283 lines
41 KiB
XML
283 lines
41 KiB
XML
<document id="9000523DA7141A00B334566570E1DE7F" ID-DOI="10.5281/zenodo.7710087" ID-ISSN="1942-1354" ID-Zenodo-Dep="7710087" ID-ZooBank="F5B8AFEF-30F7-43ED-86C0-D0DD03D3568A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678329485458" checkinUser="felipe" docAuthor="Heffern, Daniel, Botero, Juan Pablo & Santos-Silva, Antonio" docDate="2023" docId="0E45161CFFC59F4AFF14DCD3D327F806" docLanguage="en" docName="InsectaMundi.2023.0966.1–17.pdf" docOrigin="Insecta Mundi 2023 (966)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Pentanodes clavatus Heffern, Botero and Santos-Silva 2023, new species" docType="treatment" docVersion="3" lastPageNumber="7" masterDocId="F27C6E64FFC39F42FFD4DD70D718FF80" masterDocTitle="New species and taxonomical and geographical notes on Cerambycidae and Disteniidae (Coleoptera: Chrysomeloidea)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="5" updateTime="1699461707745" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="CCB5CDD2F0462D7D80E5EFF1051C8938">New species and taxonomical and geographical notes on Cerambycidae and Disteniidae (Coleoptera: Chrysomeloidea)</mods:title>
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<mods:name id="BCB8C89B3867C38BD071CD7755E59630" type="personal">
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<mods:namePart id="D2915D31F6D0D50176BB711585D2CEB2">Heffern, Daniel</mods:namePart>
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<mods:affiliation id="7F240D58CD6845F9CC007C0BF55169EB">10531 Goldfield Lane, Houston, TX 77064, USA</mods:affiliation>
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<mods:namePart id="9431345FAD6BEB5F68EDE0045F553EA6">Botero, Juan Pablo</mods:namePart>
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<mods:affiliation id="35789D11B3BBF30C67EA3DB133BD9092">Sistemática Molecular, Laboratorio de Entomología, Pontificia Universidad Javeriana. Bogotá</mods:affiliation>
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<mods:roleTerm id="25D211766DA9CD12BA887FD7BEAFAF73">Author</mods:roleTerm>
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<mods:namePart id="A81B4C0CB91EA36331CE6C5E77FAFD03">Santos-Silva, Antonio</mods:namePart>
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<mods:affiliation id="A3D83CBF7805946662EBFB614A9F6ECE">Museu de Zoologia, Universidade de São Paulo, São Paulo, SP, Brazil</mods:affiliation>
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<mods:title id="E2394FB814CF88DE5752A4F15196341A">Insecta Mundi</mods:title>
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<mods:part id="A0611A953F128D26F3FF2884330231A7">
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<mods:date id="CEAB045D051222738625A5917BDF872F">2023</mods:date>
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<mods:number id="DF38E8ECF26AC3A7B92E33FB54207B76">2023-01-06</mods:number>
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<mods:detail id="8ACDD5D50D50F618EBA45035CCB0D43A" type="volume">
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<mods:number id="94E566B3117EABC84152B985ED81FB10">2023</mods:number>
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<mods:number id="41CD0B8C7F9027CE147B3E63F373CC04">966</mods:number>
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<subSubSection id="CEF6F481FFC59F44FF14DCD3D312FE40" box="[192,1034,419,448]" pageId="6" pageNumber="5" type="nomenclature">
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<paragraph id="8653A70AFFC59F44FF14DCD3D312FE40" blockId="6.[192,1034,419,481]" box="[192,1034,419,448]" pageId="6" pageNumber="5">
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<heading id="DD1B1066FFC59F44FF14DCD3D312FE40" box="[192,1034,419,448]" fontSize="12" level="2" pageId="6" pageNumber="5" reason="2">
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<emphasis id="B4987B18FFC59F44FF14DCD3D312FE40" bold="true" box="[192,1034,419,448]" pageId="6" pageNumber="5">
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<taxonomicName id="41ECDC89FFC59F44FF14DCD3D471FE40" ID-CoL="BS93R" authority="Heffern, Botero and Santos-Silva" authorityName="Heffern, Botero and Santos-Silva" authorityYear="2023" box="[192,873,419,448]" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="clavatus" status="sp. nov.">
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<emphasis id="B4987B18FFC59F44FF14DCD3D6A6FE40" bold="true" box="[192,446,419,448]" italics="true" pageId="6" pageNumber="5">Pentanodes clavatus</emphasis>
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Heffern, Botero and Santos-Silva
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</taxonomicName>
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,
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<taxonomicNameLabel id="AFABC663FFC59F44FCA1DCD3D312FE40" box="[885,1034,419,448]" pageId="6" pageNumber="5" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="CEF6F481FFC59F4AFF14DCB7D47FFA0C" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5" type="description">
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<paragraph id="8653A70AFFC59F44FF14DCB7D67BFE61" blockId="6.[192,1034,419,481]" box="[192,355,455,481]" pageId="6" pageNumber="5">
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(
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<figureCitation id="1ED7BB8FFFC59F44FF1CDCB7D63DFE61" box="[200,293,455,481]" captionStart="Figures 1–7" captionStartId="3.[192,279,1833,1859]" captionTargetBox="[206,1425,213,1803]" captionTargetId="figure-61@3.[206,1425,213,1803]" captionTargetPageId="3" captionText="Figures 1–7. Pentanodes spp. 1–6) Pentanodes clavatus, holotype male: 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Head, frontal view; 5) Head and pronotum, oblique view; 6) Pronotum and basal antennomeres. 7) Pentanodes tropipennis (Chemsak, 1997), male from Costa Rica, pronotum and basal antennomeres." figureDoi="http://doi.org/10.5281/zenodo.7710089" httpUri="https://zenodo.org/record/7710089/files/figure.png" pageId="6" pageNumber="5">Fig. 1–6</figureCitation>
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,
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<figureCitation id="1ED7BB8FFFC59F44FEE4DCB7D643FE61" box="[304,347,455,481]" captionStart="Figures 8–14" captionStartId="5.[192,279,1833,1859]" captionTargetBox="[258,1374,213,1803]" captionTargetId="figure-52@5.[258,1374,213,1803]" captionTargetPageId="5" captionText="Figures 8–14. Pentanodes spp. 8–9) Pentanodes clavatus, paratype female: 8) Dorsal habitus; 9) Ventral habitus. 10–14) Pentanodes baldwini, holotype male: 10) Lateral habitus; 11) Dorsal habitus; 12) Ventral habitus; 13) Head, frontal view; 14) Eye lobe and pronotum." figureDoi="http://doi.org/10.5281/zenodo.7710091" httpUri="https://zenodo.org/record/7710091/files/figure.png" pageId="6" pageNumber="5">8–9</figureCitation>
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)
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</paragraph>
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<paragraph id="8653A70AFFC59F44FF14DC81D371FCBF" blockId="6.[191,1443,497,1927]" pageId="6" pageNumber="5">
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<emphasis id="B4987B18FFC59F44FF14DC81D51DFD8C" bold="true" box="[192,517,497,524]" pageId="6" pageNumber="5">
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Description.
