64FFDB6C1CF96AFEAEF885" ID-DOI="http://doi.org/10.5281/zenodo.5049252" ID-Zenodo-Dep="5049252" httpUri="https://zenodo.org/record/5049252/files/figure.png" pageId="35" pageNumber="36" startId="35.[264,368,1718,1742]" targetBox="[278,1311,690,1697]" targetPageId="35">
<paragraph id="E42B3682ED64FFDB6C1CF96AFEAEF885" blockId="35.[264,1323,1718,1811]" pageId="35" pageNumber="36">
<emphasis id="D6E0EA90ED64FFDB6C1CF96AFE44F958" bold="true" box="[264,411,1718,1742]" pageId="35" pageNumber="36">FIGURE 10.</emphasis>
—
<taxonomicName id="23944D01ED64FFDB6CD4F96BFCA8F95B" authority="de Man, 1888" authorityName="De Man" authorityYear="1888" box="[448,887,1718,1742]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED64FFDB6CD4F96BFD11F958" box="[448,718,1719,1742]" italics="true" pageId="35" pageNumber="36">Pachygrapsus planifrons</emphasis>
<bibRefCitation id="80054B73ED64FFDB6FCCF96AFCA8F95B" author="De Man, J. G." box="[728,887,1718,1741]" pageId="35" pageNumber="36" pagination="215 - 600" refId="ref31574" refString="De Man, J. G. (1888) Bericht uber die im indischen Archipel von Dr. J. Brock gesammelten, Decapoden und Stomatopoden. Archiv fur Naturgeschichte, 53, 215 - 600, pl. 7 - 22." type="journal article" year="1888">de Man, 1888</bibRefCitation>
</taxonomicName>
. Male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490): a) dorsal view; b) front; c) left chela, outer face; d) right P2; e) right P5; f) abdomen.
</paragraph>
</caption>
<paragraph id="E42B3682ED63FFDC6C1CFEC0FE48FEA3" blockId="36.[264,1323,284,470]" box="[264,407,284,309]" pageId="36" pageNumber="37">
<emphasis id="D6E0EA90ED63FFDC6C1CFEC0FE48FEA3" box="[264,407,284,309]" italics="true" pageId="36" pageNumber="37">Distribution</emphasis>
</paragraph>
<paragraph id="E42B3682ED63FFDC6C2CFE9FFE85FE40" blockId="36.[264,1323,284,470]" pageId="36" pageNumber="37">
<emphasis id="D6E0EA90ED63FFDC6C2CFE9FFCE2FECB" bold="true" box="[312,829,323,349]" pageId="36" pageNumber="37">West Indian Ocean to east Pacific islands</emphasis>
:
<collectingCountry id="9C837612ED63FFDC6E59FE98FC11FEC8" box="[845,974,324,350]" name="Seychelles" pageId="36" pageNumber="37">Seychelles</collectingCountry>
; Christmas I,, CocosKeeling Is.;
<collectingCountry id="9C837612ED63FFDC6C22FEB0FE7EFE10" box="[310,417,364,390]" name="Thailand" pageId="36" pageNumber="37">Thailand</collectingCountry>
,
<collectingCountry id="9C837612ED63FFDC6CBBFEB0FDFAFE10" box="[431,549,364,390]" name="Indonesia" pageId="36" pageNumber="37">Indonesia</collectingCountry>
; South
<collectingCountry id="9C837612ED63FFDC6F95FEB0FD18FE10" box="[641,711,364,390]" name="China" pageId="36" pageNumber="37">China</collectingCountry>
Sea (Anambas Is.,
<collectingRegion id="2650F860ED63FFDC6EA7FEB0FBD7FE10" box="[947,1032,364,390]" country="China" name="Hainan" pageId="36" pageNumber="37">Hainan</collectingRegion>
I., Xisha or Paracel Is);
<collectingCountry id="9C837612ED63FFDC6C1CFE48FE91FE38" box="[264,334,404,430]" name="Japan" pageId="36" pageNumber="37">Japan</collectingCountry>
; Caroline Is.; Mariana Is.;
<collectingCountry id="9C837612ED63FFDC6F99FE48FD33FE38" box="[653,748,404,430]" name="Kiribati" pageId="36" pageNumber="37">Kiribati</collectingCountry>
; Funafuti; Hawaiian Is.; Tuamotu Archip.;
<collectingCountry id="9C837612ED63FFDC69F8FE48FE89FE40" name="Clipperton Island" pageId="36" pageNumber="37">Clipperton</collectingCountry>
.
