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<mods:title id="4A833249F1297750A6F330079969D827">A revision of the genus Pachygrapsus Randall, 1840 (Crustacea: Decapoda: Brachyura, Grapsidae), with special reference to the Southwest Pacific species</mods:title>
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<taxonomicName id="23944D01ED4FFFF06C1CF9B1FD06F911" ID-CoL="4BSBZ" authority="Randall, 1840" authorityName="Randall" authorityYear="1840" box="[264,729,1645,1671]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4FFFF06C1CF9B1FDFAF911" bold="true" box="[264,549,1645,1671]" italics="true" pageId="8" pageNumber="9">Pachygrapsus crassipes</emphasis>
<bibRefCitation id="80054B73ED4FFFF06F39F9B1FD06F911" author="Randall, J. W." box="[557,729,1645,1671]" pageId="8" pageNumber="9" pagination="106 - 148" refId="ref33057" refString="Randall, J. W. (1840) Catalogue of the Crustacea brought by Thomas Nuttall and J. K. Townsend, from the west coast of North America and the Sandwich Islands, with description of such species as are apparently new, among which are included several species of different localities, previously existing in the collection of the Academy. Journal of the Academy of Natural Science of Philadelphia, 1839 (1840), 8 (1), 106 - 148, pl. 3 - 7." type="journal article" year="1840">Randall, 1840</bibRefCitation>
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(
<figureCitation id="7CAF2A07ED4FFFF06FFDF9B1FC55F911" box="[745,906,1645,1671]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="8" pageNumber="9">Figures 2ae</figureCitation>
,
<figureCitation id="7CAF2A07ED4FFFF06E83F9B1FC1BF911" box="[919,964,1645,1671]" captionStart="FIGURE 14" captionStartId="47.[264,368,1403,1427]" captionTargetBox="[274,1309,325,1344]" captionTargetId="figure-211@47.[265,1322,294,1379]" captionTargetPageId="47" captionText="FIGURE 14. —Infraorbital margin: a) Pachygrapsus fakaravensis Rathbun (1907), ov. female 15.0 x 16.5 mm, Taiaro, French Polynesia (MNHN B25782); b) Pachygrapsus laevimanus Stimpson, 1858, neotype male 18.0 x 21.7 mm, Sydney, Australia (AM P7478); c) Pachygrapsus minutus A. Milne­Edwards, 1873, male 5.2 x 7.0 mm, Moruroa, French Polynesia (MNHN B29273); d) Pachygrapsus planifrons de Man, 1888, male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), male 12.1 x 15.3 mm, Taiaro, French Polynesia (MNHN B29279); f) Pachygrapsus corrugatus (von Martens, 1872) ov. female 13.6 x 15.2 mm, Virgin Is. (USNM 72340); g) Pachygrapsus crassipes Randall, 1840, male 31.9 x 36.6 mm, Sagami Bay, Japan (NSMT Cr7516); h) Pachygrapsus gracilis (de Saussure, 1858), ov. female 8.5 x 11 mm, Angola (MNHN B12903); i) Pachygrapsus loveridgei Chace, 1966, male 8.1 x 10 mm, St. Helena I. (MNHN B17836); j) Pachygrapsus marmoratus (Fabricius, 1787), female 12.7 x 14.3 mm, Préfailles, France (MNHN B22654); k) Pachygrapsus maurus (Lucas, 1846), female 12.7 x 14.3 mm, Banyuls, France (MNHN B12933); l) Pachygrapsus transversus (Gibbes, 1850), male 17.0 x 22.5 mm, French Guiana (MNHN B16013). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049262" httpUri="https://zenodo.org/record/5049262/files/figure.png" pageId="8" pageNumber="9">
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,
<figureCitation id="7CAF2A07ED4FFFF06EC6F9B1FC20F911" box="[978,1023,1645,1671]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. Milne­Edwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, gl, from Chace, 1966, fig. 1011; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="8" pageNumber="9">
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,
<figureCitation id="7CAF2A07ED4FFFF06918F9B1FBF5F911" box="[1036,1066,1645,1671]" captionStart="FIGURE 16" captionStartId="49.[264,368,1320,1344]" captionTargetBox="[267,1320,730,1296]" captionTargetId="figure-253@49.[267,1320,726,1296]" captionTargetPageId="49" captionText="FIGURE 16. —Geographic distribution of: a) Pachygrapsus crassipes Randall, 1840 (Stars); b) Pachygrapsus transversus (Gibbes, 1850) (full dots); and c) P. laevimanus Stimpson, 1858 (empty dots). ab, mostly from literature; c) from collections examined during this study. Occurrence of P. laevimanus at Easter I. uncertain (?) and must be confirmed by examination of G1 (see text)." figureDoi="http://doi.org/10.5281/zenodo.5049268" httpUri="https://zenodo.org/record/5049268/files/figure.png" pageId="8" pageNumber="9">16</figureCitation>
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<emphasis id="D6E0EA90ED4FFFF06C1CF965FDD5F946" box="[264,522,1721,1744]" italics="true" pageId="8" pageNumber="9">Pachygrapsus crassipes</emphasis>
<bibRefCitation id="80054B73ED4FFFF06F07F964FD3EF959" author="Randall, J. W." box="[531,737,1720,1743]" pageId="8" pageNumber="9" pagination="106 - 148" refId="ref33057" refString="Randall, J. W. (1840) Catalogue of the Crustacea brought by Thomas Nuttall and J. K. Townsend, from the west coast of North America and the Sandwich Islands, with description of such species as are apparently new, among which are included several species of different localities, previously existing in the collection of the Academy. Journal of the Academy of Natural Science of Philadelphia, 1839 (1840), 8 (1), 106 - 148, pl. 3 - 7." type="journal article" year="1840">Randall, 1840: 127</bibRefCitation>
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[
<typeStatus id="3B2F8820ED4FFFF06FFBF964FCFFF959" box="[751,800,1720,1743]" pageId="8" pageNumber="9">type</typeStatus>
locality: Hawaiian Is. but according to
<bibRefCitation id="80054B73ED4FFFF069C4F964FEA0F979" author="Rathbun, M. J." pageId="8" pageNumber="9" pagination="1 - 461" refId="ref33490" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, i - xxii, 1 - 461, pl. 1 - 161." type="journal article" year="1918">Rathbun (1918)</bibRefCitation>
and Edmonson (1959) this locality must be erroneous]. —
<treatmentCitation id="65351093ED4FFFF06904F904FB39F979" author="Stimpson, W." box="[1040,1254,1752,1775]" page="48" pageId="8" pageNumber="9" year="1858">
<bibRefCitation id="80054B73ED4FFFF06904F904FB39F979" author="Stimpson, W." box="[1040,1254,1752,1775]" pageId="8" pageNumber="9" pagination="93 - 110" refId="ref34073" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit W. Stimpson. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences, Philadelphia, 10, 93 - 110 [39 - 56]." type="journal article" year="1858">Stimpson, 1858: 48</bibRefCitation>
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[102] (California,
<collectingCountry id="9C837612ED4FFFF06CAFF924FE23F899" box="[443,508,1784,1807]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
); 1907: 116 (
<collectingCountry id="9C837612ED4FFFF06F9DF924FD13F899" box="[649,716,1784,1807]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
). —
<treatmentCitation id="65351093ED4FFFF06FEDF924FC0EF899" author="Ortmann, A." box="[761,977,1784,1807]" page="708" pageId="8" pageNumber="9" year="1894">
<bibRefCitation id="80054B73ED4FFFF06FEDF924FC0EF899" author="Ortmann, A." box="[761,977,1784,1807]" pageId="8" pageNumber="9" pagination="683 - 772" refId="ref32512" refString="Ortmann, A. (1894) Die Decapoden - Krebse des Strassburger Museums. VIII Theil. Abtheilung: Brachyura (Brachyura genuina Boas), III. Unterabtheilung: Cancroidea, 2. Section: Cancrinea, 2. Gruppe: Catametopa. Zoologischen Jahrbuchern, Iena, Abtheilung fur Systematik, 8, 683 - 772, pl. 23." type="journal article" year="1894">Ortmann, 1894: 708</bibRefCitation>