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<typeStatus id="595719A8FFC59F44FE8DDC82D6DEFD8C" box="[345,454,498,524]" pageId="6" pageNumber="5" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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(
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<figureCitation id="1ED7BB8FFFC59F44FDC7DC81D56AFD8B" box="[531,626,497,523]" captionStart="Figures 1–7" captionStartId="3.[192,279,1833,1859]" captionTargetBox="[206,1425,213,1803]" captionTargetId="figure-61@3.[206,1425,213,1803]" captionTargetPageId="3" captionText="Figures 1–7. Pentanodes spp. 1–6) Pentanodes clavatus, holotype male: 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Head, frontal view; 5) Head and pronotum, oblique view; 6) Pronotum and basal antennomeres. 7) Pentanodes tropipennis (Chemsak, 1997), male from Costa Rica, pronotum and basal antennomeres." figureDoi="http://doi.org/10.5281/zenodo.7710089" httpUri="https://zenodo.org/record/7710089/files/figure.png" pageId="6" pageNumber="5">Fig. 1–6</figureCitation>
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). Integument mostly black dorsally, mostly dark brown ventrally; ventral surface of head gradually dark reddish brown toward prothorax; palpomeres reddish brown, lighter on apex; anteclypeus and labrum dark reddish brown; scape, pedicel, and antennomeres IX–XI dark brown; antennomeres III–IV dark reddish brown on narrow basal area, dark brown on remaining surface; antennomeres V–VI orangish brown on narrow basal area, dark brown on remaining surface; antennomere V orangish brown; antennomere VII orangish brown, except brown apical area. Elytra with arched eburneous band located about middle extending from near epipleural margin to near suture. Apex of mesoventral process, trochanters and base of metafemora dark reddish brown; pro- and mesofemora, remaining surface of metafemora dark brown; tibiae dark brown, slightly lighter toward apex on metatibiae; tarsomeres I–II dark brown; tarsomeres III–IV reddish brown; tarsomere V dark reddish brown, except reddish brown apex; tarsal claws reddish brown.
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</paragraph>
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<paragraph id="8653A70AFFC59F44FED4DE38D32CF8C2" blockId="6.[191,1443,497,1927]" pageId="6" pageNumber="5">
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<emphasis id="B4987B18FFC59F44FED4DE38D65DFCE2" bold="true" box="[256,325,840,866]" pageId="6" pageNumber="5">Head.</emphasis>
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Frons coarsely rugose-punctate on wide central area; sides elevated, carina-shaped, punctures close to eyes finer, slightly sparser than on wide central area; area close to antennal tubercles slightly depressed; with short, sparse yellowish-white setae on wide central area, slightly longer close to eyes; with one long, erect yellowish seta on each side close to postclypeus. Vertex and area behind antenna with sculpturing and setae as on wide central area of frons; with one long, erect yellowish seta on each side of vertex. Area behind eyes tumid, coarsely rugose-punctate close to eyes, subsmooth close to prothorax; with sparse, both short and long yellowish-white setae on tumid area, glabrous close to prothorax. Genae coarsely rugose-punctate, punctures and roughness finer than on wide central area of frons; with both short and decumbent, and long and somewhat short yellowishwhite setae. Gulamentum smooth, glabrous posteriorly, coarsely, transversely striate anteriorly, except slightly depressed, striate-punctate area close to anterior margin; with sparse, both short and decumbent, and long and erect yellowish-white setae. Maxillary palpomere IV and labial palpomere III securiform. Wide central area of postclypeus coarsely rugose-punctate, punctures finer, shallower, and roughness finer than on central area of frons; with short, sparse, decumbent yellowish-white setae. Sides of postclypeus smooth, glabrous. Labrum glabrous close to anteclypeus, with long tuft of yellowish-brown setae on each side of central area, and short, bristly yellowish-white setae on remaining surface. Anterior 2/3 of outer surface of mandibles coarsely rugose-punctate, with sparse, short, decumbent yellowish-white setae, and thick, long, erect yellowish-white setae interspersed; posterior third smooth, glabrous. Superior area of eyes slightly rounded, projected upward; in frontal view, distance between eyes 0.62 times distance between outer margins of eyes. Antennae 1.55 times elytral length, almost reaching elytral apex. Scape subcylindrical; finely, sparsely punctate dorsally, except smooth apex, subsmooth on remaining surface; with sparse, short, decumbent brownish setae and long, erect yellowish-white setae interspersed dorsally, except glabrous smooth area, glabrous on remaining surface, except a few long, erect yellowish-white setae on posterior third of ventral surface. Pedicel with sparse, long, erect yellowish-white setae throughout, and a few short, decumbent setae of same color interspersed. Antennomeres III–VI pedunculateclavate, peduncle of III longer; with short, decumbent white setae, not obscuring integument, and moderately long, erect setae of same color interspersed dorsally, a few short, decumbent white setae laterally, and long, erect yellowish-white setae on apex of dorsal and ventral surfaces. Antennomeres VII–XI with abundant whitish pubescence not obscuring integument, and short, erect setae of same color interspersed; dorsal and ventral apex of VII with long, erect yellowish-white setae, and dorsal apex of VII with moderately long, erect yellowish-white setae on dorsal apex. Antennal formula based on length of antennomere III: scape = 0.89; pedicel = 0.20; IV = 0.72; V = 0.76; VI = 0.72; VII = 0.58; VIII = 0.48; IX = 0.48; X = 0.41; XI = 0.52.