</paragraph>
<paragraph id="E42B3682ED63FFDC6C1CFDD0FC27FDD8" blockId="36.[264,1016,524,590]" box="[264,1016,524,590]" pageId="36" pageNumber="37">
<emphasis id="D6E0EA90ED63FFDC6C1CFDD0FEBEFDB3" box="[264,353,524,549]" italics="true" pageId="36" pageNumber="37">Habitat</emphasis>
Common in the high intertidal zone, under rocks and debris.
</paragraph>
<paragraph id="E42B3682ED63FFDC6C1CFD58FEB1FD0B" blockId="36.[264,1323,644,1630]" box="[264,366,644,669]" pageId="36" pageNumber="37">
<emphasis id="D6E0EA90ED63FFDC6C1CFD58FEB1FD0B" box="[264,366,644,669]" italics="true" pageId="36" pageNumber="37">Remarks</emphasis>
</paragraph>
<paragraph id="E42B3682ED63FFDC6C2CFD70FB4AFB90" blockId="36.[264,1323,644,1630]" pageId="36" pageNumber="37">
<taxonomicName id="23944D01ED63FFDC6C2CFD70FD81FD53" authorityName="De Man" authorityYear="1888" box="[312,606,684,709]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6C2CFD70FD81FD53" box="[312,606,684,709]" italics="true" pageId="36" pageNumber="37">Pachygrapsus planifrons</emphasis>
</taxonomicName>
is unique in possessing rows of bristles at the tips of the fingers of the chelae. This character alone is sufficient to distinguish it from all other species of
<taxonomicName id="23944D01ED63FFDC6C3CFD20FE11FC83" authorityName="Randall" authorityYear="1840" box="[296,462,764,789]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0EA90ED63FFDC6C3CFD20FE11FC83" box="[296,462,764,789]" italics="true" pageId="36" pageNumber="37">Pachygrapsus</emphasis>
</taxonomicName>
. It is similar to
<taxonomicName id="23944D01ED63FFDC6F9DFD20FCDDFC83" authorityName="A. Milne-Edwards" authorityYear="1873" box="[649,770,764,789]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="D6E0EA90ED63FFDC6F9DFD20FCDDFC83" box="[649,770,764,789]" italics="true" pageId="36" pageNumber="37">P. minutus</emphasis>
</taxonomicName>
in its small size, but the two can be easily separated. The exorbital angle is produced and more spinous in
<taxonomicName id="23944D01ED63FFDC6EC7FCF8FBB8FCAB" authorityName="De Man" authorityYear="1888" box="[979,1127,804,829]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6EC7FCF8FBB8FCAB" box="[979,1127,804,829]" italics="true" pageId="36" pageNumber="37">P. planifrons</emphasis>
</taxonomicName>
than in
<taxonomicName id="23944D01ED63FFDC69DDFCF8FEF5FCF3" authorityName="A. Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="D6E0EA90ED63FFDC69DDFCF8FEF5FCF3" italics="true" pageId="36" pageNumber="37">P. minutus</emphasis>
</taxonomicName>
, the lateral margins of the carapace are concave in
<taxonomicName id="23944D01ED63FFDC6EA2FC90FB90FCF3" authorityName="De Man" authorityYear="1888" box="[950,1103,844,869]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6EA2FC90FB90FCF3" box="[950,1103,844,869]" italics="true" pageId="36" pageNumber="37">P. planifrons</emphasis>
</taxonomicName>
but straight in
<taxonomicName id="23944D01ED63FFDC6801FC90FEBAFC1B" authorityName="A. Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="D6E0EA90ED63FFDC6801FC90FEBAFC1B" italics="true" pageId="36" pageNumber="37">P. minutus</emphasis>
</taxonomicName>
, the anterior margin of the front is almost straight in
<taxonomicName id="23944D01ED63FFDC6EC4FCA8FBBCFC1B" authorityName="De Man" authorityYear="1888" box="[976,1123,884,909]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6EC4FCA8FBBCFC1B" box="[976,1123,884,909]" italics="true" pageId="36" pageNumber="37">P. planifrons</emphasis>
</taxonomicName>
but sinuous in
<taxonomicName id="23944D01ED63FFDC6801FCA8FEBAFC23" authorityName="A. Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="D6E0EA90ED63FFDC6801FCA8FEBAFC23" italics="true" pageId="36" pageNumber="37">P. minutus</emphasis>
</taxonomicName>
, the lower margin of the P5 merus is smooth in
<taxonomicName id="23944D01ED63FFDC6EB0FC40FBE5FC23" authorityName="De Man" authorityYear="1888" box="[932,1082,924,949]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6EB0FC40FBE5FC23" box="[932,1082,924,949]" italics="true" pageId="36" pageNumber="37">P. planifrons</emphasis>
</taxonomicName>
while having a submedian tubercle in
<taxonomicName id="23944D01ED63FFDC6CFAFC18FDB6FC4B" authorityName="A. Milne-Edwards" authorityYear="1873" box="[494,617,964,989]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="D6E0EA90ED63FFDC6CFAFC18FDB6FC4B" box="[494,617,964,989]" italics="true" pageId="36" pageNumber="37">P. minutus</emphasis>
</taxonomicName>
and the distal end of the G1 has a short vertical gutterlike process in
<taxonomicName id="23944D01ED63FFDC6C91FC30FE20FB93" authorityName="A. Milne-Edwards" authorityYear="1873" box="[389,511,1004,1029]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="D6E0EA90ED63FFDC6C91FC30FE20FB93" box="[389,511,1004,1029]" italics="true" pageId="36" pageNumber="37">P. minutus</emphasis>
</taxonomicName>