</treatmentCitation>
(California;
<collectingCountry id="9C837612ED4FFFF0694FF924FB43F899" box="[1115,1180,1784,1807]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
). —
<treatmentCitation id="65351093ED4FFFF069DEF924FE7AF8B9" author="Rathbun, M. J." page="241" pageId="8" pageNumber="9" year="1918">
<bibRefCitation id="80054B73ED4FFFF069DEF924FE7AF8B9" author="Rathbun, M. J." pageId="8" pageNumber="9" pagination="1 - 461" refId="ref33490" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, i - xxii, 1 - 461, pl. 1 - 161." type="journal article" year="1918">Rathbun, 1918: 241</bibRefCitation>
</treatmentCitation>
(California; Galápagos Is.,
<collectingCountry id="9C837612ED4FFFF06FC0F8C4FCCCF8B9" box="[724,787,1816,1839]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
,
<collectingCountry id="9C837612ED4FFFF06E34F8C4FCBAF8B9" box="[800,869,1816,1839]" name="South Korea" pageId="8" pageNumber="9">Korea</collectingCountry>
; but not
<collectingCountry id="9C837612ED4FFFF06ED2F8C4FBDDF8B9" box="[966,1026,1816,1839]" name="Chile" pageId="8" pageNumber="9">Chile</collectingCountry>
, see distribution). —
<treatmentCitation id="65351093ED4FFFF069F3F8C4FE48F8D9" author="Tesch, J. J." page="76" pageId="8" pageNumber="9" year="1918">
<bibRefCitation id="80054B73ED4FFFF069F3F8C4FE48F8D9" author="Tesch, J. J." pageId="8" pageNumber="9" pagination="1 - 148" refId="ref34227" refString="Tesch, J. J. (1918) The Decapoda Brachyura of the Siboga expedition. I. Hymenosomidae, Retroplumidae, Ocypodidae, Grapsidae, and Gecarcinidae. Siboga - Expeditie, 39 c, 1 - 148, pl. 1 - 6." type="journal article" year="1918">Tesch, 1918: 76</bibRefCitation>
</treatmentCitation>
(Key). —
<treatmentCitation id="65351093ED4FFFF06F1EF8E4FD1DF8D9" author="Garth, J. S." box="[522,706,1848,1871]" page="103" pageId="8" pageNumber="9" year="1957">
<bibRefCitation id="80054B73ED4FFFF06F1EF8E4FD1DF8D9" author="Garth, J. S." box="[522,706,1848,1871]" pageId="8" pageNumber="9" pagination="1 - 128" refId="ref30372" refString="Garth, J. S. (1957) Reports of the Lund University Chile Expedition 1948 - 49, n ° 29. The crustacea Decapoda Brachyura of Chile. Lunds Universitets A rsskrift, n. f., Avd. 2, 53 (7), 1 - 128." type="journal article" year="1957">Garth, 1957: 103</bibRefCitation>
</treatmentCitation>
(invalid record in
<collectingCountry id="9C837612ED4FFFF06E84F8E4FC12F8D9" box="[912,973,1848,1871]" name="Chile" pageId="8" pageNumber="9">Chile</collectingCountry>
; see distribution). —Edmonson, 1959: 168 (California). —
<treatmentCitation id="65351093ED4FFFF06F45F884FCDAF8F9" author="Sakai, T." box="[593,773,1880,1903]" page="634" pageId="8" pageNumber="9" year="1976">
<bibRefCitation id="80054B73ED4FFFF06F45F884FCDAF8F9" author="Sakai, T." box="[593,773,1880,1903]" pageId="8" pageNumber="9" pagination="1 - 251" refId="ref33764" refString="Sakai, T. (1976) Crabs of Japan and the adjacent seas. Tokyo, Kodansha, English volume, i - xxix, 773 pp.; Japanese volume, 461 pp.; plates volume, pl. 1 - 251." type="book chapter" year="1976">Sakai, 1976: 634</bibRefCitation>
</treatmentCitation>
, pl. 218, fig. 1 (
<collectingCountry id="9C837612ED4FFFF06EBAF884FC30F8F9" box="[942,1007,1880,1903]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
,
<collectingCountry id="9C837612ED4FFFF06EEEF884FBE0F8F9" box="[1018,1087,1880,1903]" name="South Korea" pageId="8" pageNumber="9">Korea</collectingCountry>
; references). —
<treatmentCitation id="65351093ED4FFFF169FCF884FE38FEA5" author="Dai, A. &amp; Yang, S." lastPageId="9" lastPageNumber="10" page="511" pageId="8" pageNumber="9" year="1991">
<bibRefCitation id="80054B73ED4FFFF169FCF884FE38FEA5" author="Dai, A. &amp; Yang, S." lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" refId="ref29508" refString="Dai, A. &amp; Yang, S. (1991) Crabs of the China seas. China Ocean Press Beijing; Springer-Verlag Berlin, Heidelberg, New York, Tokyo (English Edition), 682 pp." type="book" year="1991">Dai &amp; Yang, 1991: 511</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="9C837612ED4EFFF16CE1FEC0FDE9FEA5" box="[501,566,284,307]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
sea,
<collectingRegion id="2650F860ED4EFFF16F7AFEC0FD32FEA5" box="[622,749,284,307]" country="China" name="Guangdong" pageId="9" pageNumber="10">Guangdong</collectingRegion>
,
<collectingRegion id="2650F860ED4EFFF16FECFEC0FCE0FEA5" box="[760,831,284,307]" country="China" name="Fujian" pageId="9" pageNumber="10">Fujian</collectingRegion>
, and
<collectingRegion id="2650F860ED4EFFF16E63FEC0FC04FEA5" box="[887,987,284,307]" country="China" name="Zhejiang" pageId="9" pageNumber="10">Zhejiang</collectingRegion>
). —
<treatmentCitation id="65351093ED4EFFF1691CFEC0FB2EFEA5" author="Hendrickx, M. E." box="[1032,1265,284,307]" page="140" pageId="9" pageNumber="10" year="1995">
<bibRefCitation id="80054B73ED4EFFF1691CFEC0FB2EFEA5" author="Hendrickx, M. E." box="[1032,1265,284,307]" pageId="9" pageNumber="10" pagination="125 - 150" refId="ref30852" refString="Hendrickx, M. E. (1995) Checklist of brachyuran crabs (Crustacea: Decapoda) from the Eastern Tropical Pacific. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 65, 125 - 150." type="journal article" year="1995">Hendrickx, 1995: 140</bibRefCitation>
</treatmentCitation>
(east Pacific from Oregon to Gulf of California and Rocas Alijos; references). —
<treatmentCitation id="65351093ED4EFFF1694FFEE0FE6EFEE5" author="Cuesta, J. A. &amp; Schubart, C. D." page="1501" pageId="9" pageNumber="10" year="1998">
<bibRefCitation id="80054B73ED4EFFF1694FFEE0FE6EFEE5" author="Cuesta, J. A. &amp; Schubart, C. D." pageId="9" pageNumber="10" pagination="1499 - 1508" refId="ref29456" refString="Cuesta, J. A. &amp; Schubart, C. D. (1998) Morphological and molecular differentiation between three allopatric populations of the littoral crabs Pachygrapsus transversus (Gibbes, 1850) (Brachyura, Grapsidae). Journal of Natural History, 32, 1499 - 1508." type="journal article" year="1998">Cuesta &amp; Schubart, 1998: 1501</bibRefCitation>
</treatmentCitation>
(California). —
<treatmentCitation id="65351093ED4EFFF16F49FE80FCFAFEE5" author="Ng, P. K. L. &amp; Wang, C. H. &amp; Ho, P. H. &amp; Shih, H. T." box="[605,805,348,371]" page="40" pageId="9" pageNumber="10" year="2001">
<bibRefCitation id="80054B73ED4EFFF16F49FE80FCFAFEE5" author="Ng, P. K. L. &amp; Wang, C. H. &amp; Ho, P. H. &amp; Shih, H. T." box="[605,805,348,371]" pageId="9" pageNumber="10" pagination="1 - 86" refId="ref32394" refString="Ng, P. K. L., Wang, C. H., Ho, P. H. &amp; Shih, H. T. (2001) An annotated checklist of the brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum, Special Publication Series, 11, 1 - 86." type="journal article" year="2001">
Ng
<emphasis id="D6E0EA90ED4EFFF16F90FE80FD6BFEE5" box="[644,692,348,371]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
., 2001: 40
</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="9C837612ED4EFFF16E27FE80FC59FEE5" box="[819,902,348,371]" name="Taiwan" pageId="9" pageNumber="10">Taiwan</collectingCountry>
; references).