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</paragraph>
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<paragraph id="8653A70AFFC59F4AFED4DA3AD6C1FB12" blockId="6.[191,1443,497,1927]" lastBlockId="8.[191,1442,214,1308]" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5">
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<emphasis id="B4987B18FFC59F44FED4DA3AD643F8E4" bold="true" box="[256,347,1866,1892]" pageId="6" pageNumber="5">Thorax.</emphasis>
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Prothorax distinctly longer than wide; sides rounded between apex of anterior and base of posterior quarter, distinctly narrowed on posterior quarter. Pronotum coarsely, longitudinally striate-punctate on wide central area; striae denser centrally; coarsely, shallowly punctate close to anterior margin; coarsely rugose-punctate on posterior quarter, except subsmooth on wide central area close to posterior margin; anterior 3/4 with a few short, decumbent brownish setae, and sparse, long, erect yellowish-white setae laterally; posterior quarter with yellowish-white pubescence not obscuring integument. Sides of prothorax longitudinally striate-punctate close to pronotum, sculpturing finer than on pronotum, gradually smooth toward prosternum; with a few long, erect yellowish-white setae superiorly, glabrous inferiorly. Prosternum sparsely, coarsely, shallowly punctate, except central area close to procoxal cavities with abundant, somewhat fine punctures, and finer striae on central apex of anterior third; with a few long and short yellowish-white setae, absent on wide central area, except abundant yellowish-white pubescence not obscuring integument posteriorly close to procoxal cavities. Prosternal process distinctly narrowed centrally, strongly widened on posterior third; narrowest area 0.14 times procoxal width; with a few short brownish setae. Mesoventrite strongly, obliquely inclined from anterior quarter toward mesoventral process; with white pubescence not obscuring integument anteriorly and laterally, glabrous on remaining surface; with a few long, erect yellowish-white setae near mesocoxal cavities. Mesanepisternum with dense white pubescence; mesepimeron with dense white pubescence close to elytra, glabrous close to mesocoxae. Mesoventral process with white pubescence not obscuring integument; apex widened, bilobed, strongly emarginate centrally; narrowest area 0.42 times mesocoxal width. Metanepisternum almost glabrous, except apex with dense white pubescence. Metaventrite with a few short yellowish setae, and a few long, erect white setae, except dense white pubescence on sides of posterior area. Scutellum with short, decumbent, sparse yellowish-white setae.
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<emphasis id="B4987B18FFCB9F4AFA85DE50D2B8FCBA" bold="true" box="[1361,1440,800,826]" pageId="8" pageNumber="7">Elytra.</emphasis>
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Gradually inclined from base to near middle, then convex toward apex; centrobasal crest elevated, convex dorsally; anterior 3/4 coarsely, sparsely punctate; posterior quarter finely, sparsely punctate, punctures distinctly more abundant than on anterior 3/4; punctures on anterior 3/4 with long, erect yellowish-white seta, nearly all seta yellower basally, but some setae entirely yellowish-brown; posterior quarter with moderately abundant and short yellowish-white setae, and long, erect setae of same color interspersed.
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<emphasis id="B4987B18FFCB9F4AFC27DEBCD328FC66" bold="true" box="[1011,1072,972,998]" pageId="8" pageNumber="7">Legs.</emphasis>
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Femora with sparse, long, erect whitish setae dorsally and laterally, glabrous ventrally. Tibiae with sparse, short, fine yellowish-white setae, and moderately abundant, long, erect white setae, except sides of posterior half of protibiae with abundant white pubescence, bristly, yellowish pubescence on ventral surface of posterior half of protibiae, and somewhat abundant yellowish-white pubescence on ventral surface of posterior third of meso- and metatibiae. Metatarsomere I as long as II–III together.
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</paragraph>
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<caption id="D293F782FFC49F45FF14DB95D29EF806" ID-DOI="http://doi.org/10.5281/zenodo.7710093" ID-Zenodo-Dep="7710093" httpUri="https://zenodo.org/record/7710093/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,278,1765,1791]" targetBox="[288,1343,217,1736]" targetPageId="7">
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<paragraph id="8653A70AFFC49F45FF14DB95D29EF806" blockId="7.[191,1441,1765,1927]" pageId="7" pageNumber="6">
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<emphasis id="B4987B18FFC49F45FF14DB95D670F97F" bold="true" box="[192,360,1765,1791]" pageId="7" pageNumber="6">Figures 15–22.</emphasis>
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<taxonomicName id="41ECDC89FFC49F45FEBADB96D515F880" authorityName="Latreille" authorityYear="1802" box="[366,525,1766,1792]" class="Insecta" family="Cerambycidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="family">Cerambycidae</taxonomicName>
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spp.