(
<figureCitation id="7CAF2A07ED63FFDC6F1BFC30FDAEFB90" box="[527,625,1004,1030]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. MilneEdwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, g–l, from Chace, 1966, fig. 10–11; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="36" pageNumber="37">Fig. 15c</figureCitation>
) but a petallike tip in
<taxonomicName id="23944D01ED63FFDC6E69FC30FBCEFB93" authorityName="De Man" authorityYear="1888" box="[893,1041,1004,1029]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6E69FC30FBCEFB93" box="[893,1041,1004,1029]" italics="true" pageId="36" pageNumber="37">P. planifrons</emphasis>
</taxonomicName>
(
<figureCitation id="7CAF2A07ED63FFDC6935FC30FB58FB90" box="[1057,1159,1004,1030]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. MilneEdwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, g–l, from Chace, 1966, fig. 10–11; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="36" pageNumber="37">Fig. 15d</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED63FFDC6C2CFBC8FB24FA00" blockId="36.[264,1323,644,1630]" pageId="36" pageNumber="37">
<bibRefCitation id="80054B73ED63FFDC6C2CFBC8FDC8FBB8" author="Chace, F. A." box="[312,535,1044,1070]" pageId="36" pageNumber="37" pagination="623 - 660" refId="ref29268" refString="Chace, F. A. (1966) Decapod crustaceans from St. Helena Island, South Atlantic. Proceedings of the United States National Museum, 118, 623 - 660." type="journal article" year="1966">Chace (1966: 645)</bibRefCitation>
indicated that
<taxonomicName id="23944D01ED63FFDC6FD8FBC8FCADFBBB" authorityName="Randall" authorityYear="1840" box="[716,882,1044,1069]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="D6E0EA90ED63FFDC6FD8FBC8FCADFBBB" box="[716,882,1044,1069]" italics="true" pageId="36" pageNumber="37">Pachygrapsus</emphasis>
</taxonomicName>
was in need of revision and figured the gonopods of 8 species in the USNM collection. These figures have been very useful for the present study and some of them are reproduced in
<figureCitation id="7CAF2A07ED63FFDC6EB1FBB8FBC3FBE8" box="[933,1052,1124,1150]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. MilneEdwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, g–l, from Chace, 1966, fig. 10–11; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="36" pageNumber="37">Figure 15</figureCitation>
. Some inconsistencies were found with the figure of the G1 of
<taxonomicName id="23944D01ED63FFDC6FC8FB50FCAEFB33" authorityName="De Man" authorityYear="1888" box="[732,881,1164,1189]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6FC8FB50FCAEFB33" box="[732,881,1164,1189]" italics="true" pageId="36" pageNumber="37">P. planifrons</emphasis>
</taxonomicName>
(
<bibRefCitation id="80054B73ED63FFDC6E94FB50FB8DFB30" author="Chace, F. A." box="[896,1106,1164,1190]" pageId="36" pageNumber="37" pagination="623 - 660" refId="ref29268" refString="Chace, F. A. (1966) Decapod crustaceans from St. Helena Island, South Atlantic. Proceedings of the United States National Museum, 118, 623 - 660." type="journal article" year="1966">Chace, 1966: 645</bibRefCitation>
, fig. 11f, a male cl
<quantity id="236C9B67ED63FFDC6C1CFB68FEBDFB58" box="[264,354,1204,1230]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="36" pageNumber="37" unit="mm" value="9.5">9.5 mm</quantity>
from Honolulu), however. It has a gutterlike tip instead of tuliplike as shown in
<figureCitation id="7CAF2A07ED63FFDC6C1CFB00FE54FB60" box="[264,395,1244,1270]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. MilneEdwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, g–l, from Chace, 1966, fig. 10–11; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="36" pageNumber="37">Figure 15d</figureCitation>
, even if the G1 illustrated here is from the same specimen as that of Chace, as its size and collection details were the same, and the G1 was already dissected in the jar. We therefore assume that there was a mistake made when Chace’s plate was being prepared. The tuliplike shape of the
<taxonomicName id="23944D01ED63FFDC6F86FA88FCFAFAFB" authorityName="De Man" authorityYear="1888" box="[658,805,1364,1389]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6F86FA88FCFAFAFB" box="[658,805,1364,1389]" italics="true" pageId="36" pageNumber="37">P. planifrons</emphasis>
</taxonomicName>
G1 was also found in other males examined here, as well as in one illustrated for a Chinese specimen (
<bibRefCitation id="80054B73ED63FFDC6EA6FAA0FBA7FA00" author="Chen, H." box="[946,1144,1404,1430]" pageId="36" pageNumber="37" pagination="117 - 147" refId="ref29360" refString="Chen, H. (1980) Studies on the crabs of the Xisha Islands, Guangdong Province, China. II. Studia Marina Sinica, 17, 117 - 147, pl. 1 - 4 (in Chinese)." type="journal article" year="1980">Chen, 1980: 140</bibRefCitation>
, fig. 225).