</treatmentCitationGroup>
</paragraph>
<paragraph id="E42B3682ED4EFFF16C1CFEA0FE1CFE25" blockId="9.[264,1323,284,467]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="C48411ACED4EFFF16C1CFEA0FE1CFE25" pageId="9" pageNumber="10">
<treatmentCitation id="65351093ED4EFFF16C1CFEA0FCD0FE05" author="Milne Edwards, H." box="[264,783,380,403]" page="170" pageId="9" pageNumber="10" year="1853">
<taxonomicName id="23944D01ED4EFFF16C1CFEA0FCD0FE05" ID-CoL="6KSW7" authority="H. Milne Edwards, 1853: 170" authorityName="H. Milne Edwards" authorityPageNumber="170" authorityYear="1853" box="[264,783,380,403]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="eydouxi">
<emphasis id="D6E0EA90ED4EFFF16C1CFEA0FE61FE05" box="[264,446,380,403]" italics="true" pageId="9" pageNumber="10">Grapsus eydouxi</emphasis>
H.
<bibRefCitation id="80054B73ED4EFFF16CFFFEA0FCD0FE05" author="Milne Edwards, H." box="[491,783,380,403]" pageId="9" pageNumber="10" pagination="163 - 228" refId="ref32020" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodiens. Annales des Sciences Naturelles, Zoologie, Paris, ser. 3, 20, 163 - 228, pl. 6 - 11." type="journal article" year="1853">Milne Edwards, 1853: 170</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[136] (
<collectingCountry id="9C837612ED4EFFF16E76FEA0FC7FFE05" box="[866,928,380,403]" name="Chile" pageId="9" pageNumber="10">Chile</collectingCountry>
; probably an erroneous locality, see distribution).
</treatmentCitationGroup>
</paragraph>
<paragraph id="E42B3682ED4EFFF16C1CFE60FB47FE45" blockId="9.[264,1323,284,467]" box="[264,1176,444,467]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="C48411ACED4EFFF16C1CFE60FB47FE45" box="[264,1176,444,467]" pageId="9" pageNumber="10">
<treatmentCitation id="65351093ED4EFFF16C1CFE60FC96FE45" author="Milne Edwards, H." box="[264,841,444,467]" page="173" pageId="9" pageNumber="10" year="1853">
<taxonomicName id="23944D01ED4EFFF16C1CFE60FC96FE45" ID-CoL="6PGKC" authority="H. Milne Edwards, 1853: 173" authorityName="H. Milne Edwards" authorityPageNumber="173" authorityYear="1853" box="[264,841,444,467]" class="Malacostraca" family="Grapsidae" genus="Leptograpsus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gonagrus">
<emphasis id="D6E0EA90ED4EFFF16C1CFE60FDDAFE45" box="[264,517,444,467]" italics="true" pageId="9" pageNumber="10">Leptograpsus gonagrus</emphasis>
H.
<bibRefCitation id="80054B73ED4EFFF16F38FE60FC96FE45" author="Milne Edwards, H." box="[556,841,444,467]" pageId="9" pageNumber="10" pagination="163 - 228" refId="ref32020" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodiens. Annales des Sciences Naturelles, Zoologie, Paris, ser. 3, 20, 163 - 228, pl. 6 - 11." type="journal article" year="1853">Milne Edwards, 1853: 173</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[139] (
<typeStatus id="3B2F8820ED4EFFF16E83FE60FC19FE45" box="[919,966,444,467]" pageId="9" pageNumber="10">type</typeStatus>
locality unknown).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC8E6509ED4EFFF36C1CFDD8FE72FC48" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="E42B3682ED4EFFF16C1CFDD8FB5BFDD0" blockId="9.[264,1156,516,582]" pageId="9" pageNumber="10">
<materialsCitation id="54FC3CDFED4EFFF16C1CFDD8FB5EFDD0" collectionCode="ANSP" collectorName="P. Callomon" pageId="9" pageNumber="10" specimenCode="CA 901" specimenCount="1" typeStatus="holotype">
<emphasis id="D6E0EA90ED4EFFF16C1CFDD8FE76FD8B" box="[264,425,516,541]" italics="true" pageId="9" pageNumber="10">
<typeStatus id="3B2F8820ED4EFFF16C1CFDD8FEE0FD8B" box="[264,319,516,541]" pageId="9" pageNumber="10">Type</typeStatus>
material
</emphasis>
Male preserved in alcohol (
<collectionCode id="8285AE47ED4EFFF16F6CFDF0FD1AFDD0" box="[632,709,556,582]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="9" pageNumber="10">ANSP</collectionCode>
<specimenCode id="B4329EF9ED4EFFF16FD9FDF0FCF5FDD0" box="[717,810,556,582]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="9" pageNumber="10" type="Museum">CA 901</specimenCode>
) (
<collectorName id="49615354ED4EFFF16E57FDF0FC08FDD0" box="[835,983,556,582]" pageId="9" pageNumber="10">P. Callomon</collectorName>
, pers. comm.)
</materialsCitation>
.
</paragraph>
<paragraph id="E42B3682ED4EFFF16C1CFDA0FE34FD03" blockId="9.[264,1323,636,902]" box="[264,491,636,661]" pageId="9" pageNumber="10">
<emphasis id="D6E0EA90ED4EFFF16C1CFDA0FE34FD03" box="[264,491,636,661]" italics="true" pageId="9" pageNumber="10">Material Examined</emphasis>
</paragraph>
<paragraph id="E42B3682ED4EFFF16C2CFD78FBDEFC10" blockId="9.[264,1323,636,902]" pageId="9" pageNumber="10">
<emphasis id="D6E0EA90ED4EFFF16C2CFD78FD85FD28" bold="true" box="[312,602,676,702]" pageId="9" pageNumber="10">East Pacific. California</emphasis>
:
<materialsCitation id="54FC3CDFED4EFFF16F7FFD78FC22FC10" accessionNumber="B12892" collectingDate="1905-06" collectingDateMax="1980-08-15" collectingDateMin="1905-06" collectionCode="MNHN, NSMT" collectorName="J. E. Benedict &amp; M. Takeda &amp; T. Habe" country="Japan" county="West Pacifi" location="Sagami Bay" municipality="Enoshima" pageId="9" pageNumber="10" specimenCode="USNM 32234" specimenCount="1" stateProvince="Chiba">
Pacific Grove, 1 M 27.0 x
<quantity id="236C9B67ED4EFFF16EB1FD78FBD1FD28" box="[933,1038,676,702]" metricMagnitude="-2" metricUnit="m" metricValue="3.08" pageId="9" pageNumber="10" unit="mm" value="30.8">30.8 mm</quantity>
(
<collectionCode id="8285AE47ED4EFFF1690AFD78FBA6FD28" box="[1054,1145,676,702]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="10">MNHN</collectionCode>
B12892); coll.