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<emphasis id="B4987B18FFC49F45FD90DB95D571F97F" bold="true" box="[580,617,1765,1791]" pageId="7" pageNumber="6">15)</emphasis>
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<taxonomicName id="41ECDC89FFC49F45FDA4DB96D4C1F880" authority="(Bates, 1885)" authorityName="sensu Giesbert and Chemsak" authorityYear="1997" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[624,985,1765,1792]" class="Insecta" family="Cerambycidae" genus="Euderces" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cribellatus">
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<emphasis id="B4987B18FFC49F45FDA4DB96D45BF880" box="[624,835,1766,1792]" italics="true" pageId="7" pageNumber="6">Euderces cribellatus</emphasis>
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(
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<bibRefCitation id="E27DDAFBFFC49F45FC84DB96D4C9F880" author="Bates HW" box="[848,977,1765,1792]" pageId="7" pageNumber="6" pagination="249 - 436" refId="ref10368" refString="Bates HW. 1885. Supplement to Longicornia. p. 249 - 436. In: Godman FD, Salvin O (eds.). Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis; London. xii + 525 p." type="book chapter" year="1885">Bates, 1885</bibRefCitation>
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)
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</taxonomicName>
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, female from Panama, dorsal habitus;
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<emphasis id="B4987B18FFC49F45FAAEDB95D287F97F" bold="true" box="[1402,1439,1765,1791]" pageId="7" pageNumber="6">16)</emphasis>
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<taxonomicName id="41ECDC89FFC49F45FF14DA77D429F8A1" authority="sensu Giesbert and Chemsak (1997)" authorityName="sensu Giesbert and Chemsak" authorityYear="1997" baseAuthorityName="Bates" baseAuthorityYear="1885" box="[192,817,1799,1825]" class="Insecta" family="Cerambycidae" genus="Euderces" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cribellatus">
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<emphasis id="B4987B18FFC49F45FF14DA77D6C0F8A1" box="[192,472,1799,1825]" italics="true" pageId="7" pageNumber="6">Euderces cribellatus sensu</emphasis>
|
||
<bibRefCitation id="E27DDAFBFFC49F45FE34DA77D429F8A1" author="Giesbert EF & Chemsak, JA" box="[480,817,1799,1825]" pageId="7" pageNumber="6" pagination="211 - 286" refId="ref10527" refString="Giesbert EF, Chemsak, JA. 1997. A review of the genus Euderces LeConte (Coleoptera, Cerambycidae, Tillomorphini). Proceedings of the California Academy of Sciences 49 (8): 211 - 286." type="journal article" year="1997">Giesbert and Chemsak (1997)</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B4987B18FFC49F45FCE9DA77D495F8A1" bold="true" box="[829,909,1799,1825]" pageId="7" pageNumber="6">17–18)</emphasis>
|
||
<taxonomicName id="41ECDC89FFC49F45FC47DA77D215F8A1" authority="McCarty, 2001" authorityName="McCarty" authorityYear="2001" box="[915,1293,1799,1825]" class="Insecta" family="Cerambycidae" genus="Oreodera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="noguerai">
|
||
<emphasis id="B4987B18FFC49F45FC47DA77D347F8A1" box="[915,1119,1799,1825]" italics="true" pageId="7" pageNumber="6">Oreodera noguerai</emphasis>
|
||
<bibRefCitation id="E27DDAFBFFC49F45FBBCDA77D215F8A1" author="McCarty JD" box="[1128,1293,1799,1825]" pageId="7" pageNumber="6" pagination="13 - 34" refId="ref10655" refString="McCarty JD. 2001. A review of the Mexican species of Oreodera Audinet-Serville (Coleoptera, Cerambycidae). Occasional Papers of the Consortium Coleopterorum 4 (1): 13 - 34." type="journal article" year="2001">McCarty, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
, female from Mexico (Jalisco):
|
||
<emphasis id="B4987B18FFC49F45FEAADA58D6BBF8C2" bold="true" box="[382,419,1832,1858]" pageId="7" pageNumber="6">17)</emphasis>
|
||
Dorsal habitus;
|
||
<emphasis id="B4987B18FFC49F45FD82DA58D563F8C2" bold="true" box="[598,635,1832,1858]" pageId="7" pageNumber="6">18)</emphasis>
|
||
Apex of protibia.
|
||
<emphasis id="B4987B18FFC49F45FCEBDA58D47CF8C2" bold="true" box="[831,868,1832,1858]" pageId="7" pageNumber="6">19)</emphasis>
|
||
<taxonomicName id="41ECDC89FFC49F45FCBFDA59D27AF8C3" authority="Martins and Monne, 1993" authorityName="Martins and Monne" authorityYear="1993" box="[875,1378,1833,1859]" class="Insecta" family="Cerambycidae" genus="Oreodera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="albilatera">
|
||
<emphasis id="B4987B18FFC49F45FCBFDA59D324F8C3" box="[875,1084,1833,1859]" italics="true" pageId="7" pageNumber="6">Oreodera albilatera</emphasis>
|
||
Martins and Monné, 1993
|
||
</taxonomicName>
|
||
, apex of protibia.