</paragraph>
<paragraph id="E42B3682ED63FFDC6C2CFA78FE7CF9C8" blockId="36.[264,1323,644,1630]" pageId="36" pageNumber="37">
<bibRefCitation id="80054B73ED63FFDC6C2CFA78FD9BFA28" author="Borradaile, L. A." box="[312,580,1444,1470]" pageId="36" pageNumber="37" pagination="568 - 596" refId="ref28916" refString="Borradaile, L. A. (1900) On some Crustacea from the South Pacific. Part IV. The Crabs. Proceedings of the Zoological Society, London, 568 - 596." type="book chapter" year="1900">Borradaile (1900: 592</bibRefCitation>
, pl. 42, fig. 7) described
<taxonomicName id="23944D01ED63FFDC6E68FA78FC3FFA2B" authorityName="Borradaile" authorityYear="1900" box="[892,992,1444,1469]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="D6E0EA90ED63FFDC6E68FA78FC3FFA2B" box="[892,992,1444,1469]" italics="true" pageId="36" pageNumber="37">P. laevis</emphasis>
</taxonomicName>
from a female collected at
<collectingRegion id="2650F860ED63FFDC6C1CFA10FEB0FA70" box="[264,367,1484,1510]" country="Tuvalu" name="Funafuti" pageId="36" pageNumber="37">Funafuti</collectingRegion>
. The characters described by Borradaile are those of
<taxonomicName id="23944D01ED63FFDC6EF0FA10FBA6FA73" authorityName="De Man" authorityYear="1888" box="[996,1145,1484,1509]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6EF0FA10FBA6FA73" box="[996,1145,1484,1509]" italics="true" pageId="36" pageNumber="37">P. planifrons</emphasis>
</taxonomicName>
, especially the characteristic rows of bristles at the tips of the fingers of the chelae. The
<typeStatus id="3B2F8820ED63FFDC6979FA28FB40F998" box="[1133,1183,1524,1550]" pageId="36" pageNumber="37">type</typeStatus>
of
<taxonomicName id="23944D01ED63FFDC69DCFA28FAF5F99B" authorityName="Borradaile" authorityYear="1900" box="[1224,1322,1524,1549]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="D6E0EA90ED63FFDC69DCFA28FAF5F99B" box="[1224,1322,1524,1549]" italics="true" pageId="36" pageNumber="37">P. laevis</emphasis>
</taxonomicName>
(
<collectionCode id="8285AE47ED63FFDC6C04F9C0FEB9F9A0" box="[272,358,1564,1590]" pageId="36" pageNumber="37">UMZC</collectionCode>
<date id="902A1042ED63FFDC6C7AF9C0FE2CF9A0" box="[366,499,1564,1590]" pageId="36" pageNumber="37" value="1897-11-01">11.01.1897</date>
) was examined and it can be confirmed that it is a junior synonym of
<taxonomicName id="23944D01ED63FFDC6C1CF998FE43F9CB" authorityName="De Man" authorityYear="1888" box="[264,412,1604,1629]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="planifrons">
<emphasis id="D6E0EA90ED63FFDC6C1CF998FE43F9CB" box="[264,412,1604,1629]" italics="true" pageId="36" pageNumber="37">P. planifrons</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>