<collectorName id="49615354ED4EFFF16C1CFD10FE6EFD70" box="[264,433,716,742]" pageId="9" pageNumber="10">J. E. Benedict</collectorName>
,
<date id="902A1042ED4EFFF16CABFD10FDE4FD70" box="[447,571,716,742]" pageId="9" pageNumber="10" value="1905-06">
<collectingDate id="806EE9AAED4EFFF16CABFD10FDE4FD70" box="[447,571,716,742]" pageId="9" pageNumber="10" value="1905-06">June 1905</collectingDate>
</date>
, 3 M 26.5 x 29.837.2 x
<quantity id="236C9B67ED4EFFF16E64FD10FC02FD70" box="[880,989,716,742]" metricMagnitude="-2" metricUnit="m" metricValue="4.29" pageId="9" pageNumber="10" unit="mm" value="42.9">42.9 mm</quantity>
, 4 ov. F 27.3 x 30.131.9 x
<quantity id="236C9B67ED4EFFF16C1CFD28FEABFC98" box="[264,372,756,782]" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" pageId="9" pageNumber="10" unit="mm" value="36.8">36.8 mm</quantity>
, 2 F 25.6 x 28.4, 34.0 x
<quantity id="236C9B67ED4EFFF16F83FD28FCDCFC98" box="[663,771,756,782]" metricMagnitude="-2" metricUnit="m" metricValue="3.96" pageId="9" pageNumber="10" unit="mm" value="39.6">39.6 mm</quantity>
, plus numerous other specimens not examined (
<specimenCode id="B4329EF9ED4EFFF16C04FCC0FE63FCA0" box="[272,444,796,822]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">USNM 32234</specimenCode>
).
<emphasis id="D6E0EA90ED4EFFF16CDAFCC0FD85FCA0" bold="true" box="[462,602,796,822]" pageId="9" pageNumber="10">
<collectingCounty id="0D4A4E0EED4EFFF16CDAFCC0FD85FCA0" box="[462,602,796,822]" pageId="9" pageNumber="10">West Pacifi</collectingCounty>
</emphasis>
c.
<emphasis id="D6E0EA90ED4EFFF16F61FCC0FD1EFCA0" bold="true" box="[629,705,796,822]" pageId="9" pageNumber="10">
<collectingCountry id="9C837612ED4EFFF16F61FCC0FD1EFCA0" box="[629,705,796,822]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
</emphasis>
:
<collectingMunicipality id="044FACF8ED4EFFF16FC5FCC0FC96FCA0" box="[721,841,796,822]" pageId="9" pageNumber="10">Enoshima</collectingMunicipality>
,
<location id="E14B6059ED4EFFF16E40FCC0FC3CFCA0" LSID="urn:lsid:plazi:treatment:6C3D8794ED4FFFF36C1CF9B1FCC5F998:E14B6059ED4EFFF16E40FCC0FC3CFCA0" box="[852,995,796,822]" country="Japan" county="West Pacifi" municipality="Enoshima" name="Sagami Bay" pageId="9" pageNumber="10" stateProvince="Chiba">Sagami Bay</location>
, coll.
<collectorName id="49615354ED4EFFF1693CFCC0FB78FCA0" box="[1064,1191,796,822]" pageId="9" pageNumber="10">M. Takeda</collectorName>
,
<date id="902A1042ED4EFFF169A6FCC0FE9AFCC8" pageId="9" pageNumber="10" value="1980-08-15">
<collectingDate id="806EE9AAED4EFFF169A6FCC0FE9AFCC8" pageId="9" pageNumber="10" value="1980-08-15">15 August 1980</collectingDate>
</date>
, 1 M 31.9 x
<quantity id="236C9B67ED4EFFF16CC3FC98FDE0FCC8" box="[471,575,836,862]" metricMagnitude="-2" metricUnit="m" metricValue="3.66" pageId="9" pageNumber="10" unit="mm" value="36.6">36.6 mm</quantity>
(
<collectionCode id="8285AE47ED4EFFF16F59FC98FD7FFCC8" box="[589,672,836,862]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="9" pageNumber="10">NSMT</collectionCode>
Cr7516). —
<location id="E14B6059ED4EFFF16E22FC98FC55FCC8" LSID="urn:lsid:plazi:treatment:6C3D8794ED4FFFF36C1CF9B1FCC5F998:E14B6059ED4EFFF16E22FC98FC55FCC8" box="[822,906,836,862]" country="Japan" county="West Pacifi" municipality="Enoshima" name="Sotobo" pageId="9" pageNumber="10" stateProvince="Chiba">Sotobo</location>
,
<collectingRegion id="2650F860ED4EFFF16E82FC98FB81FCC8" box="[918,1118,836,862]" country="Japan" name="Chiba" pageId="9" pageNumber="10">Chiba Prefecture</collectingRegion>
, coll.
<collectorName id="49615354ED4EFFF169B7FC98FADDFCC8" box="[1187,1282,836,862]" pageId="9" pageNumber="10">T. Habe</collectorName>
,
<date id="902A1042ED4EFFF16819FC98FE46FC10" pageId="9" pageNumber="10" value="1975-03-25">
<collectingDate id="806EE9AAED4EFFF16819FC98FE46FC10" pageId="9" pageNumber="10" value="1975-03-25">25 March 1975</collectingDate>
</date>
, 2 M 16.8 x 18.9, 24.9 x
<quantity id="236C9B67ED4EFFF16FABFCB0FCF9FC10" box="[703,806,876,902]" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="9" pageNumber="10" unit="mm" value="29.3">29.3 mm</quantity>
(
<collectionCode id="8285AE47ED4EFFF16E22FCB0FC56FC10" box="[822,905,876,902]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="9" pageNumber="10">NSMT</collectionCode>
Cr 6822)
</materialsCitation>
.
</paragraph>
<paragraph id="E42B3682ED4EFFF16C1CFC60FEA0FC43" blockId="9.[264,1324,956,1742]" box="[264,383,956,981]" pageId="9" pageNumber="10">
<emphasis id="D6E0EA90ED4EFFF16C1CFC60FEA0FC43" box="[264,383,956,981]" italics="true" pageId="9" pageNumber="10">Diagnosis</emphasis>
</paragraph>
<paragraph id="E42B3682ED4EFFF16C2CFC38FD91FB50" blockId="9.[264,1324,956,1742]" pageId="9" pageNumber="10">
Carapace with dorsal surface weakly convex, striated on gastric, hepatic, branchial regions; striae without setae. Cardiac, intestinal regions smooth or only with faint striae. Lateral margins convergent posteriorly, with single tooth behind exorbital angle (
<figureCitation id="7CAF2A07ED4EFFF169D0FBE8FAC4FBD8" box="[1220,1307,1076,1102]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="9" pageNumber="10">Fig. 2a</figureCitation>
). Front 0.6 times exorbital width, anterior margin slightly sinuous with 2 lobules distally (
<figureCitation id="7CAF2A07ED4EFFF16C05FB58FEB5FB08" box="[273,362,1156,1182]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="9" pageNumber="10">Fig. 2b</figureCitation>
). Infraorbital margin weakly dentate on inner half, deeply dentate on outer half; outer notch deep (
<figureCitation id="7CAF2A07ED4EFFF16CCFFB70FD9EFB50" box="[475,577,1196,1222]" captionStart="FIGURE 14" captionStartId="47.[264,368,1403,1427]" captionTargetBox="[274,1309,325,1344]" captionTargetId="figure-211@47.[265,1322,294,1379]" captionTargetPageId="47" captionText="FIGURE 14. —Infraorbital margin: a) Pachygrapsus fakaravensis Rathbun (1907), ov. female 15.0 x 16.5 mm, Taiaro, French Polynesia (MNHN B25782); b) Pachygrapsus laevimanus Stimpson, 1858, neotype male 18.0 x 21.7 mm, Sydney, Australia (AM P7478); c) Pachygrapsus minutus A. Milne­Edwards, 1873, male 5.2 x 7.0 mm, Moruroa, French Polynesia (MNHN B29273); d) Pachygrapsus planifrons de Man, 1888, male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), male 12.1 x 15.3 mm, Taiaro, French Polynesia (MNHN B29279); f) Pachygrapsus corrugatus (von Martens, 1872) ov. female 13.6 x 15.2 mm, Virgin Is. (USNM 72340); g) Pachygrapsus crassipes Randall, 1840, male 31.9 x 36.6 mm, Sagami Bay, Japan (NSMT Cr7516); h) Pachygrapsus gracilis (de Saussure, 1858), ov. female 8.5 x 11 mm, Angola (MNHN B12903); i) Pachygrapsus loveridgei Chace, 1966, male 8.1 x 10 mm, St. Helena I. (MNHN B17836); j) Pachygrapsus marmoratus (Fabricius, 1787), female 12.7 x 14.3 mm, Préfailles, France (MNHN B22654); k) Pachygrapsus maurus (Lucas, 1846), female 12.7 x 14.3 mm, Banyuls, France (MNHN B12933); l) Pachygrapsus transversus (Gibbes, 1850), male 17.0 x 22.5 mm, French Guiana (MNHN B16013). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049262" httpUri="https://zenodo.org/record/5049262/files/figure.png" pageId="9" pageNumber="10">
Fig.