|
||
<emphasis id="B4987B18FFC49F45FEE9DA3AD67BF8E4" bold="true" box="[317,355,1866,1892]" pageId="7" pageNumber="6">20)</emphasis>
|
||
<taxonomicName id="41ECDC89FFC49F45FEBDDA3AD4B4F8E5" authority="Heffern, Santos-Silva and Botero, 2019" authorityName="Heffern, Santos-Silva and Botero" authorityYear="2019" box="[361,940,1866,1893]" class="Insecta" family="Cerambycidae" genus="Adetus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="croton">
|
||
<emphasis id="B4987B18FFC49F45FEBDDA3AD6E6F8E4" box="[361,510,1866,1892]" italics="true" pageId="7" pageNumber="6">Adetus croton</emphasis>
|
||
<bibRefCitation id="E27DDAFBFFC49F45FDD7DA3AD4B4F8E5" author="Heffern D & Santos-Silva A & Botero JP" box="[515,940,1866,1893]" pageId="7" pageNumber="6" pagination="561 - 574" refId="ref10567" refString="Heffern D, Santos-Silva A, Botero JP. 2019. A new genus and two new species of Apomecynini, a new species of Desmiphorini, and new records in Lamiinae and Disteniidae (Coleoptera). Zootaxa 4691 (5): 561 - 574." type="journal article" year="2019">Heffern, Santos-Silva and Botero, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
, male from Nicaragua.
|
||
<emphasis id="B4987B18FFC49F45FB7DDA3AD3D6F8E4" bold="true" box="[1193,1230,1866,1892]" pageId="7" pageNumber="6">21)</emphasis>
|
||
<taxonomicName id="41ECDC89FFC49F45FB01DA3BD6C7F806" authority="Martins and Galileo, 1991" authorityName="Martins and Galileo" authorityYear="1991" class="Insecta" family="Cerambycidae" genus="Icimauna" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="sarauaia">
|
||
<emphasis id="B4987B18FFC49F45FB01DA3BD2B9F8E4" box="[1237,1441,1867,1892]" italics="true" pageId="7" pageNumber="6">Icimauna sarauaia</emphasis>
|
||
<bibRefCitation id="E27DDAFBFFC49F45FF6BDA1CD6C7F806" author="Martins UR & Galileo MHM" box="[191,479,1900,1926]" pageId="7" pageNumber="6" pagination="813 - 824" refId="ref10615" refString="Martins UR, Galileo MHM. 1991. O genero Tyrinthia Bates, 1866 e generos afins (Coleoptera, Cerambycidae, Lamiinae, Hemilophini). Revista Brasileira de Entomologia 35 (4): 813 - 824." type="journal article" year="1991">Martins and Galileo, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
, female from Peru.
|
||
<emphasis id="B4987B18FFC49F45FD63DA1CD5C5F806" bold="true" box="[695,733,1900,1926]" pageId="7" pageNumber="6">22)</emphasis>
|
||
<taxonomicName id="41ECDC89FFC49F45FD37DA1CD33BF806" authority="Bates, 1881" authorityName="Bates" authorityYear="1881" box="[739,1059,1900,1927]" class="Insecta" family="Cerambycidae" genus="Phoebe" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="mexicana">
|
||
<emphasis id="B4987B18FFC49F45FD37DA1CD487F806" box="[739,927,1900,1926]" italics="true" pageId="7" pageNumber="6">Phoebe mexicana</emphasis>
|
||
<bibRefCitation id="E27DDAFBFFC49F45FC71DA1DD33BF806" author="Bates HW" box="[933,1059,1900,1927]" pageId="7" pageNumber="6" pagination="153 - 224" refId="ref10314" refString="Bates HW. 1881. Longicornia, Fam. Cerambycidae [continuation]. p. 153 - 224. In: Godman FD, Salvin O (eds.). Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis; London. xii + 525 p." type="book chapter" year="1881">Bates, 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
, female from Mexico (Chiapas).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8653A70AFFCB9F4AFED4D9EAD661FB57" blockId="8.[191,1442,214,1308]" pageId="8" pageNumber="7">
|
||
<emphasis id="B4987B18FFCB9F4AFED4D9EAD663FB34" bold="true" box="[256,379,1178,1204]" pageId="8" pageNumber="7">Abdomen.</emphasis>
|
||
Ventrites with sparse, decumbent yellowish-white setae, and a few long, erect white setae interspersed laterally.
|
||
</paragraph>
|
||
<paragraph id="8653A70AFFCB9F4AFED4D990D445FA9C" blockId="8.[191,1442,214,1308]" pageId="8" pageNumber="7">
|
||
<emphasis id="B4987B18FFCB9F4AFED4D990D64CFB7A" bold="true" box="[256,340,1248,1274]" pageId="8" pageNumber="7">Female</emphasis>
|
||
(
|
||
<figureCitation id="1ED7BB8FFFCB9F4AFEB5D990D6D8FB7A" box="[353,448,1248,1274]" captionStart="Figures 8–14" captionStartId="5.[192,279,1833,1859]" captionTargetBox="[258,1374,213,1803]" captionTargetId="figure-52@5.[258,1374,213,1803]" captionTargetPageId="5" captionText="Figures 8–14. Pentanodes spp. 8–9) Pentanodes clavatus, paratype female: 8) Dorsal habitus; 9) Ventral habitus. 10–14) Pentanodes baldwini, holotype male: 10) Lateral habitus; 11) Dorsal habitus; 12) Ventral habitus; 13) Head, frontal view; 14) Eye lobe and pronotum." figureDoi="http://doi.org/10.5281/zenodo.7710091" httpUri="https://zenodo.org/record/7710091/files/figure.png" pageId="8" pageNumber="7">Fig. 8–9</figureCitation>
|
||
). Similar to male, differing especially by the antennal shape (generic feature), antennae 1.2 times elytral length, reaching about posterior third of elytra.