<quantity id="236C9B67ED4EFFF16F05FB70FD9EFB50" box="[529,577,1196,1222]" metricMagnitude="-2" metricUnit="kg" metricValue="1.4" pageId="9" pageNumber="10" unit="g" value="14.0">14g</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED4EFFF16C2CFB08FE62FA20" blockId="9.[264,1324,956,1742]" pageId="9" pageNumber="10">
Outer face of chelae smooth with faint longitudinal line near lower margin, becoming clearly noticeable only on outer face of fixed finger; upper margin slightly carinated; tip of fingers spoon­like, glabrous (
<figureCitation id="7CAF2A07ED4EFFF16F74FAF8FD6AFAA8" box="[608,693,1316,1342]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="9" pageNumber="10">Fig. 2c</figureCitation>
). Ambulatory legs almost totally glabrous; carpi, propodi with several short mobile spines; dactyli with longitudal rows of long mobile spines, ending with strong, corneous claw. Lower margin of P5 merus smooth with rounded distal angle (
<figureCitation id="7CAF2A07ED4EFFF16C43FA40FE70FA20" box="[343,431,1436,1462]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="9" pageNumber="10">Fig. 2d</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED4EFFF16C2CFA18FB7AF990" blockId="9.[264,1324,956,1742]" pageId="9" pageNumber="10">
Abdominal tergites smooth; sixth somite of abdomen plus telson triangular in male (
<figureCitation id="7CAF2A07ED4EFFF16C05FA30FEB8F990" box="[273,359,1516,1542]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="9" pageNumber="10">Fig. 2e</figureCitation>
). G1 bluntly truncated with small horn­like process distally (
<figureCitation id="7CAF2A07ED4EFFF16925FA30FB47F990" box="[1073,1176,1516,1542]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. Milne­Edwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, gl, from Chace, 1966, fig. 1011; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="9" pageNumber="10">
Fig.
<quantity id="236C9B67ED4EFFF1697CFA30FB47F990" box="[1128,1176,1516,1542]" metricMagnitude="-2" metricUnit="kg" metricValue="1.5" pageId="9" pageNumber="10" unit="g" value="15.0">15g</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED4EFFF16C2CF9C8FC50F9E8" blockId="9.[264,1324,956,1742]" pageId="9" pageNumber="10">Colour: very variable on photographs examined. Carapace green to almost totally black. Chelipeds light pink to purple or black, with a marbled pattern on the chelae. Ambulatory legs light green or purple to almost totally black.</paragraph>
<paragraph id="E42B3682ED4EFFF16C2CF950FDC4F958" blockId="9.[264,1324,956,1742]" pageId="9" pageNumber="10">
Measurements: large size; carapace of specimens examined ranging from 26.5 x
<quantity id="236C9B67ED4EFFF169ECF950FEEAF958" metricMagnitude="-2" metricUnit="m" metricValue="2.98" pageId="9" pageNumber="10" unit="mm" value="29.8">29.8 mm</quantity>
to 37.2 x
<quantity id="236C9B67ED4EFFF16CBFF968FDC8F958" box="[427,535,1716,1742]" metricMagnitude="-2" metricUnit="m" metricValue="4.29" pageId="9" pageNumber="10" unit="mm" value="42.9">42.9 mm</quantity>
.
</paragraph>
<paragraph id="E42B3682ED4EFFF16C1CF8D8FE48F88B" blockId="9.[264,1322,1796,1902]" box="[264,407,1796,1821]" pageId="9" pageNumber="10">
<emphasis id="D6E0EA90ED4EFFF16C1CF8D8FE48F88B" box="[264,407,1796,1821]" italics="true" pageId="9" pageNumber="10">Distribution</emphasis>
</paragraph>
<paragraph id="E42B3682ED4EFFF16C2CF8F0FE63F8F8" blockId="9.[264,1322,1796,1902]" pageId="9" pageNumber="10">
<emphasis id="D6E0EA90ED4EFFF16C2CF8F0FE15F8D0" bold="true" box="[312,458,1836,1862]" pageId="9" pageNumber="10">East Pacific</emphasis>
. Oregon and California,
<collectingCountry id="9C837612ED4EFFF16FE4F8F0FC93F8D0" box="[752,844,1836,1862]" name="Mexico" pageId="9" pageNumber="10">Mexico</collectingCountry>
, Galápagos.
<emphasis id="D6E0EA90ED4EFFF16EF7F8F0FBA2F8D0" bold="true" box="[995,1149,1836,1862]" pageId="9" pageNumber="10">West Pacific</emphasis>
.
<collectingCountry id="9C837612ED4EFFF16998F8F0FB0AF8D0" box="[1164,1237,1836,1862]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
,
<collectingCountry id="9C837612ED4EFFF169F4F8F0FAF9F8D0" box="[1248,1318,1836,1862]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
,
<collectingCountry id="9C837612ED4EFFF16C1CF888FE8DF8F8" box="[264,338,1876,1902]" name="South Korea" pageId="9" pageNumber="10">Korea</collectingCountry>
,
<collectingCountry id="9C837612ED4EFFF16C4AF888FE67F8F8" box="[350,440,1876,1902]" name="Taiwan" pageId="9" pageNumber="10">Taiwan</collectingCountry>
.
</paragraph>
<caption id="B0EB660AED4DFFF26C1CF9C8FD29F9EF" ID-DOI="http://doi.org/10.5281/zenodo.5049236" ID-Zenodo-Dep="5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="10" pageNumber="11" startId="10.[264,368,1556,1580]" targetBox="[278,1310,296,1513]" targetPageId="10">
<paragraph id="E42B3682ED4DFFF26C1CF9C8FD29F9EF" blockId="10.[264,1323,1556,1657]" pageId="10" pageNumber="11">
<emphasis id="D6E0EA90ED4DFFF26C1CF9C8FE53F9BA" bold="true" box="[264,396,1556,1580]" pageId="10" pageNumber="11">FIGURE 2.</emphasis>
<taxonomicName id="23944D01ED4DFFF26CBBF9C9FC8CF9BA" authority="Randall, 1840" authorityName="Randall" authorityYear="1840" box="[431,851,1557,1580]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4DFFF26CBBF9C9FD6EF9BA" box="[431,689,1557,1580]" italics="true" pageId="10" pageNumber="11">Pachygrapsus crassipes</emphasis>
<bibRefCitation id="80054B73ED4DFFF26FADF9C9FC8CF9BA" author="Randall, J. W." box="[697,851,1557,1580]" pageId="10" pageNumber="11" pagination="106 - 148" refId="ref33057" refString="Randall, J. W. (1840) Catalogue of the Crustacea brought by Thomas Nuttall and J. K. Townsend, from the west coast of North America and the Sandwich Islands, with description of such species as are apparently new, among which are included several species of different localities, previously existing in the collection of the Academy. Journal of the Academy of Natural Science of Philadelphia, 1839 (1840), 8 (1), 106 - 148, pl. 3 - 7." type="journal article" year="1840">Randall, 1840</bibRefCitation>
</taxonomicName>
. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892).
</paragraph>
</caption>
<paragraph id="E42B3682ED4DFFF36C2CF96CFD4AFE10" blockId="10.[264,1323,1712,1898]" lastBlockId="11.[264,1323,284,390]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
The
<typeStatus id="3B2F8820ED4DFFF26C7AF96CFE7FF95C" box="[366,416,1712,1738]" pageId="10" pageNumber="11">type</typeStatus>
locality given by
<bibRefCitation id="80054B73ED4DFFF26F6CF96CFCF2F95C" author="Randall, J. W." box="[632,813,1712,1738]" pageId="10" pageNumber="11" pagination="106 - 148" refId="ref33057" refString="Randall, J. W. (1840) Catalogue of the Crustacea brought by Thomas Nuttall and J. K. Townsend, from the west coast of North America and the Sandwich Islands, with description of such species as are apparently new, among which are included several species of different localities, previously existing in the collection of the Academy. Journal of the Academy of Natural Science of Philadelphia, 1839 (1840), 8 (1), 106 - 148, pl. 3 - 7." type="journal article" year="1840">Randall (1840)</bibRefCitation>
, the Hawaiian Is., is certainly erroneous as this large­size and common species has never again been recorded from the islands. The
<typeStatus id="3B2F8820ED4DFFF26C1CF8DCFEE5F88C" box="[264,314,1792,1818]" pageId="10" pageNumber="11">type</typeStatus>
specimen was most probably collected on the California coast.
<taxonomicName id="23944D01ED4DFFF2692AF8DDFE9DF8D4" authorityName="Randall" authorityYear="1840" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4DFFF2692AF8DDFE9DF8D4" italics="true" pageId="10" pageNumber="11">Pachygrapsus crassipes</emphasis>
</taxonomicName>
was also listed as occurring in
<collectingCountry id="9C837612ED4DFFF26FA9F8F4FD23F8D4" box="[701,764,1832,1858]" name="Chile" pageId="10" pageNumber="11">Chile</collectingCountry>
by
<bibRefCitation id="80054B73ED4DFFF26E3FF8F4FBFDF8D4" author="Rathbun, M. J." box="[811,1058,1832,1858]" pageId="10" pageNumber="11" pagination="1 - 461" refId="ref33490" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, i - xxii, 1 - 461, pl. 1 - 161." type="journal article" year="1918">Rathbun (1918: 241)</bibRefCitation>
based on the fact that this is the
<typeStatus id="3B2F8820ED4DFFF26C9DF88CFE63F8FC" box="[393,444,1872,1898]" pageId="10" pageNumber="11">type</typeStatus>
locality given for
<taxonomicName id="23944D01ED4DFFF26FB6F88DFB30F8FC" authority="H. Milne Edwards (1853: 170)" authorityName="H. Milne Edwards" authorityPageNumber="170" authorityYear="1853" box="[674,1263,1872,1898]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="eydouxi">
<emphasis id="D6E0EA90ED4DFFF26FB6F88DFCB4F8FC" box="[674,875,1873,1898]" italics="true" pageId="10" pageNumber="11">Grapsus eydouxi</emphasis>
H.
<bibRefCitation id="80054B73ED4DFFF26E88F88CFB30F8FC" author="Milne Edwards, H." box="[924,1263,1872,1898]" pageId="10" pageNumber="11" pagination="163 - 228" refId="ref32020" refString="Milne Edwards, H. (1853) Memoire sur la famille des Ocypodiens. Annales des Sciences Naturelles, Zoologie, Paris, ser. 3, 20, 163 - 228, pl. 6 - 11." type="journal article" year="1853">Milne Edwards (1853: 170)</bibRefCitation>
</taxonomicName>
. The
<typeStatus id="3B2F8820ED4CFFF36C1CFEC0FEE5FEA0" box="[264,314,284,310]" pageId="11" pageNumber="12">type</typeStatus>
of
<taxonomicName id="23944D01ED4CFFF36C73FEC0FE3AFEA3" authorityName="H. Milne Edwards" authorityYear="1853" box="[359,485,284,309]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="eydouxi">
<emphasis id="D6E0EA90ED4CFFF36C73FEC0FE3AFEA3" box="[359,485,284,309]" italics="true" pageId="11" pageNumber="12">G. eydouxi</emphasis>
</taxonomicName>
has not been found at the MNHN.
<bibRefCitation id="80054B73ED4CFFF36E8EFEC0FBABFEA0" author="Garth, J. S." box="[922,1140,284,310]" pageId="11" pageNumber="12" pagination="1 - 128" refId="ref30372" refString="Garth, J. S. (1957) Reports of the Lund University Chile Expedition 1948 - 49, n ° 29. The crustacea Decapoda Brachyura of Chile. Lunds Universitets A rsskrift, n. f., Avd. 2, 53 (7), 1 - 128." type="journal article" year="1957">Garth (1957: 105)</bibRefCitation>
considered the
<typeStatus id="3B2F8820ED4CFFF36C1CFE98FEE5FEC8" box="[264,314,324,350]" pageId="11" pageNumber="12">type</typeStatus>
locality of
<collectingCountry id="9C837612ED4CFFF36CD0FE98FDDBFEC8" box="[452,516,324,350]" name="Chile" pageId="11" pageNumber="12">Chile</collectingCountry>
for
<taxonomicName id="23944D01ED4CFFF36F2CFE98FD6BFECB" authorityName="H. Milne Edwards" authorityYear="1853" box="[568,692,324,349]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="eydouxi">
<emphasis id="D6E0EA90ED4CFFF36F2CFE98FD6BFECB" box="[568,692,324,349]" italics="true" pageId="11" pageNumber="12">G. eydouxi</emphasis>
</taxonomicName>
to be erroneous, and thus he did not include
<taxonomicName id="23944D01ED4CFFF369C4FE98FE9DFE13" authorityName="Randall" authorityYear="1840" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4CFFF369C4FE98FE9DFE13" italics="true" pageId="11" pageNumber="12">P. crassipes</emphasis>
</taxonomicName>
as part of the Chilean fauna.
</paragraph>
<paragraph id="E42B3682ED4CFFF36C1CFE60FDB3FE68" blockId="11.[264,620,444,510]" box="[264,620,444,510]" pageId="11" pageNumber="12">
<emphasis id="D6E0EA90ED4CFFF36C1CFE60FEBDFE43" box="[264,354,444,469]" italics="true" pageId="11" pageNumber="12">Habitat</emphasis>
Intertidal on rocky shores.