|
||
</paragraph>
|
||
<paragraph id="8653A70AFFCB9F4AFF14D85CD47FFA0C" blockId="8.[192,1440,1324,1420]" pageId="8" pageNumber="7">
|
||
<emphasis id="B4987B18FFCB9F4AFF14D85CD42AFAC6" bold="true" box="[192,818,1324,1351]" pageId="8" pageNumber="7">
|
||
Dimensions in mm (
|
||
<typeStatus id="595719A8FFCB9F4AFE63D85CD538FAC6" box="[439,544,1324,1350]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
male/
|
||
<typeStatus id="595719A8FFCB9F4AFDBED85CD5CAFAC6" box="[618,722,1324,1350]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
|
||
female)
|
||
</emphasis>
|
||
. Total length, 5.05/5.80; prothoracic length, 1.30/1.55; anterior prothoracic width, 0.80/1.00; posterior prothoracic width, 0.75/0.85; maximum prothoracic width, 0.90/1.15; humeral width, 1.10/1.40; elytral length, 2.95/3.60.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CEF6F481FFCB9F4AFF14D8ECD220F99D" pageId="8" pageNumber="7" type="materials_examined">
|
||
<paragraph id="8653A70AFFCB9F4AFF14D8ECD220F99D" blockId="8.[192,1440,1436,1566]" pageId="8" pageNumber="7">
|
||
<emphasis id="B4987B18FFCB9F4AFF14D8ECD673FA36" bold="true" box="[192,363,1436,1462]" pageId="8" pageNumber="7">Type material.</emphasis>
|
||
<materialsCitation id="3684AD57FFCB9F4AFEA7D8ECD569FA59" collectingDate="2004-12-15" collectingDateMax="2004-12-17" collectingDateMin="2004-12-15" collectionCode="TAMU, DHCO" collectorName="Z. Mracek" country="Ecuador" location="Tandapi" municipality="Santo Domingo" pageId="8" pageNumber="7" specimenCount="1" stateProvince="Pichincha" typeStatus="holotype">
|
||
<typeStatus id="595719A8FFCB9F4AFEA7D8ECD6C3FA36" box="[371,475,1436,1462]" pageId="8" pageNumber="7" type="holotype">Holotype</typeStatus>
|
||
male from
|
||
<collectingCountry id="FEFBE79AFFCB9F4AFDB0D8ECD5F2FA36" box="[612,746,1436,1462]" name="Ecuador" pageId="8" pageNumber="7">ECUADOR</collectingCountry>
|
||
,
|
||
<collectingRegion id="442869E8FFCB9F4AFD23D8ECD48BFA36" box="[759,915,1436,1462]" country="Ecuador" name="Pichincha" pageId="8" pageNumber="7">PICHINCHA</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="66373D70FFCB9F4AFC76D8ECD34FFA36" box="[930,1111,1436,1462]" pageId="8" pageNumber="7">Santo Domingo</collectingMunicipality>
|
||
,
|
||
<location id="8333F1D1FFCB9F4AFBB0D8ECD3D9FA36" LSID="urn:lsid:plazi:treatment:0E45161CFFC59F4AFF14DCD3D327F806:8333F1D1FFCB9F4AFBB0D8ECD3D9FA36" box="[1124,1217,1436,1462]" country="Ecuador" municipality="Santo Domingo" name="Tandapi" pageId="8" pageNumber="7" stateProvince="Pichincha">Tandapi</location>
|
||
,
|
||
<date id="F25281CAFFCB9F4AFB19D8ECD264FA37" box="[1229,1404,1436,1463]" pageId="8" pageNumber="7" value="2004-12-15" valueMax="2004-12-17" valueMin="2004-12-15">
|
||
<collectingDate id="E2167822FFCB9F4AFB19D8ECD264FA37" box="[1229,1404,1436,1463]" pageId="8" pageNumber="7" value="2004-12-15" valueMax="2004-12-17" valueMin="2004-12-15">15–17.XII.2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="2B19C2DCFFCB9F4AFA5DD8EDD60DFA58" pageId="8" pageNumber="7">Z. Mráček</collectorName>
|
||
leg. (
|
||
<collectionCode id="E0FD3FCFFFCB9F4AFE9BD8CFD6BAFA59" box="[335,418,1471,1497]" pageId="8" pageNumber="7">TAMU</collectionCode>
|
||
, formerly
|
||
<collectionCode id="E0FD3FCFFFCB9F4AFDC7D8CFD571FA59" box="[531,617,1471,1497]" pageId="8" pageNumber="7">DHCO</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="595719A8FFCB9F4AFDAED8CFD5C4FA59" box="[634,732,1471,1497]" pageId="8" pageNumber="7" type="paratype">Paratype</typeStatus>
|
||
female,
|
||
<collectingCountry id="FEFBE79AFFCB9F4AFCEDD8CFD4A7FA59" box="[825,959,1471,1497]" name="Ecuador" pageId="8" pageNumber="7">ECUADOR</collectingCountry>
|
||
,
|
||
<collectingRegion id="442869E8FFCB9F4AFC1DD8CFD37DFA59" box="[969,1125,1471,1497]" country="Ecuador" name="Pichincha" pageId="8" pageNumber="7">PICHINCHA</collectingRegion>
|
||
: Santo Domingo de los Colorados,
|
||
<date id="F25281CAFFCB9F4AFECED891D670FA7B" box="[282,360,1505,1531]" pageId="8" pageNumber="7" value="1981-05">V.1981</date>
|
||
, no collector indicated (
|
||
<collectionCode id="E0FD3FCFFFCB9F4AFDAFD891D5D0FA7B" box="[635,712,1505,1531]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="7" type="Museum">MZSP</collectionCode>
|
||
). The
|
||
<typeStatus id="595719A8FFCB9F4AFCDAD891D477FA7B" box="[782,879,1505,1531]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
|
||
female has an additional label with the following information: “On
|
||
<taxonomicName id="41ECDC89FFCB9F4AFE5DDB74D6E5F99E" box="[393,509,1540,1566]" class="Magnoliopsida" family="Rosaceae" genus="Prunus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="7" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B4987B18FFCB9F4AFE5DDB74D6C0F99D" box="[393,472,1540,1565]" italics="true" pageId="8" pageNumber="7">Prunus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
“Kapuli.” This tree is typical of high altitude and is introduced in this city.”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CEF6F481FFCB9F4AFF14DB5ED38FF9C8" box="[192,1175,1582,1608]" pageId="8" pageNumber="7" type="etymology">
|
||
<paragraph id="8653A70AFFCB9F4AFF14DB5ED38FF9C8" blockId="8.[192,1175,1582,1608]" box="[192,1175,1582,1608]" pageId="8" pageNumber="7">
|
||
<emphasis id="B4987B18FFCB9F4AFF14DB5ED65CF9C8" bold="true" box="[192,324,1582,1608]" pageId="8" pageNumber="7">Etymology.</emphasis>
|
||
Latin, “clavatus” (club-shaped); allusive to the shape of the antennomere III.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CEF6F481FFCB9F4AFF14DB28D327F806" pageId="8" pageNumber="7" type="discussion">
|
||
<paragraph id="8653A70AFFCB9F4AFF14DB28D327F806" blockId="8.[192,1442,1624,1926]" pageId="8" pageNumber="7">
|
||
<emphasis id="B4987B18FFCB9F4AFF14DB28D637F9F2" bold="true" box="[192,303,1624,1650]" pageId="8" pageNumber="7">Remarks.</emphasis>
|
||
<taxonomicName id="41ECDC89FFCB9F4AFEE3DB28D50DF9F2" authority="Heffern & Botero & Santos-Silva, 2023" authorityName="Heffern & Botero & Santos-Silva" authorityYear="2023" box="[311,533,1624,1650]" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="clavatus" status="sp. nov.">
|
||
<emphasis id="B4987B18FFCB9F4AFEE3DB28D50DF9F2" box="[311,533,1624,1650]" italics="true" pageId="8" pageNumber="7">Pentanodes clavatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B4987B18FFCB9F4AFDC9DB28D5BFF9F2" bold="true" box="[541,679,1624,1650]" pageId="8" pageNumber="7">
|
||
<taxonomicNameLabel id="AFABC663FFCB9F4AFDC9DB28D5BFF9F2" box="[541,679,1624,1650]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="41ECDC89FFCB9F4AFC96DB29D3B9F9F2" authority="(Chemsak, 1997)" baseAuthorityName="Chemsak" baseAuthorityYear="1997" box="[834,1185,1624,1650]" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="tropipennis">
|
||
<emphasis id="B4987B18FFCB9F4AFC96DB29D4CFF9F2" box="[834,983,1625,1650]" italics="true" pageId="8" pageNumber="7">P. tropipennis</emphasis>
|
||
(Chemsak, 1997)
|
||
</taxonomicName>
|
||
, but differs as follows: Antennomere III distinctly pedunculate-clavate in males (
|
||
<figureCitation id="1ED7BB8FFFCB9F4AFC9ADB0BD488F915" box="[846,912,1659,1685]" captionStart="Figures 1–7" captionStartId="3.[192,279,1833,1859]" captionTargetBox="[206,1425,213,1803]" captionTargetId="figure-61@3.[206,1425,213,1803]" captionTargetPageId="3" captionText="Figures 1–7. Pentanodes spp. 1–6) Pentanodes clavatus, holotype male: 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Head, frontal view; 5) Head and pronotum, oblique view; 6) Pronotum and basal antennomeres. 7) Pentanodes tropipennis (Chemsak, 1997), male from Costa Rica, pronotum and basal antennomeres." figureDoi="http://doi.org/10.5281/zenodo.7710089" httpUri="https://zenodo.org/record/7710089/files/figure.png" pageId="8" pageNumber="7">Fig. 6</figureCitation>
|
||
); pronotum mostly striate-alveolate (
|
||
<figureCitation id="1ED7BB8FFFCB9F4AFAE6DB0BD26DF915" box="[1330,1397,1659,1685]" captionStart="Figures 1–7" captionStartId="3.[192,279,1833,1859]" captionTargetBox="[206,1425,213,1803]" captionTargetId="figure-61@3.[206,1425,213,1803]" captionTargetPageId="3" captionText="Figures 1–7. Pentanodes spp. 1–6) Pentanodes clavatus, holotype male: 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Head, frontal view; 5) Head and pronotum, oblique view; 6) Pronotum and basal antennomeres. 7) Pentanodes tropipennis (Chemsak, 1997), male from Costa Rica, pronotum and basal antennomeres." figureDoi="http://doi.org/10.5281/zenodo.7710089" httpUri="https://zenodo.org/record/7710089/files/figure.png" pageId="8" pageNumber="7">Fig. 6</figureCitation>
|
||
). In
|
||
<taxonomicName id="41ECDC89FFCB9F4AFF15DBEED64DF937" baseAuthorityName="Chemsak" baseAuthorityYear="1997" box="[193,341,1694,1719]" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="tropipennis">
|
||
<emphasis id="B4987B18FFCB9F4AFF15DBEED64DF937" box="[193,341,1694,1719]" italics="true" pageId="8" pageNumber="7">P. tropipennis</emphasis>
|
||
</taxonomicName>
|
||
, antennomere III is gradually widened from base to apex in males (