</paragraph>
<paragraph id="E42B3682ED4CFFF36C1CFDE8FEB1FDDB" blockId="11.[264,1324,564,1550]" box="[264,366,564,589]" pageId="11" pageNumber="12">
<emphasis id="D6E0EA90ED4CFFF36C1CFDE8FEB1FDDB" box="[264,366,564,589]" italics="true" pageId="11" pageNumber="12">Remarks</emphasis>
</paragraph>
<paragraph id="E42B3682ED4CFFF36C2CFD80FE72FC48" blockId="11.[264,1324,564,1550]" pageId="11" pageNumber="12">
<emphasis id="D6E0EA90ED4CFFF36C2CFD80FD80FDE3" box="[312,607,604,629]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="23944D01ED4CFFF36C2CFD80FD84FDE3" authorityName="Randall" authorityYear="1840" box="[312,603,604,629]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">Pachygrapsus crassipes</taxonomicName>
,
</emphasis>
together with
<taxonomicName id="23944D01ED4CFFF36E0EFD80FC48FDE3" baseAuthorityName="de Saussure" baseAuthorityYear="1858" box="[794,919,604,629]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="D6E0EA90ED4CFFF36E0EFD80FC48FDE3" box="[794,919,604,629]" italics="true" pageId="11" pageNumber="12">P. gracilis</emphasis>
</taxonomicName>
,
<taxonomicName id="23944D01ED4CFFF36EBFFD80FB8DFDE3" authorityName="Stimpson" authorityYear="1858" box="[939,1106,604,629]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="laevimanus">
<emphasis id="D6E0EA90ED4CFFF36EBFFD80FB8DFDE3" box="[939,1106,604,629]" italics="true" pageId="11" pageNumber="12">P. laevimanus</emphasis>
</taxonomicName>
,
<taxonomicName id="23944D01ED4CFFF36973FD80FADEFDE3" authorityName="Chace" authorityYear="1966" box="[1127,1281,604,629]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="loveridgei">
<emphasis id="D6E0EA90ED4CFFF36973FD80FADEFDE3" box="[1127,1281,604,629]" italics="true" pageId="11" pageNumber="12">P. loveridgei</emphasis>
</taxonomicName>
,
<taxonomicName id="23944D01ED4CFFF36801FD80FEBFFD0B" baseAuthorityName="Lucas" baseAuthorityYear="1846" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="maurus">
<emphasis id="D6E0EA90ED4CFFF36801FD80FEBFFD0B" italics="true" pageId="11" pageNumber="12">P. maurus</emphasis>
</taxonomicName>
, and
<emphasis id="D6E0EA90ED4CFFF36CB4FD58FD94FD0B" box="[416,587,644,669]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="23944D01ED4CFFF36CB4FD58FD98FD0B" baseAuthorityName="Gibbes" baseAuthorityYear="1850" box="[416,583,644,669]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="transversus">P. transversus</taxonomicName>
,
</emphasis>
belongs to the group of species with only one tooth behind the exorbital angle. It clearly differs from these species by the shape of its G1 (
<figureCitation id="7CAF2A07ED4CFFF3697BFD70FB19FD50" box="[1135,1222,684,710]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. Milne­Edwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, gl, from Chace, 1966, fig. 1011; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="11" pageNumber="12">Fig. 15</figureCitation>
). Of this group it is morphologically close to
<taxonomicName id="23944D01ED4CFFF36FBBFD08FC8DFD7B" baseAuthorityName="Gibbes" baseAuthorityYear="1850" box="[687,850,724,749]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="D6E0EA90ED4CFFF36FBBFD08FC8DFD7B" box="[687,850,724,749]" italics="true" pageId="11" pageNumber="12">P. transversus</emphasis>
</taxonomicName>
, another large­size species. The two can be separated by the presence of striae on the cardiac and intestinal regions of the carapace being marked only in
<taxonomicName id="23944D01ED4CFFF36F18FCF8FD6EFCAB" baseAuthorityName="Gibbes" baseAuthorityYear="1850" box="[524,689,804,829]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="D6E0EA90ED4CFFF36F18FCF8FD6EFCAB" box="[524,689,804,829]" italics="true" pageId="11" pageNumber="12">P. transversus</emphasis>
</taxonomicName>
(
<figureCitation id="7CAF2A07ED4CFFF36FD6FCF8FCC8FCA8" box="[706,791,804,830]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="11" pageNumber="12">Fig. 2a</figureCitation>
,
<figureCitation id="7CAF2A07ED4CFFF36E30FCF8FC8DFCA8" box="[804,850,804,830]" captionStart="FIGURE 13" captionStartId="45.[264,367,1640,1664]" captionTargetBox="[270,1287,290,1616]" captionTargetId="figure-103@45.[266,1288,280,1633]" captionTargetPageId="45" captionText="FIGURE 13. —Pachygrapsus transversus (Gibbes, 1850). Male 17 x 22.5 mm, French Guiana (MNHN B16013): a) dorsal view; b) front; c) left chela outer face, d) right P5, e) male abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049258" httpUri="https://zenodo.org/record/5049258/files/figure.png" pageId="11" pageNumber="12">13a</figureCitation>
); the longitudinal stria on the outer face of the chela is unclear in
<taxonomicName id="23944D01ED4CFFF36F3BFC90FD66FCF3" authorityName="Randall" authorityYear="1840" box="[559,697,844,869]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4CFFF36F3BFC90FD66FCF3" box="[559,697,844,869]" italics="true" pageId="11" pageNumber="12">P. crassipes</emphasis>
</taxonomicName>
, but deep in
<taxonomicName id="23944D01ED4CFFF36E46FC90FC2AFCF3" baseAuthorityName="Gibbes" baseAuthorityYear="1850" box="[850,1013,844,869]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="D6E0EA90ED4CFFF36E46FC90FC2AFCF3" box="[850,1013,844,869]" italics="true" pageId="11" pageNumber="12">P. transversus</emphasis>
</taxonomicName>
(
<figureCitation id="7CAF2A07ED4CFFF36911FC90FB85FCF0" box="[1029,1114,844,870]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="11" pageNumber="12">Fig. 2c</figureCitation>
,
<figureCitation id="7CAF2A07ED4CFFF36973FC90FB4CFCF0" box="[1127,1171,844,870]" captionStart="FIGURE 13" captionStartId="45.[264,367,1640,1664]" captionTargetBox="[270,1287,290,1616]" captionTargetId="figure-103@45.[266,1288,280,1633]" captionTargetPageId="45" captionText="FIGURE 13. —Pachygrapsus transversus (Gibbes, 1850). Male 17 x 22.5 mm, French Guiana (MNHN B16013): a) dorsal view; b) front; c) left chela outer face, d) right P5, e) male abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049258" httpUri="https://zenodo.org/record/5049258/files/figure.png" pageId="11" pageNumber="12">13c</figureCitation>
); the ventrodistal angle of the P5 merus is rounded in
<taxonomicName id="23944D01ED4CFFF36E16FCA8FC52FC1B" authorityName="Randall" authorityYear="1840" box="[770,909,884,909]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4CFFF36E16FCA8FC52FC1B" box="[770,909,884,909]" italics="true" pageId="11" pageNumber="12">P. crassipes</emphasis>
</taxonomicName>
, but it has spines in
<taxonomicName id="23944D01ED4CFFF36991FCA8FAF5FC1B" baseAuthorityName="Gibbes" baseAuthorityYear="1850" box="[1157,1322,884,909]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="D6E0EA90ED4CFFF36991FCA8FAF5FC1B" box="[1157,1322,884,909]" italics="true" pageId="11" pageNumber="12">P. transversus</emphasis>
</taxonomicName>
(
<figureCitation id="7CAF2A07ED4CFFF36C05FC40FEB7FC20" box="[273,360,924,950]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="11" pageNumber="12">Fig. 2d</figureCitation>
,
<figureCitation id="7CAF2A07ED4CFFF36C62FC40FE7AFC20" box="[374,421,924,950]" captionStart="FIGURE 13" captionStartId="45.[264,367,1640,1664]" captionTargetBox="[270,1287,290,1616]" captionTargetId="figure-103@45.[266,1288,280,1633]" captionTargetPageId="45" captionText="FIGURE 13. —Pachygrapsus transversus (Gibbes, 1850). Male 17 x 22.5 mm, French Guiana (MNHN B16013): a) dorsal view; b) front; c) left chela outer face, d) right P5, e) male abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049258" httpUri="https://zenodo.org/record/5049258/files/figure.png" pageId="11" pageNumber="12">13d</figureCitation>
); and the male abdomen is regularly triangular at its tip only in
<taxonomicName id="23944D01ED4CFFF3698BFC40FAF5FC23" authorityName="Randall" authorityYear="1840" box="[1183,1322,924,949]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4CFFF3698BFC40FAF5FC23" box="[1183,1322,924,949]" italics="true" pageId="11" pageNumber="12">P. crassipes</emphasis>
</taxonomicName>
(
<figureCitation id="7CAF2A07ED4CFFF36C05FC18FEBAFC48" box="[273,357,964,990]" captionStart="FIGURE 2" captionStartId="10.[264,368,1556,1580]" captionTargetBox="[278,1310,296,1513]" captionTargetId="figure-130@10.[266,1322,281,1528]" captionTargetPageId="10" captionText="FIGURE 2. —Pachygrapsus crassipes Randall, 1840. a) dorsal view; b) front; c) left chela outer face, d) left P5, e) male abdomen. a) male 26.5 x 29.8 mm, California (USNM 32234); be, male 27.0 x 30.8 mm, California (MNHN B12892)." figureDoi="http://doi.org/10.5281/zenodo.5049236" httpUri="https://zenodo.org/record/5049236/files/figure.png" pageId="11" pageNumber="12">Fig. 2e</figureCitation>
,
<figureCitation id="7CAF2A07ED4CFFF36C66FC18FE7FFC48" box="[370,416,964,990]" captionStart="FIGURE 13" captionStartId="45.[264,367,1640,1664]" captionTargetBox="[270,1287,290,1616]" captionTargetId="figure-103@45.[266,1288,280,1633]" captionTargetPageId="45" captionText="FIGURE 13. —Pachygrapsus transversus (Gibbes, 1850). Male 17 x 22.5 mm, French Guiana (MNHN B16013): a) dorsal view; b) front; c) left chela outer face, d) right P5, e) male abdomen." figureDoi="http://doi.org/10.5281/zenodo.5049258" httpUri="https://zenodo.org/record/5049258/files/figure.png" pageId="11" pageNumber="12">13e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AC8E6509ED4CFFF36C2CFC30FCC5F998" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="E42B3682ED4CFFF36C2CFC30FCC5F998" blockId="11.[264,1324,564,1550]" pageId="11" pageNumber="12">
<taxonomicName id="23944D01ED4CFFF36C2CFC30FD8DFB93" ID-CoL="4BSBZ" authorityName="Randall" authorityYear="1840" box="[312,594,1004,1029]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4CFFF36C2CFC30FD8DFB93" box="[312,594,1004,1029]" italics="true" pageId="11" pageNumber="12">Pachygrapsus crassipes</emphasis>
</taxonomicName>
is common on the Pacific coasts of North and South America from Oregon to
<collectingCountry id="9C837612ED4CFFF36CD1FBC8FDC1FBB8" box="[453,542,1044,1070]" name="Mexico" pageId="11" pageNumber="12">Mexico</collectingCountry>
and the Galápagos Is.
<bibRefCitation id="80054B73ED4CFFF36E3CFBC8FBFAFBB8" author="Stimpson, W." box="[808,1061,1044,1070]" pageId="11" pageNumber="12" pagination="1 - 240" refId="ref34180" refString="Stimpson, W. (1907) Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition, 1853 - 1856. Smithsonian Miscellaneous Collections, 49 (1717), 1 - 240, pl. 1 - 26." type="journal article" year="1907">Stimpson (1907: 116)</bibRefCitation>
was the first to record it from the west Pacific, indicating that he was “unable to find a distinguishing character, however minute, between the specimens from
<collectingCountry id="9C837612ED4CFFF36E64FBB8FC6CFBE8" box="[880,947,1124,1150]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
and those from California”.
<taxonomicName id="23944D01ED4CFFF36C1CFB50FDF6FB33" ID-CoL="4BSBZ" authorityName="Randall" authorityYear="1840" box="[264,553,1164,1189]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4CFFF36C1CFB50FDF6FB33" box="[264,553,1164,1189]" italics="true" pageId="11" pageNumber="12">Pachygrapsus crassipes</emphasis>
</taxonomicName>
was reported several times thereafter from the west Pacific, including
<collectingCountry id="9C837612ED4CFFF36C95FB68FE17FB58" box="[385,456,1204,1230]" name="South Korea" pageId="11" pageNumber="12">Korea</collectingCountry>
(
<bibRefCitation id="80054B73ED4CFFF36CC8FB68FD4EFB58" author="Rathbun, M. J." box="[476,657,1204,1230]" pageId="11" pageNumber="12" pagination="1 - 461" refId="ref33490" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, i - xxii, 1 - 461, pl. 1 - 161." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
),
<collectingCountry id="9C837612ED4CFFF36FBDFB68FD34FB58" box="[681,747,1204,1230]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
(
<bibRefCitation id="80054B73ED4CFFF36FEBFB68FC4DFB58" author="Sakai, T." box="[767,914,1204,1230]" pageId="11" pageNumber="12" pagination="1 - 251" refId="ref33764" refString="Sakai, T. (1976) Crabs of Japan and the adjacent seas. Tokyo, Kodansha, English volume, i - xxix, 773 pp.; Japanese volume, 461 pp.; plates volume, pl. 1 - 251." type="book chapter" year="1976">Sakai, 1976</bibRefCitation>
),
<collectingCountry id="9C837612ED4CFFF36EBEFB68FC2FFB58" box="[938,1008,1204,1230]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
(Dai &amp; Yang, 1911), and
<collectingCountry id="9C837612ED4CFFF36C1CFB00FE81FB60" box="[264,350,1244,1270]" name="Taiwan" pageId="11" pageNumber="12">Taiwan</collectingCountry>
(
<bibRefCitation id="80054B73ED4CFFF36C79FB00FDFFFB60" author="Ng, P. K. L. &amp; Wang, C. H. &amp; Ho, P. H. &amp; Shih, H. T." box="[365,544,1244,1270]" pageId="11" pageNumber="12" pagination="1 - 86" refId="ref32394" refString="Ng, P. K. L., Wang, C. H., Ho, P. H. &amp; Shih, H. T. (2001) An annotated checklist of the brachyuran crabs from Taiwan (Crustacea: Decapoda). National Taiwan Museum, Special Publication Series, 11, 1 - 86." type="journal article" year="2001">
Ng
<emphasis id="D6E0EA90ED4CFFF36C8DFB00FE13FB63" box="[409,460,1244,1269]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
., 2001
</bibRefCitation>
). This amphi­Pacific distribution is rather unusual, the fauna of the east and west Pacific normally being distinct. The G1 of Asian
<taxonomicName id="23944D01ED4CFFF36903FAD8FB7BFA8B" ID-CoL="4BSBZ" authorityName="Randall" authorityYear="1840" box="[1047,1188,1284,1309]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4CFFF36903FAD8FB7BFA8B" box="[1047,1188,1284,1309]" italics="true" pageId="11" pageNumber="12">P. crassipes</emphasis>
</taxonomicName>
appears to have never been figured or compared with the American
<taxonomicName id="23944D01ED4CFFF36EBFFAF0FBEAFAD3" ID-CoL="4BSBZ" authorityName="Randall" authorityYear="1840" box="[939,1077,1324,1349]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4CFFF36EBFFAF0FBEAFAD3" box="[939,1077,1324,1349]" italics="true" pageId="11" pageNumber="12">P. crassipes</emphasis>
</taxonomicName>
. The examination of the G1 of Japanese specimens (NSMT Cr7516, Cr 6822) shows them to be very similar to the American
<taxonomicName id="23944D01ED4CFFF36CA5FAA0FDE3FA03" ID-CoL="4BSBZ" authorityName="Randall" authorityYear="1840" box="[433,572,1404,1429]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crassipes">
<emphasis id="D6E0EA90ED4CFFF36CA5FAA0FDE3FA03" box="[433,572,1404,1429]" italics="true" pageId="11" pageNumber="12">P. crassipes</emphasis>
</taxonomicName>
. Despite this, preliminary DNA analyses by C. Schubart (pers. comm.) indicate genetic differences between the Asian and American populations and, if these are confirmed by additional data (morphology, larval development, colour pattern), the two could be considered separate species.
</paragraph>
</subSubSection>
</treatment>
</document>