|
||
<figureCitation id="1ED7BB8FFFCB9F4AFB89DBEDD3B8F937" box="[1117,1184,1693,1719]" captionStart="Figures 1–7" captionStartId="3.[192,279,1833,1859]" captionTargetBox="[206,1425,213,1803]" captionTargetId="figure-61@3.[206,1425,213,1803]" captionTargetPageId="3" captionText="Figures 1–7. Pentanodes spp. 1–6) Pentanodes clavatus, holotype male: 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Head, frontal view; 5) Head and pronotum, oblique view; 6) Pronotum and basal antennomeres. 7) Pentanodes tropipennis (Chemsak, 1997), male from Costa Rica, pronotum and basal antennomeres." figureDoi="http://doi.org/10.5281/zenodo.7710089" httpUri="https://zenodo.org/record/7710089/files/figure.png" pageId="8" pageNumber="7">Fig. 7</figureCitation>
|
||
), and the pronotum is finely densely longitudinally striate (
|
||
<figureCitation id="1ED7BB8FFFCB9F4AFD89DBB0D5B9F95A" box="[605,673,1728,1754]" captionStart="Figures 1–7" captionStartId="3.[192,279,1833,1859]" captionTargetBox="[206,1425,213,1803]" captionTargetId="figure-61@3.[206,1425,213,1803]" captionTargetPageId="3" captionText="Figures 1–7. Pentanodes spp. 1–6) Pentanodes clavatus, holotype male: 1) Dorsal habitus; 2) Ventral habitus; 3) Lateral habitus; 4) Head, frontal view; 5) Head and pronotum, oblique view; 6) Pronotum and basal antennomeres. 7) Pentanodes tropipennis (Chemsak, 1997), male from Costa Rica, pronotum and basal antennomeres." figureDoi="http://doi.org/10.5281/zenodo.7710089" httpUri="https://zenodo.org/record/7710089/files/figure.png" pageId="8" pageNumber="7">Fig. 7</figureCitation>
|
||
). This new species differs from
|
||
<taxonomicName id="41ECDC89FFCB9F4AFBDDDBB1D696F97C" authority="Heffern, Botero and Santos-Silva, 2020" authorityName="Heffern, Botero and Santos-Silva" authorityYear="2020" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="nicaraguensis">
|
||
<emphasis id="B4987B18FFCB9F4AFBDDDBB1D3A0F95A" box="[1033,1208,1729,1754]" italics="true" pageId="8" pageNumber="7">P. nicaraguensis</emphasis>
|
||
Heffern, Botero and Santos-Silva, 2020
|
||
</taxonomicName>
|
||
(known only by the female
|
||
<typeStatus id="595719A8FFCB9F4AFD18DB92D436F97C" box="[716,814,1762,1788]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="90EA6C83FFCB9F4AFCBCDB93D4FCF97C" box="[872,996,1762,1789]" pageId="8" pageNumber="7" type="female">two female</specimenCount>
|
||
<typeStatus id="595719A8FFCB9F4AFC38DB92D344F97C" box="[1004,1116,1762,1788]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
), by the dorsal surface of the head without somewhat abundant whitish setae (present in
|
||
<taxonomicName id="41ECDC89FFCB9F4AFCB4DA75D315F89E" authorityName="Heffern, Botero and Santos-Silva" authorityYear="2020" box="[864,1037,1797,1822]" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="nicaraguensis">
|
||
<emphasis id="B4987B18FFCB9F4AFCB4DA75D315F89E" box="[864,1037,1797,1822]" italics="true" pageId="8" pageNumber="7">P. nicaraguensis</emphasis>
|
||
</taxonomicName>
|
||
), pronotum mostly striate-alveolate (sculpturing distinctly finer and denser in
|
||
<taxonomicName id="41ECDC89FFCB9F4AFD72DA58D44BF8C1" authorityName="Heffern, Botero and Santos-Silva" authorityYear="2020" box="[678,851,1832,1857]" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="nicaraguensis">
|
||
<emphasis id="B4987B18FFCB9F4AFD72DA58D44BF8C1" box="[678,851,1832,1857]" italics="true" pageId="8" pageNumber="7">P. nicaraguensis</emphasis>
|
||
</taxonomicName>
|
||
), scutellum distinctly smaller, 0.15 times as wide as the posterior width of the prothorax (wider, 0.25 times as wide as the posterior width of the prothorax in
|
||
<taxonomicName id="41ECDC89FFCB9F4AFA84DA3AD63EF806" authorityName="Heffern, Botero and Santos-Silva" authorityYear="2020" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="nicaraguensis">
|
||
<emphasis id="B4987B18FFCB9F4AFA84DA3AD63EF806" italics="true" pageId="8" pageNumber="7">P.nicaraguensis</emphasis>
|
||
</taxonomicName>
|
||
), and elytral yellowish-white band arched (straight in
|
||
<taxonomicName id="41ECDC89FFCB9F4AFC57DA1DD337F806" authorityName="Heffern, Botero and Santos-Silva" authorityYear="2020" box="[899,1071,1901,1926]" class="Insecta" family="Cerambycidae" genus="Pentanodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="nicaraguensis">
|
||
<emphasis id="B4987B18FFCB9F4AFC57DA1DD337F806" box="[899,1071,1901,1926]" italics="true" pageId="8" pageNumber="7">P. nicaraguensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |