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<mods:title id="D90298FF6ECBA1168267E4AED8ACEAD5">Revision of Epyris afer (Magretti, 1884), comb. rev. (Hymenoptera: Bethylidae) with new synonymy and description of two new species</mods:title>
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<taxonomicName id="4196D7E015543D48C491CF63D2A5F9AB" authority="Magretti, 1884" authorityName="Magretti" authorityYear="1884" box="[151,496,1551,1577]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115543D48C491CF63D248F9AB" bold="true" box="[151,285,1551,1577]" italics="true" pageId="1" pageNumber="18">Epyris afer</emphasis>
(
<bibRefCitation id="E207D19215543D48C52ACF63D2BCF9AB" author="Magretti" box="[300,489,1551,1577]" pageId="1" pageNumber="18" refString="Magretti, P. (1884) Risultati di raccolte imenotterologiche nell' Africa orientale. Annali de Museo Civico di Storia Naturale d'Genova, 21 (1), 523 - 636." type="journal article" year="1884">Magretti, 1884</bibRefCitation>
)
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, comb. rev.
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</paragraph>
<paragraph id="8629AC6315543D48C491CF5ED1F2F9C8" blockId="1.[151,679,1551,1610]" box="[151,679,1586,1610]" pageId="1" pageNumber="18">
(
<figureCitation id="1EADB0E615543D48C499CF5ED25FF9C8" box="[159,266,1586,1610]" captionStart="FIGURES 1 – 6" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[151,1436,195,1882]" captionTargetId="figure@6.[151,1436,195,1883]" captionTargetPageId="6" captionText="FIGURES 1 – 6. Head. 1. Epyris afer Ƥ, lectotype, dorsal view. 2 – 3. E. enerterus sp. nov. Ƥ, holotype. 2. Dorsal view; 3. Ventral view. 4. E. afer 3, paralectotype, dorsal view. 5 – 6. E. penatii sp. nov. 3, holotype. 5. Dorsal view; 6. Ventral view. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204625/files/figure.png" pageId="1" pageNumber="18">Figs.1, 4</figureCitation>
, 
<figureCitation id="1EADB0E615543D48C510CF5ED204F9C8" box="[278,337,1586,1610]" captionStart="FIGURES 7 – 18" captionStartId="7.[151,264,1085,1108]" captionTargetBox="[184,1407,198,1061]" captionTargetId="figure@7.[178,1409,193,1064]" captionTargetPageId="7" captionText="FIGURES 7 – 18. Mandible. 7 – 9. Epyris afer Ƥ, lectotype. 7. Frontal view; 8. Ventral view; 9. Dorsal view. 10 – 12. E. afer 3, paralectotype. 10. Frontal view; 11. Ventral view; 12. Dorsal view. 13 – 15. E. enerterus sp. nov. Ƥ, holotype. 13. Frontal view; 14. Ventral view; 15. Dorsal view. 16 – 18. E. penatii sp. nov. 3, holotype. 16. Frontal view; 17. Ventral view; 18. Dorsal view. (Scale bar = 200 µm)." httpUri="https://zenodo.org/record/204626/files/figure.png" pageId="1" pageNumber="18">7–12</figureCitation>
, 
<figureCitation id="1EADB0E615543D48C55ACF5ED2F1F9C8" box="[348,420,1586,1610]" captionStart="FIGURES 19 – 22" captionStartId="9.[151,264,1914,1937]" captionTargetBox="[151,1436,193,1892]" captionTargetId="figure@9.[151,1436,193,1893]" captionTargetPageId="9" captionText="FIGURES 19 – 22. Mesosoma in dorsal view and mesopleuron. 19 – 20. Epyris afer Ƥ, lectotype. 19. Mesosoma; 20. Mesopleuron. 21 – 22. E. enerterus sp. nov. Ƥ, holotype. 21. Mesosoma; 22. Mesopleuron. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204627/files/figure.png" pageId="1" pageNumber="18">19, 20</figureCitation>
, 
<figureCitation id="1EADB0E615543D48C5B6CF5ED2ACF9C8" box="[432,505,1586,1610]" captionStart="FIGURES 23 – 26" captionStartId="10.[151,264,1865,1888]" captionTargetBox="[151,1436,193,1844]" captionTargetId="figure@10.[151,1436,193,1844]" captionTargetPageId="10" captionText="FIGURES 23 – 26. Mesosoma in dorsal view and mesopleuron. 23 – 24. Epyris afer 3, paralectotype. 23. Mesosoma; 24. Mesopleuron. 25 – 26. E. penatii sp. nov. 3, holotype. 25. Mesosoma; 26. Mesopleuron. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204628/files/figure.png" pageId="1" pageNumber="18">23, 24</figureCitation>
, 
<figureCitation id="1EADB0E615543D48C602CF5ED119F9C8" box="[516,588,1586,1610]" captionStart="FIGURES 27 – 30" captionStartId="11.[151,264,1358,1381]" captionTargetBox="[151,1436,195,1336]" captionTargetId="figure@11.[151,1436,195,1336]" captionTargetPageId="11" captionText="FIGURES 27 – 30. Petiole in ventral view. 27. Epyris afer Ƥ, lectotype. 28. E. enerterus sp. nov. Ƥ, holotype. 29. E. afer 3, paralectotype. 30. E. penatii sp. nov. 3, holotype. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204629/files/figure.png" pageId="1" pageNumber="18">27, 29</figureCitation>
, 
<figureCitation id="1EADB0E615543D48C65ECF5ED1CAF9C8" box="[600,671,1586,1610]" captionStart="FIGURES 31 – 36" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[151,1415,208,1890]" captionTargetId="figure@12.[151,1436,193,1894]" captionTargetPageId="12" captionText="FIGURES 31 – 36. Genitalia. 31 – 33. Epyris afer 3, paralectotype. 31. Dorsal view; 32. Ventral view. 33. Lateral view. 34 – 36. E. penatii sp. nov. 3, holotype. 34. Dorsal view; 35. Ventral view. 36. Lateral view. (Scale bar = 250 µm)" httpUri="https://zenodo.org/record/204630/files/figure.png" pageId="1" pageNumber="18">31–33</figureCitation>
)
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<taxonomicName id="4196D7E015543D48C491CF17D151F912" authority="Magretti 1884: 533" authorityName="Magretti" authorityPageNumber="533" authorityYear="1884" box="[151,516,1658,1680]" class="Insecta" genus="Pristocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="B4E2707115543D48C491CF17D261F912" box="[151,308,1658,1680]" italics="true" pageId="1" pageNumber="18">Pristocera afra</emphasis>
<treatmentCitation id="07378A7215543D48C53ACF16D151F912" author="Magretti" box="[316,516,1658,1680]" page="533" pageId="1" pageNumber="18" year="1884">
<bibRefCitation id="E207D19215543D48C53ACF16D287F912" author="Magretti" box="[316,466,1658,1680]" pageId="1" pageNumber="18" refString="Magretti, P. (1884) Risultati di raccolte imenotterologiche nell' Africa orientale. Annali de Museo Civico di Storia Naturale d'Genova, 21 (1), 523 - 636." type="journal article" year="1884">Magretti 1884</bibRefCitation>
: 533
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–534, pl. 1 fig. 1, Ƥ 3, 
<typeStatus id="592D12C115543D48C6F9CF16D009F912" box="[767,860,1658,1680]" pageId="1" pageNumber="18" type="lectotype">lectotype</typeStatus>
(designated here), Ƥ, examined; 
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<bibRefCitation id="E207D19215543D48C0B0CF16D63FF912" author="Dalla" box="[1206,1386,1658,1680]" pageId="1" pageNumber="18" refString="Dalla Torre de, C. G. (1898) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Volumen V: Chalcididae et Proctotrupidae. Sumptibus Guilelmi Engelmann, Lipsiae, i - x + 1 - 598." type="book chapter" year="1898">Dalla Torre 1898</bibRefCitation>
: 561
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(catalog); 
<treatmentCitation id="07378A7215543D48C528CFF5D11CF92D" author="Gordh" box="[302,585,1689,1711]" page="133" pageId="1" pageNumber="18" year="1990">
<bibRefCitation id="E207D19215543D48C528CFF5D14CF92D" author="Gordh" box="[302,537,1689,1711]" pageId="1" pageNumber="18" refString="Gordh, G. &amp; Moczar, L. (1990) A catalog of the world Bethylidae (Hymenoptera: Aculeata). Memories of the American Entomological Institute, 46, 1 - 323." type="journal article" year="1990">Gordh &amp; Móczár 1990</bibRefCitation>
: 133
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(catalog).
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</paragraph>
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<treatmentCitationGroup id="A6868B4D15543D48C491CFD5D76CF94C" box="[151,1081,1720,1742]" pageId="1" pageNumber="18">
<taxonomicName id="4196D7E015543D48C491CFD5D28CF94F" authority="Magretti 1897: 319" authorityName="Magretti" authorityPageNumber="319" authorityYear="1897" box="[151,473,1720,1742]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115543D48C491CFD5D25DF94C" box="[151,264,1720,1742]" italics="true" pageId="1" pageNumber="18">Epyris afer</emphasis>
: 
<treatmentCitation id="07378A7215543D48C513CFD4D28CF94F" author="Magretti" box="[277,473,1720,1742]" page="319" pageId="1" pageNumber="18" year="1897">
<bibRefCitation id="E207D19215543D48C513CFD4D2FCF94F" author="Magretti" box="[277,425,1720,1742]" pageId="1" pageNumber="18" refString="Magretti, P. (1897) Viaggio di Leonardo Fea in Birmania e regioni Vicine 74. Imenotteri. Parte Seconda. Trigonalidi, Betilidi, Crisididi colla descrizione de parecchie nuove specie. Annali de Museo Civico di Storia Naturale d'Genova. Series 2, 17, 308 - 326." type="journal article" year="1897">Magretti 1897</bibRefCitation>
: 319
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</taxonomicName>
(part.); 
<treatmentCitation id="07378A7215543D48C62DCFD4D1B5F94C" author="Kieffer" box="[555,736,1720,1742]" page="396" pageId="1" pageNumber="18" year="1904">
<bibRefCitation id="E207D19215543D48C62DCFD4D1E5F94F" author="Kieffer" box="[555,688,1720,1742]" pageId="1" pageNumber="18" refString="Kieffer, J. J. (1904) Description de nouveaux Dryininae et Bethylinae du Musee Civique de Genes. Annali del Museo civico di Storia Naturale di Genova, 41, 351 - 412." type="journal article" year="1904">Kieffer 1904</bibRefCitation>
: 396
</treatmentCitation>
(key); 
<treatmentCitation id="07378A7215543D48C72FCFD4D084F94F" author="Kieffer" box="[809,977,1720,1742]" page="27" pageId="1" pageNumber="18" year="1908">
<bibRefCitation id="E207D19215543D48C72FCFD4D0FBF94F" author="Kieffer" box="[809,942,1720,1742]" pageId="1" pageNumber="18" refString="Kieffer, J. J. (1908) Hymenoptera. Fam. Bethylidae. In: Wytsman, P., Genera Insectorum. Fasc. 76. V. Verteneuil &amp; L. Desmet Imprimeurs-Editeurs, Bruxelles, 50 pp. + 3 pls." type="book" year="1908">Kieffer 1908</bibRefCitation>
: 27
</treatmentCitation>
(catalog).
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</paragraph>
<paragraph id="8629AC6315543D48C491CFB4D660F96E" blockId="1.[151,1437,1658,2018]" box="[151,1333,1750,1773]" pageId="1" pageNumber="18">
<treatmentCitationGroup id="A6868B4D15543D48C491CFB4D660F96E" box="[151,1333,1750,1773]" pageId="1" pageNumber="18">
<emphasis id="B4E2707115543D48C491CFB4D271F96F" box="[151,292,1751,1773]" italics="true" pageId="1" pageNumber="18">
Lytepyris 
<taxonomicName id="4196D7E015543D48C4FDCFBBD271F96F" authority="Kieffer 1913: 108" authorityName="Kieffer" authorityPageNumber="108" authorityYear="1913" box="[251,292,1751,1773]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="afer">afer</taxonomicName>
</emphasis>
: 
<treatmentCitation id="07378A7215543D48C537CFBAD2BCF96E" author="Kieffer" box="[305,489,1750,1772]" page="108" pageId="1" pageNumber="18" year="1913">
<bibRefCitation id="E207D19215543D48C537CFBAD2E3F96E" author="Kieffer" box="[305,438,1750,1772]" pageId="1" pageNumber="18" refString="Kieffer, J. J. (1913) Nouveau microhymenoperes de l'Afrique equatoriale. Laboratorio di Zoologia Generate e Agraria delia R. Scuola Superiore d'Agricoltura in Portici. Bolletino, 7, 105 - 108." type="journal article" year="1913">Kieffer 1913</bibRefCitation>
: 108
</treatmentCitation>
; 
<treatmentCitation id="07378A7215543D48C5F5CFBAD1FDF96E" author="Kieffer" box="[499,680,1750,1772]" page="416" pageId="1" pageNumber="18" year="1914">
<bibRefCitation id="E207D19215543D48C5F5CFBAD12DF96E" author="Kieffer" box="[499,632,1750,1772]" pageId="1" pageNumber="18" refString="Kieffer, J. J. (1914) Bethylinae. Das Tierreich, 41, 228 - 595." type="journal article" year="1914">Kieffer 1914</bibRefCitation>
: 416
</treatmentCitation>
–417 (key, redescription); 
<treatmentCitation id="07378A7215543D48C7B4CFBAD798F96E" author="Gordh" box="[946,1229,1750,1772]" page="133" pageId="1" pageNumber="18" year="1990">
<bibRefCitation id="E207D19215543D48C7B4CFBAD7C8F96E" author="Gordh" box="[946,1181,1750,1772]" pageId="1" pageNumber="18" refString="Gordh, G. &amp; Moczar, L. (1990) A catalog of the world Bethylidae (Hymenoptera: Aculeata). Memories of the American Entomological Institute, 46, 1 - 323." type="journal article" year="1990">Gordh &amp; Móczár 1990</bibRefCitation>
: 133
</treatmentCitation>
(catalog).
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</paragraph>
<paragraph id="8629AC6315543D48C491CF9AD171F889" blockId="1.[151,1437,1658,2018]" box="[151,548,1781,1803]" pageId="1" pageNumber="18">
<treatmentCitationGroup id="A6868B4D15543D48C491CF9AD171F889" box="[151,548,1781,1803]" pageId="1" pageNumber="18">
<emphasis id="B4E2707115543D48C491CF9AD272F889" box="[151,295,1781,1803]" italics="true" pageId="1" pageNumber="18">
Disepyris 
<taxonomicName id="4196D7E015543D48C4F8CF99D272F889" authority="Terayama 2004: 97" authorityName="Terayama" authorityPageNumber="97" authorityYear="2004" box="[254,295,1781,1803]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="afer">afer</taxonomicName>
</emphasis>
: 
<treatmentCitation id="07378A7215543D48C532CF9AD2ADF889" author="Terayama" box="[308,504,1782,1803]" page="97" pageId="1" pageNumber="18" year="2004">
<bibRefCitation id="E207D19215543D48C532CF9AD281F889" author="Terayama" box="[308,468,1782,1803]" pageId="1" pageNumber="18" refString="Terayama, M. (2004) Formosiepyris a new genus of the family Bethylidae (Hymenoptera, Bethylidae) from the Oriental region with a proposal of a new synonymy of genus. Liberal Arts, 12, 91 - 99." type="journal article" year="2004">Terayama 2004</bibRefCitation>
: 97
</treatmentCitation>
–98.
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</paragraph>
<paragraph id="8629AC6315543D48C491CE79D11BF8E5" blockId="1.[151,1437,1658,2018]" pageId="1" pageNumber="18">
<treatmentCitationGroup id="A6868B4D15543D48C491CE79D11BF8E5" pageId="1" pageNumber="18">
<taxonomicName id="4196D7E015543D48C491CE79D156F8AB" authority="Kieffer 1904: 396" authorityName="Kieffer" authorityPageNumber="396" authorityYear="1904" box="[151,515,1811,1834]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B4E2707115543D48C491CE79D268F8A8" box="[151,317,1812,1834]" italics="true" pageId="1" pageNumber="18">Epyris pilosipes</emphasis>
<treatmentCitation id="07378A7215543D48C540CE7FD156F8AB" author="Kieffer" box="[326,515,1811,1833]" page="396" pageId="1" pageNumber="18" year="1904">
<bibRefCitation id="E207D19215543D48C540CE7FD29BF8AB" author="Kieffer" box="[326,462,1811,1833]" pageId="1" pageNumber="18" refString="Kieffer, J. J. (1904) Description de nouveaux Dryininae et Bethylinae du Musee Civique de Genes. Annali del Museo civico di Storia Naturale di Genova, 41, 351 - 412." type="journal article" year="1904">Kieffer 1904</bibRefCitation>
: 396
</treatmentCitation>
</taxonomicName>
, 404–405 (key, description), Ƥ (
<typeStatus id="592D12C115543D48C75DCE78D0FEF8AB" box="[859,939,1812,1833]" pageId="1" pageNumber="18" type="syntype">syntype</typeStatus>
examined); 
<treatmentCitation id="07378A7215543D48C036CE7FD78AF8AB" author="Kieffer" box="[1072,1247,1811,1833]" page="28" pageId="1" pageNumber="18" year="1908">
<bibRefCitation id="E207D19215543D48C036CE7FD7ECF8AB" author="Kieffer" box="[1072,1209,1811,1833]" pageId="1" pageNumber="18" refString="Kieffer, J. J. (1908) Hymenoptera. Fam. Bethylidae. In: Wytsman, P., Genera Insectorum. Fasc. 76. V. Verteneuil &amp; L. Desmet Imprimeurs-Editeurs, Bruxelles, 50 pp. + 3 pls." type="book" year="1908">Kieffer 1908</bibRefCitation>
: 28
</treatmentCitation>
(catalog); 
<treatmentCitation id="07378A7215543D48C155CE7FD27AF8CA" author="Kieffer" page="310" pageId="1" pageNumber="18" year="1914">
<bibRefCitation id="E207D19215543D48C155CE7FD3A8F8CA" author="Kieffer" pageId="1" pageNumber="18" refString="Kieffer, J. J. (1914) Bethylinae. Das Tierreich, 41, 228 - 595." type="journal article" year="1914">Kieffer 1914</bibRefCitation>
: 310
</treatmentCitation>
, 330, fig. 140 (key, redescription); 
<treatmentCitation id="07378A7215543D48C697CE5ED011F8CA" author="Kurian" box="[657,836,1842,1864]" page="269" pageId="1" pageNumber="18" year="1954">
<bibRefCitation id="E207D19215543D48C697CE5ED041F8CA" author="Kurian" box="[657,788,1842,1864]" pageId="1" pageNumber="18" refString="Kurian, C. (1954) Catalogue of Oriental Bethyloidea. Agra University Journal of Research, 3 (1), 253 - 288." type="journal article" year="1954">Kurian 1954</bibRefCitation>
: 269
</treatmentCitation>
(catalog); 
<treatmentCitation id="07378A7215543D48C7B7CE5ED72DF8CA" author="Richards" box="[945,1144,1842,1864]" page="357" pageId="1" pageNumber="18" year="1955">
<bibRefCitation id="E207D19215543D48C7B7CE5ED712F8CA" author="Richards" box="[945,1095,1842,1864]" pageId="1" pageNumber="18" refString="Richards, O. W. (1955) On the Bethyloidea (Hymen.) of Israel. Bulletin of the Research Council of Israel, 4, 357 - 359." type="journal article" year="1955">Richards 1955</bibRefCitation>
: 357
</treatmentCitation>
, 358 (distribution); 
<treatmentCitation id="07378A7215543D48C147CE5ED2D5F8E5" author="Gordh" page="101" pageId="1" pageNumber="18" year="1990">
<bibRefCitation id="E207D19215543D48C147CE5ED204F8E5" author="Gordh" pageId="1" pageNumber="18" refString="Gordh, G. &amp; Moczar, L. (1990) A catalog of the world Bethylidae (Hymenoptera: Aculeata). Memories of the American Entomological Institute, 46, 1 - 323." type="journal article" year="1990">Gordh &amp; Móczár 1990</bibRefCitation>
: 101
</treatmentCitation>
(catalog). 
<emphasis id="B4E2707115543D48C5EBCE3DD11BF8E5" bold="true" box="[493,590,1873,1896]" pageId="1" pageNumber="18">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8629AC6315543D48C491CE1DD7A9F807" blockId="1.[151,1437,1658,2018]" box="[151,1276,1903,1926]" pageId="1" pageNumber="18">
<treatmentCitationGroup id="A6868B4D15543D48C491CE1DD7A9F807" box="[151,1276,1903,1926]" pageId="1" pageNumber="18">
<taxonomicName id="4196D7E015543D48C491CE1DD282F807" authority="Kieffer 1905: 111" authorityName="Kieffer" authorityPageNumber="111" authorityYear="1905" box="[151,471,1904,1926]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B4E2707115543D48C491CE1DD248F804" box="[151,285,1904,1926]" italics="true" pageId="1" pageNumber="18">Epyris analis</emphasis>
Kieffer 1905: 111
</taxonomicName>
-112, Ƥ (
<typeStatus id="592D12C115543D48C637CE1CD1DEF804" box="[561,651,1904,1926]" pageId="1" pageNumber="18" type="holotype">holotype</typeStatus>
examined), nom. praeocc., non 
<treatmentCitation id="07378A7215543D48C7D6CE1CD7C4F807" author="Cresson" box="[976,1169,1904,1925]" page="193" pageId="1" pageNumber="18" year="1872">
<bibRefCitation id="E207D19215543D48C7D6CE1CD70BF807" author="Cresson" box="[976,1118,1904,1925]" pageId="1" pageNumber="18" refString="Cresson, E. T. (1872) Hymenoptera Texana. Transactions of the American Entomological Society, 4, 153 - 292." type="journal article" year="1872">Cresson 1872</bibRefCitation>
: 193
</treatmentCitation>
. 
<emphasis id="B4E2707115543D48C09DCE03D7A9F807" bold="true" box="[1179,1276,1903,1926]" pageId="1" pageNumber="18">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8629AC6315543D48C491CEFCD7C3F841" blockId="1.[151,1437,1658,2018]" pageId="1" pageNumber="18">
<treatmentCitationGroup id="A6868B4D15543D48C491CEFCD7C3F841" pageId="1" pageNumber="18">
<taxonomicName id="4196D7E015543D48C491CEFCD2A5F826" authority="Brues 1906: 143" authorityName="Brues" authorityPageNumber="143" authorityYear="1906" box="[151,496,1934,1957]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="secundus">
<emphasis id="B4E2707115543D48C491CEFCD26BF827" box="[151,318,1935,1957]" italics="true" pageId="1" pageNumber="18">Epyris secundus</emphasis>
<treatmentCitation id="07378A7215543D48C543CEE3D2A5F826" author="Brues" box="[325,496,1934,1956]" page="143" pageId="1" pageNumber="18" year="1906">
<bibRefCitation id="E207D19215543D48C543CEE3D2EAF826" author="Brues" box="[325,447,1934,1956]" pageId="1" pageNumber="18" refString="Brues, C. (1906) Notes and descriptions of North American parasitic Hymenoptera. II. Bulletin of Wisconsin Natural History Society, 4 (4), 143 - 152." type="journal article" year="1906">Brues 1906</bibRefCitation>
: 143
</treatmentCitation>
</taxonomicName>
(new name for 
<taxonomicName id="4196D7E015543D48C691CEFCD0E7F827" authority="Kieffer 1905: 111" authorityName="Kieffer" authorityPageNumber="111" authorityYear="1905" box="[663,946,1934,1957]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B4E2707115543D48C691CEFCD1A6F827" box="[663,755,1935,1957]" italics="true" pageId="1" pageNumber="18">E. analis</emphasis>
Kieffer 1905: 111
</taxonomicName>
-112); 
<treatmentCitation id="07378A7215543D48C7F3CEE2D7F4F826" author="Kieffer" box="[1013,1185,1934,1956]" page="27" pageId="1" pageNumber="18" year="1908">
<bibRefCitation id="E207D19215543D48C7F3CEE2D728F826" author="Kieffer" box="[1013,1149,1934,1956]" pageId="1" pageNumber="18" refString="Kieffer, J. J. (1908) Hymenoptera. Fam. Bethylidae. In: Wytsman, P., Genera Insectorum. Fasc. 76. V. Verteneuil &amp; L. Desmet Imprimeurs-Editeurs, Bruxelles, 50 pp. + 3 pls." type="book" year="1908">Kieffer 1908</bibRefCitation>
: 27
</treatmentCitation>
(catalog); 
<treatmentCitation id="07378A7215543D48C114CEE2D3A5F841" author="Kieffer" page="312" pageId="1" pageNumber="18" year="1914">
<bibRefCitation id="E207D19215543D48C114CEE2D6CCF826" author="Kieffer" box="[1298,1433,1934,1956]" pageId="1" pageNumber="18" refString="Kieffer, J. J. (1914) Bethylinae. Das Tierreich, 41, 228 - 595." type="journal article" year="1914">Kieffer 1914</bibRefCitation>
: 312
</treatmentCitation>
, 326-327 (key, redescription); 
<treatmentCitation id="07378A7215543D48C620CEC1D18CF841" author="Kurian" box="[550,729,1965,1987]" page="270" pageId="1" pageNumber="18" year="1954">
<bibRefCitation id="E207D19215543D48C620CEC1D1FCF841" author="Kurian" box="[550,681,1965,1987]" pageId="1" pageNumber="18" refString="Kurian, C. (1954) Catalogue of Oriental Bethyloidea. Agra University Journal of Research, 3 (1), 253 - 288." type="journal article" year="1954">Kurian 1954</bibRefCitation>
: 270
</treatmentCitation>
(catalog); 
<treatmentCitation id="07378A7215543D48C740CEC1D0B9F841" author="Kurian" box="[838,1004,1965,1987]" page="90" pageId="1" pageNumber="18" year="1955">
<bibRefCitation id="E207D19215543D48C740CEC1D09CF841" author="Kurian" box="[838,969,1965,1987]" pageId="1" pageNumber="18" refString="Kurian, C. (1955) Bethyloidea (Hymenoptera) from Indian. Agra University Journal of Research, 4, 67 - 155." type="journal article" year="1955">Kurian 1955</bibRefCitation>
: 90
</treatmentCitation>
(key). 
<emphasis id="B4E2707115543D48C032CEC1D7C3F841" bold="true" box="[1076,1174,1965,1988]" pageId="1" pageNumber="18">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8629AC6315543D48C491CEA1D7ABF863" blockId="1.[151,1437,1658,2018]" box="[151,1278,1995,2018]" pageId="1" pageNumber="18">
<treatmentCitationGroup id="A6868B4D15543D48C491CEA1D7ABF863" box="[151,1278,1995,2018]" pageId="1" pageNumber="18">
<taxonomicName id="4196D7E015543D48C491CEA1D146F863" authority="Cameron 1906: 288" authorityName="Cameron" authorityPageNumber="288" authorityYear="1906" box="[151,531,1995,2018]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B4E2707115543D48C491CEA1D215F860" box="[151,320,1996,2018]" italics="true" pageId="1" pageNumber="18">Epyris rugicollis</emphasis>
<treatmentCitation id="07378A7215543D48C541CEA0D146F863" author="Cameron" box="[327,531,1995,2017]" page="288" pageId="1" pageNumber="18" year="1906">
<bibRefCitation id="E207D19215543D48C541CEA0D2B5F863" author="Cameron" box="[327,480,1995,2017]" pageId="1" pageNumber="18" refString="Cameron, P. (1906) On the Tenthredinidae &amp; parasitic Hymenoptera collected in Baluchistan by Major C. G. Nurse. Journal of Bombay Natural History Society, 17 (2), 89 - 288." type="journal article" year="1906">
<collectingCountry id="FE81ECF315543D48C541CEA0D2F6F863" box="[327,419,1996,2017]" name="Cameroon" pageId="1" pageNumber="18">Cameron</collectingCountry>
1906
</bibRefCitation>
: 288
</treatmentCitation>
</taxonomicName>
, Ƥ (
<typeStatus id="592D12C115543D48C63ACEA7D1C3F863" box="[572,662,1995,2017]" pageId="1" pageNumber="18" type="holotype">holotype</typeStatus>
examined); 
<treatmentCitation id="07378A7215543D48C713CEA7D765F863" author="Gordh" box="[789,1072,1995,2017]" page="127" pageId="1" pageNumber="18" year="1990">
<bibRefCitation id="E207D19215543D48C713CEA7D755F863" author="Gordh" box="[789,1024,1995,2017]" pageId="1" pageNumber="18" refString="Gordh, G. &amp; Moczar, L. (1990) A catalog of the world Bethylidae (Hymenoptera: Aculeata). Memories of the American Entomological Institute, 46, 1 - 323." type="journal article" year="1990">Gordh &amp; Móczár 1990</bibRefCitation>
: 127
</treatmentCitation>
(catalog). 
<emphasis id="B4E2707115543D48C09ACEA7D7ABF863" bold="true" box="[1180,1278,1995,2018]" pageId="1" pageNumber="18">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8629AC6315573D4BC491C9F4D05BFF4E" blockId="2.[151,1397,152,266]" pageId="2" pageNumber="19">
<treatmentCitationGroup id="A6868B4D15573D4BC491C9F4D05BFF4E" pageId="2" pageNumber="19">
<taxonomicName id="4196D7E015573D4BC491C9F4D17BFF2C" authority="Kieffer 1908: 26" authorityName="Kieffer" authorityPageNumber="26" authorityYear="1908" box="[151,558,152,174]" class="Insecta" family="Bethylidae" genus="Isobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="rugicolle">
<emphasis id="B4E2707115573D4BC491C9F4D22DFF2C" box="[151,376,152,174]" italics="true" pageId="2" pageNumber="19">Isobrachium rugicolle</emphasis>
: 
<treatmentCitation id="07378A7215573D4BC580C9F4D17BFF2C" author="Kieffer" box="[390,558,152,174]" page="26" pageId="2" pageNumber="19" year="1908">
<bibRefCitation id="E207D19215573D4BC580C9F4D15EFF2F" author="Kieffer" box="[390,523,152,174]" pageId="2" pageNumber="19" refString="Kieffer, J. J. (1908) Hymenoptera. Fam. Bethylidae. In: Wytsman, P., Genera Insectorum. Fasc. 76. V. Verteneuil &amp; L. Desmet Imprimeurs-Editeurs, Bruxelles, 50 pp. + 3 pls." type="book" year="1908">Kieffer 1908</bibRefCitation>
: 26
</treatmentCitation>
</taxonomicName>
(catalog); 
<treatmentCitation id="07378A7215573D4BC69AC9F4D006FF2C" author="Kieffer" box="[668,851,152,174]" page="368" pageId="2" pageNumber="19" year="1914">
<bibRefCitation id="E207D19215573D4BC69AC9F4D075FF2F" author="Kieffer" box="[668,800,152,174]" pageId="2" pageNumber="19" refString="Kieffer, J. J. (1914) Bethylinae. Das Tierreich, 41, 228 - 595." type="journal article" year="1914">Kieffer 1914</bibRefCitation>
: 368
</treatmentCitation>
, 369 (key, redescription); 
<treatmentCitation id="07378A7215573D4BC05AC9F4D65BFF2F" author="Kurian" box="[1116,1294,152,174]" page="273" pageId="2" pageNumber="19" year="1954">
<bibRefCitation id="E207D19215573D4BC05AC9F4D78BFF2C" author="Kurian" box="[1116,1246,152,174]" pageId="2" pageNumber="19" refString="Kurian, C. (1954) Catalogue of Oriental Bethyloidea. Agra University Journal of Research, 3 (1), 253 - 288." type="journal article" year="1954">Kurian 1954</bibRefCitation>
: 273
</treatmentCitation>
(catalog). 
<taxonomicName id="4196D7E015573D4BC491C9DBD1F3FF4E" authority="Gordh &amp; Moczar 1990: 127" authorityName="Gordh &amp; Moczar" authorityPageNumber="127" authorityYear="1990" box="[151,678,182,205]" class="Insecta" family="Bethylidae" genus="Isobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B4E2707115573D4BC491C9DBD22BFF4F" box="[151,382,183,205]" italics="true" pageId="2" pageNumber="19">Isobrachium rugicollis</emphasis>
: 
<treatmentCitation id="07378A7215573D4BC58DC9DAD1F3FF4E" author="Gordh" box="[395,678,182,204]" page="127" pageId="2" pageNumber="19" year="1990">
<bibRefCitation id="E207D19215573D4BC58DC9DAD123FF4E" author="Gordh" box="[395,630,182,204]" pageId="2" pageNumber="19" refString="Gordh, G. &amp; Moczar, L. (1990) A catalog of the world Bethylidae (Hymenoptera: Aculeata). Memories of the American Entomological Institute, 46, 1 - 323." type="journal article" year="1990">Gordh &amp; Móczár 1990</bibRefCitation>
: 127
</treatmentCitation>
</taxonomicName>
(catalog).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8629AC6315573D4BC491C9BAD7C6FE8B" blockId="2.[151,1397,152,266]" pageId="2" pageNumber="19">
<treatmentCitationGroup id="A6868B4D15573D4BC491C9BAD7C6FE8B" pageId="2" pageNumber="19">
<taxonomicName id="4196D7E015573D4BC491C9BAD15BFF68" authority="Turner 1928: 130" authorityName="Turner" authorityPageNumber="130" authorityYear="1928" box="[151,526,213,235]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="plurilineata">
<emphasis id="B4E2707115573D4BC491C9BAD202FF69" box="[151,343,213,235]" italics="true" pageId="2" pageNumber="19">Epyris plurilineata</emphasis>
<treatmentCitation id="07378A7215573D4BC55BC9BAD15BFF68" author="Turner" box="[349,526,213,235]" page="130" pageId="2" pageNumber="19" year="1928">
<bibRefCitation id="E207D19215573D4BC55BC9BAD288FF68" author="Turner" box="[349,477,213,235]" pageId="2" pageNumber="19" refString="Turner, R. E. (1928) New Hymenoptera of the family Bethylidae. The Annals and Magazine of Natural History, 1 (10), 129 - 152." type="journal article" year="1928">Turner 1928</bibRefCitation>
: 130
</treatmentCitation>
</taxonomicName>
–131, Ƥ (
<typeStatus id="592D12C115573D4BC66BC9B9D192FF69" box="[621,711,213,235]" pageId="2" pageNumber="19" type="holotype">holotype</typeStatus>
examined); 
<treatmentCitation id="07378A7215573D4BC743C9B9D735FF68" author="Gordh" box="[837,1120,213,235]" page="102" pageId="2" pageNumber="19" year="1990">
<bibRefCitation id="E207D19215573D4BC743C9B9D765FF68" author="Gordh" box="[837,1072,213,235]" pageId="2" pageNumber="19" refString="Gordh, G. &amp; Moczar, L. (1990) A catalog of the world Bethylidae (Hymenoptera: Aculeata). Memories of the American Entomological Institute, 46, 1 - 323." type="journal article" year="1990">Gordh &amp; Móczár 1990</bibRefCitation>
: 102
</treatmentCitation>
(catalog). 
<emphasis id="B4E2707115573D4BC0CBC9B8D67BFF68" bold="true" box="[1229,1326,212,235]" pageId="2" pageNumber="19">Syn. nov.</emphasis>
<taxonomicName id="4196D7E015573D4BC491C999D144FE8B" authority="Benoit 1957: 14" authorityName="Benoit" authorityPageNumber="14" authorityYear="1957" box="[151,529,243,266]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="plurilineatus">
<emphasis id="B4E2707115573D4BC491C999D234FE88" box="[151,353,244,266]" italics="true" pageId="2" pageNumber="19">Epyris plurilineatus</emphasis>
: 
<treatmentCitation id="07378A7215573D4BC568C99FD144FE8B" author="Benoit" box="[366,529,243,265]" page="14" pageId="2" pageNumber="19" year="1957">
<bibRefCitation id="E207D19215573D4BC568C99FD2BBFE8B" author="Benoit" box="[366,494,243,265]" pageId="2" pageNumber="19" refString="Benoit, P. L. G. (1957) Hymenoptera - Bethylidae. Exploration du Parc National Albert. Mission G. F. De Witte, 1933 - 1935. Bruxelles Institut des Parcs Nationaux de Congo Belge, 88, 1 - 57." type="journal article" year="1957">Benoit 1957</bibRefCitation>
: 14
</treatmentCitation>
</taxonomicName>
(distribution, variation); 
<treatmentCitation id="07378A7215573D4BC717C99FD779FE8B" author="Gordh" box="[785,1068,243,265]" page="102" pageId="2" pageNumber="19" year="1990">
<bibRefCitation id="E207D19215573D4BC717C99FD0A9FE8B" author="Gordh" box="[785,1020,243,265]" pageId="2" pageNumber="19" refString="Gordh, G. &amp; Moczar, L. (1990) A catalog of the world Bethylidae (Hymenoptera: Aculeata). Memories of the American Entomological Institute, 46, 1 - 323." type="journal article" year="1990">Gordh &amp; Móczár 1990</bibRefCitation>
: 102
</treatmentCitation>
(catalog).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CE8CFFE815573D4AC491C85BD02CFE2E" lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19" type="materials_examined">
<paragraph id="8629AC6315573D4BC491C85BD7B4FD34" blockId="2.[151,1437,310,2026]" pageId="2" pageNumber="19">
<emphasis id="B4E2707115573D4BC491C85BD29BFECD" bold="true" box="[151,462,310,335]" pageId="2" pageNumber="19">
<typeStatus id="592D12C115573D4BC491C85BD387FECD" box="[151,210,311,335]" pageId="2" pageNumber="19">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="592D12C115573D4BC5DAC85AD129FECC" box="[476,636,310,334]" pageId="2" pageNumber="19" type="lectotype">LECTOTYPE</typeStatus>
of 
<taxonomicName id="4196D7E015573D4BC6B7C85BD031FECC" box="[689,868,310,334]" class="Insecta" genus="Pristocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="B4E2707115573D4BC6B7C85BD031FECC" box="[689,868,310,334]" italics="true" pageId="2" pageNumber="19">Pristocera afra</emphasis>
</taxonomicName>
(designated here): Ƥ, [
<collectingCountry id="FE81ECF315573D4BC085C85AD658FECC" box="[1155,1293,310,334]" name="Ethiopia" pageId="2" pageNumber="19">ETHIOPIA</collectingCountry>
], 
<emphasis id="B4E2707115573D4BC120C854D6C9FECD" box="[1318,1436,312,335]" italics="true" pageId="2" pageNumber="19">Metemma,</emphasis>
[13.026817, 36.161562], 
<date id="F2288AA315573D4BC5B4C836D167FEF0" box="[434,562,346,370]" pageId="2" pageNumber="19" value="1883-03-24">24.III.1883</date>
, P. Magretti col. (
<collectionCode id="E08734A615573D4BC6F0C836D019FEF0" box="[758,844,346,370]" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale Giacomo Doria" pageId="2" pageNumber="19">MCSN</collectionCode>
). 
<typeStatus id="592D12C115573D4BC75BC836D70DFEF0" box="[861,1112,346,370]" pageId="2" pageNumber="19" type="paralectotype">PARALECTOTYPES</typeStatus>
of 
<taxonomicName id="4196D7E015573D4BC07AC830D672FEF1" box="[1148,1319,347,371]" class="Insecta" genus="Pristocera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="afra">
<emphasis id="B4E2707115573D4BC07AC830D672FEF1" box="[1148,1319,347,371]" italics="true" pageId="2" pageNumber="19">Pristocera afra</emphasis>
</taxonomicName>
: [ETHIO- PIA], 2 Ƥ, 
<emphasis id="B4E2707115573D4BC512C8ECD2DFFE15" box="[276,394,384,407]" italics="true" pageId="2" pageNumber="19">Metemma,</emphasis>
[13.026817, 36.161562], 
<date id="F2288AA315573D4BC6A8C812D07BFE14" box="[686,814,382,406]" pageId="2" pageNumber="19" value="1883-03-24">24.III.1883</date>
, P. Magretti col. (
<collectionCode id="E08734A615573D4BC7F0C812D719FE14" box="[1014,1100,382,406]" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale Giacomo Doria" pageId="2" pageNumber="19">MCSN</collectionCode>
); 1 3, 
<date id="F2288AA315573D4BC09EC812D643FE14" box="[1176,1302,382,406]" pageId="2" pageNumber="19" value="1883-03-22">22.III.1883</date>
(
<collectionCode id="E08734A615573D4BC123C812D62EFE14" box="[1317,1403,382,406]" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale Giacomo Doria" pageId="2" pageNumber="19">MCSN</collectionCode>
). 1 Ƥ, 
<emphasis id="B4E2707115573D4BC4BCC8CFD23AFE39" box="[186,367,419,443]" italics="true" pageId="2" pageNumber="19">Bahr el Salaam,</emphasis>
rive {= river banks}, [13.85, 36.1333333], 
<date id="F2288AA315573D4BC75EC8CED08CFE38" box="[856,985,418,442]" pageId="2" pageNumber="19" value="1883-03-14">14.III.1883</date>
, P. Magretti col (
<collectionCode id="E08734A615573D4BC091C8CED7B8FE38" box="[1175,1261,418,442]" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale Giacomo Doria" pageId="2" pageNumber="19">MCSN</collectionCode>
). 
<typeStatus id="592D12C115573D4BC0F9C8CED628FE38" box="[1279,1405,418,442]" pageId="2" pageNumber="19" type="syntype">SYNTYPE</typeStatus>
of 
<taxonomicName id="4196D7E015573D4BC491C8ABD204FE5C" box="[151,337,454,478]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B4E2707115573D4BC491C8ABD204FE5C" box="[151,337,454,478]" italics="true" pageId="2" pageNumber="19">Epyris pilosipes</emphasis>
</taxonomicName>
: Ƥ, GHINEA PORTOGHESE {
<collectingCountry id="FE81ECF315573D4BC6D4C8AAD0ECFE5C" box="[722,953,454,478]" name="Guinea-Bissau" pageId="2" pageNumber="19">GUINEA-BISSAU</collectingCountry>
}, 
<emphasis id="B4E2707115573D4BC7D4C8AAD77CFE5C" box="[978,1065,454,478]" italics="true" pageId="2" pageNumber="19">Bolama</emphasis>
, [11.578581, -15.483679], 
<date id="F2288AA315573D4BC176C8ABD3AAFD80" pageId="2" pageNumber="19" value="1899-06" valueMax="1899-12-31" valueMin="1899-06-00">VI– XII.1899</date>
, L. Fea col. (
<collectionCode id="E08734A615573D4BC589C886D2B0FD80" box="[399,485,490,514]" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale Giacomo Doria" pageId="2" pageNumber="19">MCSN</collectionCode>
); 
<typeStatus id="592D12C115573D4BC5F1C886D1DEFD80" box="[503,651,490,514]" pageId="2" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
of 
<taxonomicName id="4196D7E015573D4BC6A9C880D0D6FD81" box="[687,899,491,515]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="plurilineata">
<emphasis id="B4E2707115573D4BC6A9C880D0D6FD81" box="[687,899,491,515]" italics="true" pageId="2" pageNumber="19">Epyris plurilineata</emphasis>
</taxonomicName>
: Ƥ, S[OUTHERN] 
<collectingCountry id="FE81ECF315573D4BC064C886D78EFD80" box="[1122,1243,490,514]" name="Zimbabwe" pageId="2" pageNumber="19">RODESIA</collectingCountry>
{
<collectingCountry id="FE81ECF315573D4BC0EFC886D6DAFD80" box="[1257,1423,490,514]" name="Zimbabwe" pageId="2" pageNumber="19">ZIMBABWE</collectingCountry>
}, 
<emphasis id="B4E2707115573D4BC491CB62D27AFDA4" box="[151,303,526,550]" italics="true" pageId="2" pageNumber="19">Thabasilitche</emphasis>
, {coordinates not found, coordinates of Bulawayo, another occurrence of the species = -20.17, 28.58}, 
<date id="F2288AA315573D4BC4FECB5ED223FDC8" box="[248,374,562,586]" pageId="2" pageNumber="19" value="1924-07-06">6.VII.1924</date>
, R.H.R. Stevenson col. Brit[ish] Mus[eum] 1927-344, B.M. 
<typeStatus id="592D12C115573D4BC047CB5FD72AFDC9" box="[1089,1151,563,587]" pageId="2" pageNumber="19">Type</typeStatus>
Hym. 13302 (
<collectionCode id="E08734A615573D4BC130CB5ED6C5FDC8" box="[1334,1424,562,586]" pageId="2" pageNumber="19">BMNH</collectionCode>
); 
<typeStatus id="592D12C115573D4BC491CB3AD27EFDEC" box="[151,299,598,622]" pageId="2" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
of 
<taxonomicName id="4196D7E015573D4BC557CB3BD2B3FDEC" box="[337,486,598,622]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B4E2707115573D4BC557CB3BD2B3FDEC" box="[337,486,598,622]" italics="true" pageId="2" pageNumber="19">Epyris analis</emphasis>
</taxonomicName>
: Ƥ, 
<collectionCode id="E08734A615573D4BC61FCB3AD139FDEC" box="[537,620,598,622]" pageId="2" pageNumber="19">INDES</collectionCode>
[
<collectingCountry id="FE81ECF315573D4BC67CCB3BD184FDED" box="[634,721,599,623]" name="India" pageId="2" pageNumber="19">INDIA</collectingCountry>
], 
<emphasis id="B4E2707115573D4BC6E2CB3AD04AFDEC" box="[740,799,598,622]" italics="true" pageId="2" pageNumber="19">Mahi</emphasis>
{=Chennai} [13.083333, 80.283333], Ernest André collection 1914, Muséum Paris EY6341 (
<collectionCode id="E08734A615573D4BC65FCB16D1E0FD10" LSID="urn:lsid:biocol.org:col:34988" box="[601,693,634,658]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="19">MNHN</collectionCode>
). 
<typeStatus id="592D12C115573D4BC6CDCB16D00AFD10" box="[715,863,634,658]" pageId="2" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
of 
<taxonomicName id="4196D7E015573D4BC78ACB17D7C4FD11" box="[908,1169,635,659]" class="Insecta" family="Bethylidae" genus="Isobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B4E2707115573D4BC78ACB17D7C4FD11" box="[908,1169,635,659]" italics="true" pageId="2" pageNumber="19">Isobrachium rugicollis</emphasis>
</taxonomicName>
: Ƥ, [
<collectingCountry id="FE81ECF315573D4BC0D7CB16D608FD10" box="[1233,1373,634,658]" name="Pakistan" pageId="2" pageNumber="19">PAKISTAN</collectingCountry>
, 
<emphasis id="B4E2707115573D4BC16ACB17D20BFD34" italics="true" pageId="2" pageNumber="19">Balochistan province</emphasis>
] 
<emphasis id="B4E2707115573D4BC568CBF3D2EFFD34" box="[366,442,671,694]" italics="true" pageId="2" pageNumber="19">Quetta</emphasis>
, [30.1872, 67.0125], 6.02.{= 
<date id="F2288AA315573D4BC70FCBF2D030FD34" box="[777,869,670,694]" pageId="2" pageNumber="19" value="1902-06">VI.1902</date>
?}, Nurse col. 
<date id="F2288AA315573D4BC008CBF2D739FD34" box="[1038,1132,670,694]" pageId="2" pageNumber="19" value="1915" valueMax="1934">1915-34</date>
(
<collectionCode id="E08734A615573D4BC07CCBF2D780FD34" box="[1146,1237,670,694]" pageId="2" pageNumber="19">BMNH</collectionCode>
).
</paragraph>
<paragraph id="8629AC6315573D4BC4C3CBAED1D7FB9C" blockId="2.[151,1437,310,2026]" pageId="2" pageNumber="19">
<emphasis id="B4E2707115573D4BC4C3CBAED175FD59" bold="true" box="[197,544,706,731]" pageId="2" pageNumber="19">Afrotropical material (new).</emphasis>
<collectionCode id="E08734A615573D4BC62ACBAFD134FD58" box="[556,609,707,730]" httpUri="http://grbio.org/cool/xxq3-2kv6" name="University of Northern Iowa" pageId="2" pageNumber="19">UNI</collectionCode>
[TED] 
<collectionCode id="E08734A615573D4BC6B5CBAFD054FD59" box="[691,769,707,731]" pageId="2" pageNumber="19">ARAB</collectionCode>
EMIRATES, 3 Ƥ, 
<emphasis id="B4E2707115573D4BC7EACBAFD7C6FD59" box="[1004,1171,707,731]" italics="true" pageId="2" pageNumber="19">Wadi Wurayah</emphasis>
, 
<geoCoordinate id="E3A2CAA415573D4BC0AECBAED7AAFD58" box="[1192,1279,706,730]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="555" value="25.24">25.24N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC109CBAED637FD58" box="[1295,1378,706,730]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="555" value="56.17">56.17E</geoCoordinate>
, 
<date id="F2288AA315573D4BC174CBAED240FD7C" pageId="2" pageNumber="19" value="2005-04-12" valueMax="2005-04-14" valueMin="2005-04-12">12– 14.IV.2005</date>
, M[alaise] T[rap] &amp; Y[ellow] P[an] T[rap], T. P[ape] col. (
<collectionCode id="E08734A615573D4BC7D5CB8AD748FD7C" box="[979,1053,742,766]" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="2" pageNumber="19">UFES</collectionCode>
). 2 Ƥ, 
<emphasis id="B4E2707115573D4BC072CB8AD798FD7C" box="[1140,1229,742,766]" italics="true" pageId="2" pageNumber="19">Sharjah</emphasis>
, 
<geoCoordinate id="E3A2CAA415573D4BC0D9CB8AD660FD7C" box="[1247,1333,742,766]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="555" value="25.21">25.21N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC143CB8AD6C2FD7C" box="[1349,1431,742,766]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="555" value="55.42">55.42E</geoCoordinate>
, 
<date id="F2288AA315573D4BC491CA66D20EFCA0" box="[151,347,778,802]" pageId="2" pageNumber="19" value="2005-04-27" valueMax="2005-05-05" valueMin="2005-04-27">27.IV–05.V.2005</date>
, light tr[ap], A. v[an] Harten col. (
<collectionCode id="E08734A615573D4BC6E6CA66D073FCA0" box="[736,806,778,802]" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="2" pageNumber="19">UFES</collectionCode>
no. 30424, no. 30425). 13 Ƥ, 
<emphasis id="B4E2707115573D4BC07BCA67D780FCA1" box="[1149,1237,779,803]" italics="true" pageId="2" pageNumber="19">Bithnah</emphasis>
, 
<geoCoordinate id="E3A2CAA415573D4BC0E2CA66D66CFCA0" box="[1252,1337,778,802]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="555" value="25.11">25.11N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC143CA66D6C2FCA0" box="[1349,1431,778,802]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="555" value="56.14">56.14E</geoCoordinate>
, 
<date id="F2288AA315573D4BC491CA42D22FFCC4" box="[151,378,814,838]" pageId="2" pageNumber="19" value="2006-07-04" valueMax="2006-07-12" valueMin="2006-07-04">04.VII–12.VII.2006</date>
, M[alaise] T[rap], A. van Harten col. (
<collectionCode id="E08734A615573D4BC72BCA42D026FCC4" box="[813,883,814,838]" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="2" pageNumber="19">UFES</collectionCode>
no. 30483); 2 Ƥ, 
<date id="F2288AA315573D4BC03ACA42D649FCC4" box="[1084,1308,814,838]" pageId="2" pageNumber="19" value="2006-08-12" valueMax="2006-09-09" valueMin="2006-08-12">12.VIII–09.IX.2006</date>
(
<collectionCode id="E08734A615573D4BC12ACA42D627FCC4" box="[1324,1394,814,838]" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="2" pageNumber="19">UFES</collectionCode>
no. 30422); 2 Ƥ, 
<date id="F2288AA315573D4BC536CA3ED2A0FCE8" box="[304,501,850,874]" pageId="2" pageNumber="19" value="2006-10-19" valueMax="2006-11-16" valueMin="2006-10-19">19.X–16.XI.2006</date>
(
<collectionCode id="E08734A615573D4BC603CA3ED11AFCE8" box="[517,591,850,874]" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="2" pageNumber="19">UFES</collectionCode>
). 3 Ƥ, 
<emphasis id="B4E2707115573D4BC69ACA3FD055FCE9" box="[668,768,851,875]" italics="true" pageId="2" pageNumber="19">Al Ajban</emphasis>
, 
<geoCoordinate id="E3A2CAA415573D4BC709CA3ED033FCE8" box="[783,870,850,874]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="555" value="24.36">24.36N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC774CA3ED091FCE8" box="[882,964,850,874]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="555" value="55.01">55.01E</geoCoordinate>
, 
<date id="F2288AA315573D4BC7D7CA3ED7FEFCE8" box="[977,1195,850,874]" pageId="2" pageNumber="19" value="2006-06-26" valueMax="2006-07-25" valueMin="2006-06-26">26.VI–25.VII.2006</date>
, M[alaise] T[rap], A. van Harten col. (
<collectionCode id="E08734A615573D4BC55DCA1AD2F4FC0C" box="[347,417,886,910]" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="2" pageNumber="19">UFES</collectionCode>
no. 30423, no. 30593). 
<collectingCountry id="FE81ECF315573D4BC6BACA1BD076FC0C" box="[700,803,887,910]" name="Yemen" pageId="2" pageNumber="19">YEMEN</collectingCountry>
, 1 Ƥ, 
<emphasis id="B4E2707115573D4BC769CA1BD0DCFC0C" box="[879,905,887,910]" italics="true" pageId="2" pageNumber="19">Se</emphasis>
[
<emphasis id="B4E2707115573D4BC794CA1BD0CFFC0C" box="[914,922,887,910]" italics="true" pageId="2" pageNumber="19">i</emphasis>
] 
<emphasis id="B4E2707115573D4BC7A2CA14D099FC0D" box="[932,972,888,911]" italics="true" pageId="2" pageNumber="19">yun</emphasis>
, [15.94092, 48.780134], 
<date id="F2288AA315573D4BC0F4CA1AD604FC0C" box="[1266,1361,886,910]" pageId="2" pageNumber="19" value="2002-06">VI.2002</date>
, Light trap, A. van Harten &amp; G. Ba Saheh col. (
<collectionCode id="E08734A615573D4BC65CCAF6D1F6FC30" box="[602,675,922,946]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="2" pageNumber="19">CNCI</collectionCode>
). 2 Ƥ, 
<emphasis id="B4E2707115573D4BC6EBCAF7D036FC31" box="[749,867,923,947]" italics="true" pageId="2" pageNumber="19">Al Lahima</emphasis>
, [15.4, 43.5333333], 
<date id="F2288AA315573D4BC052CAF6D670FC30" box="[1108,1317,922,946]" pageId="2" pageNumber="19" value="2001-04-09" valueMax="2001-06-05" valueMin="2001-04-09">09.IV–05.VI.2001</date>
, M[alaise] T[rap], A. van Harten col. (
<collectionCode id="E08734A615573D4BC5CECAD2D144FC54" box="[456,529,958,982]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="2" pageNumber="19">CNCI</collectionCode>
). 1 Ƥ 
<emphasis id="B4E2707115573D4BC652CAD2D1D2FC54" box="[596,647,958,982]" italics="true" pageId="2" pageNumber="19">Lahj</emphasis>
, [13.066666, 44.883333], 
<date id="F2288AA315573D4BC7B6CAD2D75AFC54" box="[944,1039,958,982]" pageId="2" pageNumber="19" value="2000-09">IX.2000</date>
, M[alaise] T[rap], A. van Harten &amp; A. Sallam col. (
<collectionCode id="E08734A615573D4BC554CA8ED2CEFC78" box="[338,411,994,1018]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="2" pageNumber="19">CNCI</collectionCode>
); 1 Ƥ, 
<date id="F2288AA315573D4BC5F4CA8ED11BFC78" box="[498,590,994,1018]" pageId="2" pageNumber="19" value="2000-11">XI.2000</date>
(
<collectionCode id="E08734A615573D4BC667CA8ED1FFFC78" box="[609,682,994,1018]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="2" pageNumber="19">CNCI</collectionCode>
); 3 Ƥ, 
<date id="F2288AA315573D4BC707CA8ED029FC78" box="[769,892,994,1018]" pageId="2" pageNumber="19" value="2001-04" valueMax="2001-05-31" valueMin="2001-04-00">IV–V.2001</date>
(
<collectionCode id="E08734A615573D4BC789CA8ED08DFC78" box="[911,984,994,1018]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="2" pageNumber="19">CNCI</collectionCode>
); 3 Ƥ, 
<date id="F2288AA315573D4BC029CA8ED7A1FC78" box="[1071,1268,994,1018]" pageId="2" pageNumber="19" value="2001-05-17" valueMax="2001-06-15" valueMin="2001-05-17">17.V–15.VI.2001</date>
(
<collectionCode id="E08734A615573D4BC101CA8ED605FC78" box="[1287,1360,994,1018]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="2" pageNumber="19">CNCI</collectionCode>
); 2 Ƥ, 
<date id="F2288AA315573D4BC491CD6AD243FB9C" box="[151,278,1030,1054]" pageId="2" pageNumber="19" value="2001-09-07">07.IX.2001</date>
(
<collectionCode id="E08734A615573D4BC523CD6AD23BFB9C" box="[293,366,1030,1054]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="2" pageNumber="19">CNCI</collectionCode>
); 2 Ƥ, 
<date id="F2288AA315573D4BC5BCCD6AD14BFB9C" box="[442,542,1030,1054]" pageId="2" pageNumber="19" value="2002-05-03">3.V.2002</date>
(
<collectionCode id="E08734A615573D4BC62BCD6AD123FB9C" box="[557,630,1030,1054]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="2" pageNumber="19">CNCI</collectionCode>
).
</paragraph>
<paragraph id="8629AC6315573D4BC4C3CD46D6C9F868" blockId="2.[151,1437,310,2026]" pageId="2" pageNumber="19">
<emphasis id="B4E2707115573D4BC4C3CD46D2BDFBC1" bold="true" box="[197,488,1066,1091]" pageId="2" pageNumber="19">Oriental material (new).</emphasis>
[
<collectingCountry id="FE81ECF315573D4BC5F3CD47D1C5FBC1" box="[501,656,1067,1091]" name="Myanmar" pageId="2" pageNumber="19">MYANMAR</collectingCountry>
], 1 Ƥ, 
<emphasis id="B4E2707115573D4BC6DACD47D07EFBC1" box="[732,811,1067,1091]" italics="true" pageId="2" pageNumber="19">Bhamò</emphasis>
, [24.2667, 97.2333], 
<date id="F2288AA315573D4BC019CD46D7C4FBC0" box="[1055,1169,1066,1090]" pageId="2" pageNumber="19" value="1886-08">VIII.1886</date>
, L. Fea col. (
<collectionCode id="E08734A615573D4BC123CD46D62EFBC0" box="[1317,1403,1066,1090]" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale Giacomo Doria" pageId="2" pageNumber="19">MCSN</collectionCode>
). 1 Ƥ, 
<emphasis id="B4E2707115573D4BC4BDCD22D206FBE4" box="[187,339,1102,1126]" italics="true" pageId="2" pageNumber="19">Palon (Pegú)</emphasis>
, [17.4333, 95.9], 
<date id="F2288AA315573D4BC619CD22D1E1FBE4" box="[543,692,1102,1126]" pageId="2" pageNumber="19" value="1887-07" valueMax="1887-09-31" valueMin="1887-07-00">VII–IX.1887</date>
, L. Fea col (
<collectionCode id="E08734A615573D4BC745CD22D0CCFBE4" box="[835,921,1102,1126]" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale Giacomo Doria" pageId="2" pageNumber="19">MCSN</collectionCode>
). 
<collectingCountry id="FE81ECF315573D4BC7ABCD23D714FBE5" box="[941,1089,1103,1127]" name="Thailand" pageId="2" pageNumber="19">THAILAND</collectingCountry>
, 1 Ƥ, 
<emphasis id="B4E2707115573D4BC08ECD3CD667FBE4" box="[1160,1330,1103,1127]" italics="true" pageId="2" pageNumber="19">Mae Hong Son</emphasis>
, Namtok Mae Surin N[ational] P[ark], Walkway top of reservoir, 
<geoCoordinate id="E3A2CAA415573D4BC70FCD1ED0CFFB08" box="[777,922,1138,1162]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="19.348217">19°20.893'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC7A3CD1ED767FB08" box="[933,1074,1138,1162]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="97.98341">97°59.005'E</geoCoordinate>
, 
<date id="F2288AA315573D4BC03BCD1ED785FB08" box="[1085,1232,1138,1162]" pageId="2" pageNumber="19" value="2008-02-03" valueMax="2008-02-10" valueMin="2008-02-03">3–10.II.2008</date>
, M[alaise] T[rap], Na-maadkam &amp; M col. T3485 (
<collectionCode id="E08734A615573D4BC60CCDFAD10EFB2C" box="[522,603,1174,1198]" pageId="2" pageNumber="19">QSBG</collectionCode>
). 2 Ƥ, 
<emphasis id="B4E2707115573D4BC6A8CDFBD1B5FB2C" box="[686,736,1175,1198]" italics="true" pageId="2" pageNumber="19">Loei</emphasis>
, Phu Ruea N[ational] P[ark], Nern Pitsawong, 
<geoCoordinate id="E3A2CAA415573D4BC101CDFAD6CDFB2C" box="[1287,1432,1174,1198]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="17.4946">17°29.676'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC491CDD6D267FB50" box="[151,306,1210,1234]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="101.35155">101°21.093'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC546CDD6D2CCFB50" box="[320,409,1210,1234]" metricMagnitude="3" metricUnit="m" metricValue="1.168" pageId="2" pageNumber="19" unit="m" value="1168.0">1168 m</quantity>
, 
<date id="F2288AA315573D4BC5AECDD6D101FB50" box="[424,596,1210,1234]" pageId="2" pageNumber="19" value="2006-11-19" valueMax="2006-11-26" valueMin="2006-11-19">19–26.XI.2006</date>
, M[alaise] T[rap], Patikhom Tumtip col. T1123 (
<collectionCode id="E08734A615573D4BC089CDD6D7B5FB50" box="[1167,1248,1210,1234]" pageId="2" pageNumber="19">QSBG</collectionCode>
). 1 Ƥ, Phu Kradueng N[ational] P[ark], Road to Ta Krong waterfall of Na Noy Forest Unit, 
<geoCoordinate id="E3A2CAA415573D4BC00ECDB2D7CDFB74" box="[1032,1176,1246,1270]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="16.815216">16°48.913'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC0A3CDB2D66AFB74" box="[1189,1343,1246,1270]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="101.7939">101°47.634'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC14ACDB2D6C2FB74" box="[1356,1431,1246,1270]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="2" pageNumber="19" unit="m" value="265.0">265 m</quantity>
, 
<date id="F2288AA315573D4BC491CC6ED216FA98" box="[151,323,1282,1306]" pageId="2" pageNumber="19" value="2006-11-14" valueMax="2006-11-20" valueMin="2006-11-14">14–20.XI.2006</date>
, M[alaise] T[rap], Suthin Gong-lasae col. T1073 (
<collectionCode id="E08734A615573D4BC79ECC6ED0BCFA98" box="[920,1001,1282,1306]" pageId="2" pageNumber="19">QSBG</collectionCode>
). 1 Ƥ, 
<emphasis id="B4E2707115573D4BC044CC6FD640FA99" box="[1090,1301,1283,1307]" italics="true" pageId="2" pageNumber="19">Ubon Ratchathani</emphasis>
, Pha Taem N[ational] P[ark], Don Huay Can, 
<geoCoordinate id="E3A2CAA415573D4BC624CC4AD1E7FABC" box="[546,690,1318,1342]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="15.666933">15°40.016'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC6B9CC4AD00FFABC" box="[703,858,1318,1342]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="105.50837">105°30.502'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC761CC4AD0E4FABC" box="[871,945,1318,1342]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="2" pageNumber="19" unit="m" value="246.0">246 m</quantity>
, 
<date id="F2288AA315573D4BC7B8CC4AD708FABC" box="[958,1117,1318,1342]" pageId="2" pageNumber="19" value="2007-06-09" valueMax="2007-06-15" valueMin="2007-06-09">9–15.VI.2007</date>
, M[alaise] T[rap], Tongcam &amp; Banlu col. T2208 (
<collectionCode id="E08734A615573D4BC592CC26D2B0FAE0" box="[404,485,1354,1378]" pageId="2" pageNumber="19">QSBG</collectionCode>
); 2 Ƥ, Saeng Chan Waterfall, 
<geoCoordinate id="E3A2CAA415573D4BC74FCC26D08CFAE0" box="[841,985,1354,1378]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="15.533083">15°31.985'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC7EECC26D7D7FAE0" box="[1000,1154,1354,1378]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="105.59623">105°35.774'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC097CC26D78BFAE0" box="[1169,1246,1354,1378]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="2" pageNumber="19" unit="m" value="155.0">155 m</quantity>
, 
<date id="F2288AA315573D4BC0EBCC26D6CDFAE0" box="[1261,1432,1354,1378]" pageId="2" pageNumber="19" value="2007-03-12" valueMax="2007-03-20" valueMin="2007-03-12">12–20.III.2007</date>
, M[alaise] T[rap], Porntip Tonsu &amp; Bunlu Sapsiri col. T2146 (
<collectionCode id="E08734A615573D4BC74BCC02D0CBFA04" box="[845,926,1390,1414]" pageId="2" pageNumber="19">QSBG</collectionCode>
). 1 Ƥ, 
<emphasis id="B4E2707115573D4BC7ECCC02D72CFA04" box="[1002,1145,1390,1414]" italics="true" pageId="2" pageNumber="19">Chaiyaphum</emphasis>
, Pa Hin Ngam N[ational] P[ark], car park at Tung Dok Grajeaw, 
<geoCoordinate id="E3A2CAA415573D4BC64CCCFED18FFA28" box="[586,730,1426,1450]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="15.640634">15°38.438'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC6E0CCFED0D4FA28" box="[742,897,1426,1450]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="101.39294">101°23.576'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC78ACCFED080FA28" box="[908,981,1426,1450]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="2" pageNumber="19" unit="m" value="780.0">780 m</quantity>
, 
<date id="F2288AA315573D4BC7E6CCFED7C3FA28" box="[992,1174,1426,1450]" pageId="2" pageNumber="19" value="2006-07-12" valueMax="2006-07-18" valueMin="2006-07-12">12–18.VII.2006</date>
, M[alaise] T[rap], Kratae Sa-nog &amp; Buakaw Adnafai col. T333 (
<collectionCode id="E08734A615573D4BC67ECCDAD19CFA4C" box="[632,713,1462,1486]" pageId="2" pageNumber="19">QSBG</collectionCode>
); 2 Ƥ, creek at Tung Dok Grajeaw, 
<geoCoordinate id="E3A2CAA415573D4BC066CCDAD7A5FA4C" box="[1120,1264,1462,1486]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="15.63985">15°38.391'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC0FBCCDAD6C2FA4C" box="[1277,1431,1462,1486]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="101.393486">101°23.609'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC491CCB6D3B6FA70" box="[151,227,1498,1522]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="19" unit="m" value="750.0">750 m</quantity>
, 
<date id="F2288AA315573D4BC4F4CCB6D2FDFA70" box="[242,424,1498,1522]" pageId="2" pageNumber="19" value="2006-07-24" valueMax="2006-07-30" valueMin="2006-07-24">24–30.VII.2006</date>
, M[alaise] T[rap], Kratae Sa-nog &amp; Buakaw Adnafai col. T338 (
<collectionCode id="E08734A615573D4BC099CCB6D7A5FA70" box="[1183,1264,1498,1522]" pageId="2" pageNumber="19">QSBG</collectionCode>
); 1 Ƥ, ecotone between mix deciduous and dipterocarp forest, 
<geoCoordinate id="E3A2CAA415573D4BC6C9CC92D00AF994" box="[719,863,1534,1558]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="15.635533">15°38.132'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC776CC92D75EF994" box="[880,1035,1534,1558]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="101.3987">101°23.922'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC01BCC92D739F994" box="[1053,1132,1534,1558]" metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" pageId="2" pageNumber="19" unit="m" value="698.0">698 m</quantity>
, 
<date id="F2288AA315573D4BC078CC92D675F994" box="[1150,1312,1534,1558]" pageId="2" pageNumber="19" value="2007-02-19" valueMax="2007-02-25" valueMin="2007-02-19">19–25.II.2007</date>
, M[alaise] T[rap], Katae Sa-nog &amp; Buakaw Adnafai col. T1652 (
<collectionCode id="E08734A615573D4BC735CF4ED0D1F9B8" box="[819,900,1570,1594]" pageId="2" pageNumber="19">QSBG</collectionCode>
). 1 Ƥ, 
<emphasis id="B4E2707115573D4BC7E0CF4FD7DDF9B9" box="[998,1160,1571,1595]" italics="true" pageId="2" pageNumber="19">Kanchanaburi</emphasis>
, Khuean Srinagarindra N[ational] P[ark], Tourist center, 
<geoCoordinate id="E3A2CAA415573D4BC60FCF2AD1CFF9DC" box="[521,666,1606,1630]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="14.6356">14°38.136'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC6A2CF2AD064F9DC" box="[676,817,1606,1630]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="98.997284">98°59.837'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC73ACF2AD0D1F9DC" box="[828,900,1606,1630]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="19" unit="m" value="210.0">210 m</quantity>
, 
<date id="F2288AA315573D4BC789CF2AD711F9DC" box="[911,1092,1606,1630]" pageId="2" pageNumber="19" value="2008-07-11" valueMax="2008-07-18" valueMin="2008-07-11">11–18.VII.2008</date>
, M[alaise] T[rap], Somboon &amp; Chatchawan col. T3432 (
<collectionCode id="E08734A615573D4BC5C2CF06D140F900" box="[452,533,1642,1666]" pageId="2" pageNumber="19">QSBG</collectionCode>
). 1 Ƥ, 
<emphasis id="B4E2707115573D4BC675CF07D04AF901" box="[627,799,1643,1667]" italics="true" pageId="2" pageNumber="19">Nakhon Nayok</emphasis>
, Khao Yai N[ational] P[ark], Nhong ping khaokeaw, 
<geoCoordinate id="E3A2CAA415573D4BC491CFE2D272F924" box="[151,295,1678,1702]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="14.3849">14°23.094'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC531CFE2D287F924" box="[311,466,1678,1702]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="101.38425">101°23.055'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC5E4CFE2D165F924" box="[482,560,1678,1702]" metricMagnitude="2" metricUnit="m" metricValue="7.33" pageId="2" pageNumber="19" unit="m" value="733.0">733 m</quantity>
, 
<date id="F2288AA315573D4BC647CFE2D1B9F924" box="[577,748,1678,1702]" pageId="2" pageNumber="19" value="2007-03-12" valueMax="2007-03-19" valueMin="2007-03-12">12–19.III.2007</date>
, M[alaise] T[rap], Wirat Sukho col. T2104 (
<collectionCode id="E08734A615573D4BC0FBCFE2D61BF924" box="[1277,1358,1678,1702]" pageId="2" pageNumber="19">QSBG</collectionCode>
); 1 Ƥ, 
<date id="F2288AA315573D4BC491CFDED200F948" box="[151,341,1714,1738]" pageId="2" pageNumber="19" value="2007-03-26" valueMax="2007-04-02" valueMin="2007-03-26">26.III–2.IV.2007</date>
, Pong Sandao col. T2109 (
<collectionCode id="E08734A615573D4BC682CFDED180F948" box="[644,725,1714,1738]" pageId="2" pageNumber="19">QSBG</collectionCode>
). 1 Ƥ, 
<emphasis id="B4E2707115573D4BC726CFDFD0FAF949" box="[800,943,1715,1739]" italics="true" pageId="2" pageNumber="19">Chanthaburi</emphasis>
, Khao Khitchakut N[ational] P[ark], 
<quantity id="416E018615573D4BC153CFDED6C9F948" box="[1365,1436,1714,1738]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="19" unit="m" value="100.0">100 m</quantity>
N/Prabaht Unit, 
<geoCoordinate id="E3A2CAA415573D4BC554CFBAD2B6F96C" box="[338,483,1750,1774]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="12.8140335">12°48.842'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC5E9CFBAD129F96C" box="[495,636,1750,1774]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="102.1524">102°9.144'E</geoCoordinate>
, 
<quantity id="416E018615573D4BC68FCFBAD186F96C" box="[649,723,1750,1774]" metricMagnitude="2" metricUnit="m" metricValue="2.03" pageId="2" pageNumber="19" unit="m" value="203.0">203 m</quantity>
, 
<date id="F2288AA315573D4BC6E7CFBAD0DDF96C" box="[737,904,1750,1774]" pageId="2" pageNumber="19" value="2008-07-07" valueMax="2008-07-14" valueMin="2008-07-07">7–14.VII.2008</date>
, M[alaise] T[rap], Suthida &amp; Charoenchai col. T2973 (
<collectionCode id="E08734A615573D4BC4F4CF96D216F890" box="[242,323,1786,1810]" pageId="2" pageNumber="19">QSBG</collectionCode>
). 5 Ƥ, 
<emphasis id="B4E2707115573D4BC591CF97D1C4F891" box="[407,657,1787,1811]" italics="true" pageId="2" pageNumber="19">Prachuab Khiri Khan</emphasis>
, Khao Sam Roi Yot N[ational] P[ark], Saline wetland/Pa Gwad/N, 
<geoCoordinate id="E3A2CAA415573D4BC491CE72D3A8F8B4" box="[151,253,1822,1846]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="92" value="12.153334">12°9.2'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC50FCE72D2C3F8B4" box="[265,406,1822,1846]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="99.971634">99°58.298'E</geoCoordinate>
, 
<date id="F2288AA315573D4BC5A4CE72D118F8B4" box="[418,589,1822,1846]" pageId="2" pageNumber="19" value="2009-03-22" valueMax="2009-03-29" valueMin="2009-03-22">22–29.III.2009</date>
, M[alaise] T[rap], Yai &amp; Amnad col. T4219 (
<collectionCode id="E08734A615573D4BC056CE72D7F4F8B4" box="[1104,1185,1822,1846]" pageId="2" pageNumber="19">QSBG</collectionCode>
); 2 Ƥ, Saline wetland/ Pa Gwad/S, 
<geoCoordinate id="E3A2CAA415573D4BC523CE2ED2F2F8D8" box="[293,423,1858,1882]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="12.152884">12°9.173'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC5B2CE2ED114F8D8" box="[436,577,1858,1882]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="99.97073">99°58.244'E</geoCoordinate>
, T4221 (
<collectionCode id="E08734A615573D4BC6AECE2ED1ACF8D8" box="[680,761,1858,1882]" pageId="2" pageNumber="19">QSBG</collectionCode>
); 2 Ƥ, Nursery, 
<geoCoordinate id="E3A2CAA415573D4BC7B7CE2ED771F8D8" box="[945,1060,1858,1882]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="9" value="12.126333">12°7.58'N</geoCoordinate>
, 
<geoCoordinate id="E3A2CAA415573D4BC034CE2ED7EAF8D8" box="[1074,1215,1858,1882]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="99.95797">99°57.478'E</geoCoordinate>
, 
<date id="F2288AA315573D4BC0CACE2ED6CDF8D8" box="[1228,1432,1858,1882]" pageId="2" pageNumber="19" value="2008-06-29" valueMax="2008-07-06" valueMin="2008-06-29">29.VI–6.VII.2008</date>
, M[alaise] T[rap], Amnad &amp; Yai col. T3049 (
<collectionCode id="E08734A615573D4BC6A7CE0AD1A4F8FC" box="[673,753,1894,1918]" pageId="2" pageNumber="19">QSBG</collectionCode>
); 1 Ƥ, 
<date id="F2288AA315573D4BC741CE0AD746F8FC" box="[839,1043,1894,1918]" pageId="2" pageNumber="19" value="2008-06-30" valueMax="2008-07-01" valueMin="2008-06-30">30.VI–1.VII.2008</date>
, Pan trap, T3031 (
<collectionCode id="E08734A615573D4BC0E8CE0AD66AF8FC" box="[1262,1343,1894,1918]" pageId="2" pageNumber="19">QSBG</collectionCode>
); VIET- NAM, 1 Ƥ, 
<emphasis id="B4E2707115573D4BC51ACEE7D229F821" box="[284,380,1931,1955]" italics="true" pageId="2" pageNumber="19">Dak Lak</emphasis>
, Chu Yang Sin N[ational] P[ark], n[ea]r dam, [12.46277, 108.409464], c. 
<quantity id="416E018615573D4BC0B5CEE6D7A9F820" box="[1203,1276,1930,1954]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="19" unit="m" value="500.0">500 m</quantity>
, 
<date id="F2288AA315573D4BC101CEE6D6CDF820" box="[1287,1432,1930,1954]" pageId="2" pageNumber="19" value="2007-06-03" valueMax="2007-06-09" valueMin="2007-06-03">3–9.VI.2007</date>
, M[alaise] T[rap], C. v. Achterberg &amp; R. de Vries col. RMNH’07 (
<collectionCode id="E08734A615573D4BC771CEC2D084F844" LSID="urn:lsid:biocol.org:col:34992" box="[887,977,1966,1990]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="19">RMNH</collectionCode>
). 1 Ƥ, 
<emphasis id="B4E2707115573D4BC01DCEC2D7F5F844" box="[1051,1184,1966,1990]" italics="true" pageId="2" pageNumber="19">Ninh Thuân</emphasis>
, Núi Chiúa N[ational] P[ark], northeast part, [11.715754, 109.138353], M[alaise] T[rap], 
<quantity id="416E018615573D4BC786CEBED0A7F868" box="[896,1010,2002,2026]" metricMagnitude="2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="2" pageNumber="19" unit="m" value="120.0" valueMax="150.0" valueMin="90.0">90–150 m</quantity>
, 
<date id="F2288AA315573D4BC7F8CEBED7C8F868" box="[1022,1181,2002,2026]" pageId="2" pageNumber="19" value="2007-05-23" valueMax="2007-05-30" valueMin="2007-05-23">23–30.V.2007</date>
, C. v. Achterberg &amp; R.
</paragraph>
<paragraph id="8629AC6315563D4AC491C9FBD2E0FEE6" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="20">
de Vries col. RMNH’07 (
<collectionCode id="E08734A615563D4AC5B9C9FBD14FFF2D" LSID="urn:lsid:biocol.org:col:34992" box="[447,538,151,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="20">RMNH</collectionCode>
). 1 Ƥ, 
<emphasis id="B4E2707115563D4AC66BC9F4D188FF32" box="[621,733,152,176]" italics="true" pageId="3" pageNumber="20">Dông Nai</emphasis>
, Cát Tiên N[ational] P[ark], Bot[anical] garden, [11.470501, 107.259089], c. 
<quantity id="416E018615563D4AC552C9D0D2F4FF56" box="[340,417,188,212]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="20" unit="m" value="100.0">100 m</quantity>
, 
<date id="F2288AA315563D4AC5B6C9D0D11AFF56" box="[432,591,188,212]" pageId="3" pageNumber="20" value="2005-05-13" valueMax="2005-05-20" valueMin="2005-05-13">13–20.V.2005</date>
, M[alaise] T[rap] 14-19, C. v. Achterberg &amp; R. de Vries col. RMNH’05 (
<collectionCode id="E08734A615563D4AC499C9B3D3AFFF75" LSID="urn:lsid:biocol.org:col:34992" box="[159,250,223,247]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="20">RMNH</collectionCode>
). 2 Ƥ, Cát Tiên N[ational] P[ark], Dong trail, [11.470501, 107.259089], c. 
<quantity id="416E018615563D4AC084C9B3D785FF75" box="[1154,1232,223,247]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="20" unit="m" value="100.0">100 m</quantity>
, 
<date id="F2288AA315563D4AC0E4C9B3D6CDFF75" box="[1250,1432,223,247]" pageId="3" pageNumber="20" value="2007-04-09" valueMax="2007-05-19" valueMin="2007-04-09">9.IV–19.V.2007</date>
, M[alaise] T[rap], M. P. Quy, N. T. Manh &amp; C. v. Achterberg col. RMNH’07 (
<collectionCode id="E08734A615563D4AC015C868D73BFE9E" LSID="urn:lsid:biocol.org:col:34992" box="[1043,1134,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="20">RMNH</collectionCode>
). 1 Ƥ, Cát Tiên N[ational] P[ark], Ficus trail, [11.470501, 107.259089], c. 
<quantity id="416E018615563D4AC6B3C84BD1AAFEBD" box="[693,767,295,319]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="20" unit="m" value="100.0">100 m</quantity>
, 
<date id="F2288AA315563D4AC70BC84BD0F2FEBD" box="[781,935,295,319]" pageId="3" pageNumber="20" value="2007-04-09" valueMax="2007-04-30" valueMin="2007-04-09">9–30.IV.2007</date>
, M[alaise] T[rap], M. P. Quy &amp; N. T. Manh col. RMNH’07 (
<collectionCode id="E08734A615563D4AC549C820D2FCFEE6" LSID="urn:lsid:biocol.org:col:34992" box="[335,425,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="20">RMNH</collectionCode>
).
</paragraph>
<paragraph id="8629AC6315563D4AC4C3C81CD02CFE2E" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="20">
<emphasis id="B4E2707115563D4AC4C3C81CD22AFE0A" bold="true" box="[197,383,367,392]" pageId="3" pageNumber="20">
<typeStatus id="592D12C115563D4AC4C3C81CD255FE0A" box="[197,256,368,392]" pageId="3" pageNumber="20">Type</typeStatus>
condition.
</emphasis>
The 
<typeStatus id="592D12C115563D4AC5BBC803D176FE05" box="[445,547,367,391]" pageId="3" pageNumber="20" type="lectotype">lectotype</typeStatus>
and the three female 
<typeStatus id="592D12C115563D4AC719C803D094FE05" box="[799,961,367,391]" pageId="3" pageNumber="20" type="paralectotype">paralectotypes</typeStatus>
are in good condition. However the only 
<typeStatus id="592D12C115563D4AC491C8F8D27BFE2E" box="[151,302,404,428]" pageId="3" pageNumber="20" type="paralectotype">paralectotype</typeStatus>
male is broken in several parts, but none is missing.
</paragraph>
</subSubSection>
<subSubSection id="CE8CFFE815563D4AC4C3C8DBD249FD95" pageId="3" pageNumber="20" type="distribution">
<paragraph id="8629AC6315563D4AC4C3C8DBD249FD95" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="20">
<emphasis id="B4E2707115563D4AC4C3C8DBD20BFE52" bold="true" box="[197,350,439,464]" pageId="3" pageNumber="20">Distribution.</emphasis>
<collectingCountry id="FE81ECF315563D4AC560C8DBD1A0FE4D" box="[358,757,439,463]" name="Democratic Republic of the Congo" pageId="3" pageNumber="20">Democratic Republic of the Congo</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC704C8DBD031FE4D" box="[770,868,439,463]" name="Ethiopia" pageId="3" pageNumber="20">Ethiopia</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC776C8DBD74DFE4D" box="[880,1048,439,463]" name="Guinea-Bissau" pageId="3" pageNumber="20">Guinea-Bissau</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC023C8DBD737FE4D" box="[1061,1122,439,463]" name="India" pageId="3" pageNumber="20">India</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC068C8DBD7FAFE4D" box="[1134,1199,439,463]" name="Israel" pageId="3" pageNumber="20">Israel</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC0BDC8D4D67FFE52" box="[1211,1322,440,464]" name="Myanmar" pageId="3" pageNumber="20">Myanmar</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC130C8DBD6CDFE4D" box="[1334,1432,439,463]" name="Pakistan" pageId="3" pageNumber="20">Pakistan</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC491C8B0D27CFE76" box="[151,297,476,500]" name="South Africa" pageId="3" pageNumber="20">South Africa</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC535C8B0D22CFE76" box="[307,377,476,500]" name="Sudan" pageId="3" pageNumber="20">Sudan</collectingCountry>
and 
<collectingCountry id="FE81ECF315563D4AC5A8C8B0D17CFE76" box="[430,553,476,500]" name="Zimbabwe" pageId="3" pageNumber="20">Zimbabwe</collectingCountry>
. It is recorded for the first time from 
<collectingCountry id="FE81ECF315563D4AC7C2C8B0D77FFE76" box="[964,1066,476,500]" name="Thailand" pageId="3" pageNumber="20">Thailand</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC033C8B0D679FE76" box="[1077,1324,476,500]" name="United Arab Emirates" pageId="3" pageNumber="20">United Arab Emirates</collectingCountry>
, 
<collectingCountry id="FE81ECF315563D4AC130C8B0D6CDFE76" box="[1334,1432,476,500]" name="Vietnam" pageId="3" pageNumber="20">Vietnam</collectingCountry>
, and 
<collectingCountry id="FE81ECF315563D4AC4C0CB6CD24DFD95" box="[198,280,512,535]" name="Yemen" pageId="3" pageNumber="20">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CE8CFFE815563D4AC4C3CB4FD1BEFD2A" pageId="3" pageNumber="20" type="diagnosis">
<paragraph id="8629AC6315563D4AC4C3CB4FD1BEFD2A" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="20">
<emphasis id="B4E2707115563D4AC4C3CB4FD26AFDBE" bold="true" box="[197,319,547,572]" pageId="3" pageNumber="20">Diagnosis.</emphasis>
FEMALE. Mandible with pre-apical lower tooth, sensillae chaetica in lower margin of mandible present. Apex of clypeus with long setae. Head as wide as long. Space between inner and second pair of propodeal discal carina with a structure resembling a fovea. Metatibia with dense series of short setae in posterior face. Metasoma dark castaneous nearly black with apex orange.
</paragraph>
</subSubSection>
<subSubSection id="CE8CFFE815563D4DC4C3CBDFD027FF75" lastPageId="4" lastPageNumber="21" pageId="3" pageNumber="20" type="description">
<paragraph id="8629AC6315563D4AC4C3CBDFD0EEFD4E" blockId="3.[151,1437,151,2012]" box="[197,955,691,716]" pageId="3" pageNumber="20">
<emphasis id="B4E2707115563D4AC4C3CBDFD226FD4E" bold="true" box="[197,371,691,716]" pageId="3" pageNumber="20">Redescription.</emphasis>
<typeStatus id="592D12C115563D4AC57FCBD8D148FD4E" box="[377,541,692,716]" pageId="3" pageNumber="20" type="lectotype">LECTOTYPE</typeStatus>
. Body length 
<quantity id="416E018615563D4AC6BECBD8D044FD4E" box="[696,785,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="20" unit="mm" value="7.5">7.5 mm</quantity>
. LFW 
<quantity id="416E018615563D4AC758CBD8D0E2FD4E" box="[862,951,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="20" unit="mm" value="3.8">3.8 mm</quantity>
.
</paragraph>
<paragraph id="8629AC6315563D4AC4C3CBB4D7C3FCB5" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="20">
<emphasis id="B4E2707115563D4AC4C3CBB4D251FD72" box="[197,260,728,752]" italics="true" pageId="3" pageNumber="20">Color</emphasis>
. Head, 
<taxonomicName id="4196D7E015563D4AC566CBB5D283FD72" box="[352,470,729,752]" class="Crustacea" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">mesosoma</taxonomicName>
and procoxa dark castaneous nearly black. Metasoma dark castaneous nearly black with apex orange. Meso and metacoxae and femora dark castaneous. Scape, pedicel, flagellum, mandible, tegula, wing venation, trochanters and tibiae castaneous. Palpi light castaneous. Wings subhyaline.
</paragraph>
<paragraph id="8629AC6315563D4AC4C3CA28D1FBFB46" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="20">
<emphasis id="B4E2707115563D4AC4C3CA28D257FCDE" box="[197,258,836,860]" italics="true" pageId="3" pageNumber="20">Head</emphasis>
(
<figureCitation id="1EADB0E615563D4AC517CA28D207FCDE" box="[273,338,836,860]" captionStart="FIGURES 1 – 6" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[151,1436,195,1882]" captionTargetId="figure@6.[151,1436,195,1883]" captionTargetPageId="6" captionText="FIGURES 1 – 6. Head. 1. Epyris afer Ƥ, lectotype, dorsal view. 2 – 3. E. enerterus sp. nov. Ƥ, holotype. 2. Dorsal view; 3. Ventral view. 4. E. afer 3, paralectotype, dorsal view. 5 – 6. E. penatii sp. nov. 3, holotype. 5. Dorsal view; 6. Ventral view. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204625/files/figure.png" pageId="3" pageNumber="20">Fig. 1</figureCitation>
). Mandible (
<figureCitation id="1EADB0E615563D4AC5E2CA28D11AFCDE" box="[484,591,836,860]" captionStart="FIGURES 7 – 18" captionStartId="7.[151,264,1085,1108]" captionTargetBox="[184,1407,198,1061]" captionTargetId="figure@7.[178,1409,193,1064]" captionTargetPageId="7" captionText="FIGURES 7 – 18. Mandible. 7 – 9. Epyris afer Ƥ, lectotype. 7. Frontal view; 8. Ventral view; 9. Dorsal view. 10 – 12. E. afer 3, paralectotype. 10. Frontal view; 11. Ventral view; 12. Dorsal view. 13 – 15. E. enerterus sp. nov. Ƥ, holotype. 13. Frontal view; 14. Ventral view; 15. Dorsal view. 16 – 18. E. penatii sp. nov. 3, holotype. 16. Frontal view; 17. Ventral view; 18. Dorsal view. (Scale bar = 200 µm)." httpUri="https://zenodo.org/record/204626/files/figure.png" pageId="3" pageNumber="20">Figs. 7–9</figureCitation>
) apex as wide as base, with five rounded apical teeth, two lower wider than others, pre-apical lower tooth present; sensillae chaetica in lower margin of mandible present. Clypeus shorter than wide, median lobe angulate; median carina present; apex of clypeus with long setae; lateral lobe reduced. Transversal section of scape cylindrical; first four antennal segments ratio 23:7:7:9, segment III as long as thick; antennal sensillae not visible. Antennal scrobe projected forward, dorsally sulcate. Toruli distant from each other 1.7 × their diameter. Eye almost reaching upper mandibular condyle, prominent, glabrous. Frons coriaceous; punctures small and shallow, separated from each other by 1.5 × their diameter. WH 1.0 × LH; WF 1.3 × HE; WF 0.6 × WH; OOL 1.7 × WOT; VOL 0.6 × HE; distance of posterior ocellus to vertex crest 1.0 × 
<collectingCountry id="FE81ECF315563D4AC00ACD2CD718FBDA" box="[1036,1101,1088,1112]" name="Iceland" pageId="3" pageNumber="20">DAO</collectingCountry>
. Ocelli small, frontal angle of ocellar triangle acute. Temple parallel in dorsal view. Vertex almost straight in dorsal view, with soft callosity medially. Crest with continuous series of short setae. Occipital carina present dorsally and ventrally. Gena flattened. Hypostomal carina straight and polished.
</paragraph>
<paragraph id="8629AC6315563D4AC4C3CDBDD01EF887" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="20">
<taxonomicName id="4196D7E015563D4AC4C3CDBDD268FB6A" box="[197,317,1233,1256]" class="Crustacea" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115563D4AC4C3CDBDD268FB6A" box="[197,317,1233,1256]" italics="true" pageId="3" pageNumber="20">Mesosoma</emphasis>
</taxonomicName>
(
<figureCitation id="1EADB0E615563D4AC548CDBCD2F6FB6A" box="[334,419,1232,1256]" captionStart="FIGURES 19 – 22" captionStartId="9.[151,264,1914,1937]" captionTargetBox="[151,1436,193,1892]" captionTargetId="figure@9.[151,1436,193,1893]" captionTargetPageId="9" captionText="FIGURES 19 – 22. Mesosoma in dorsal view and mesopleuron. 19 – 20. Epyris afer Ƥ, lectotype. 19. Mesosoma; 20. Mesopleuron. 21 – 22. E. enerterus sp. nov. Ƥ, holotype. 21. Mesosoma; 22. Mesopleuron. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204627/files/figure.png" pageId="3" pageNumber="20">Fig. 19</figureCitation>
). Pronotal collar polished, with transversal groove. Pronotal disc 0.9 × longer than wide, coriaceous, subtrapezoidal, sparsely punctate, transversal carina in anterior margin absent. Mesoscutum coriaceous, lateral slightly elevate posteriorly. Notaulus touching posterior margin of mesoscutum, convergent posteriorly, gradual anteroposterior enlargement. Parapsidal furrow parallel, shorter and as deep as notaulus, not reaching posterior margin of mesoscutum. Scutellum coriaceous, scutellar groove absent, scutellar pit oval. Propodeal disc 0.9 × as long as wide; anterior carina medially narrower than laterally; anterior corner fovea-shaped; five discal carinae present, median carina complete, inner discal carina complete, space between median and inner discal carina with longitudinal ridge, second pair of discal carina incomplete and short, space between inner and second pair of discal carina with structure resembling fovea; lateral margin straight; sublateral carina absent, lateral carina present, space between sublateral and lateral carina with series of transversal ridges; posterior margin somewhat concave; posterior corner with suboval fovea; lateral of propodeum strigate; declivity of propodeum strigate, with median carina. Mesopleuron (
<figureCitation id="1EADB0E615563D4AC5E1CF30D16DF9F6" box="[487,568,1628,1652]" captionStart="FIGURES 19 – 22" captionStartId="9.[151,264,1914,1937]" captionTargetBox="[151,1436,193,1892]" captionTargetId="figure@9.[151,1436,193,1893]" captionTargetPageId="9" captionText="FIGURES 19 – 22. Mesosoma in dorsal view and mesopleuron. 19 – 20. Epyris afer Ƥ, lectotype. 19. Mesosoma; 20. Mesopleuron. 21 – 22. E. enerterus sp. nov. Ƥ, holotype. 21. Mesosoma; 22. Mesopleuron. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204627/files/figure.png" pageId="3" pageNumber="20">Fig. 20</figureCitation>
) coriaceous; subtegular groove elongate, large anteriorly and narrow posteriorly; anterior fovea closed; sub-anterior fovea closed and small; mesopleural fovea closed; mesopleural pit present; mesopleural elevation present; lower fovea open, groove of inferior margin of lower fovea present throughout extension of margin; episternal furrow with inner margin striate. Epicnemium lateromedially enlarged, with medial constriction. Pleurosternum with straight longitudinal groove.
</paragraph>
<paragraph id="8629AC6315563D4AC4C3CE7DD0EBF8F2" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="20">
<emphasis id="B4E2707115563D4AC4C3CE7DD3AFF8AA" box="[197,250,1809,1832]" italics="true" pageId="3" pageNumber="20">Legs</emphasis>
. Profemur 2.4 × longer than wide. Mesotibia spinose. Metatibia with dense series of short setae in posterior face. Tarsomeres distally spinose, spines of protarsus short and cylindrical, spines of meso and metatarsi long and flattened. Tarsal claws bidentate, inner tooth as long as apical tooth.
</paragraph>
<paragraph id="8629AC6315563D4AC4C3CE11D3E9F85E" blockId="3.[151,1437,151,2012]" pageId="3" pageNumber="20">
<emphasis id="B4E2707115563D4AC4C3CE11D3A8F816" box="[197,253,1917,1940]" italics="true" pageId="3" pageNumber="20">Wing</emphasis>
. Forewing with metacarpus present, 0.2 × as long as stigma length; radial vein curved forward, 2.6 × longer than basal; basal vein truncate; transverse median vein convex. 
<collectingCountry id="FE81ECF315563D4AC78CCECCD097F83A" box="[906,962,1952,1976]" name="India" pageId="3" pageNumber="20">Hind</collectingCountry>
wing with two basal and four apical hamuli.
</paragraph>
<paragraph id="8629AC6315513D4DC4C3C9F5D756FF56" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3C9F5D26CFF32" box="[197,313,153,176]" italics="true" pageId="4" pageNumber="21">Metasoma</emphasis>
. Ventral surface of petiole (
<figureCitation id="1EADB0E615513D4DC677C9FBD196FF2D" box="[625,707,151,175]" captionStart="FIGURES 27 – 30" captionStartId="11.[151,264,1358,1381]" captionTargetBox="[151,1436,195,1336]" captionTargetId="figure@11.[151,1436,195,1336]" captionTargetPageId="11" captionText="FIGURES 27 – 30. Petiole in ventral view. 27. Epyris afer Ƥ, lectotype. 28. E. enerterus sp. nov. Ƥ, holotype. 29. E. afer 3, paralectotype. 30. E. penatii sp. nov. 3, holotype. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204629/files/figure.png" pageId="4" pageNumber="21">Fig. 27</figureCitation>
) unsegmented, space between carinae of petiole wide. Tergum II longer than others; terga III–VI with line of transverse setae on dorsal surface.
</paragraph>
<paragraph id="8629AC6315513D4DC4C3C98DD027FF75" blockId="4.[151,1437,151,2012]" box="[197,882,223,248]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3C98DD278FF7A" box="[197,301,225,248]" italics="true" pageId="4" pageNumber="21">Variation</emphasis>
. 
<collectingCountry id="FE81ECF315513D4DC53DC9B3D226FF75" box="[315,371,223,247]" name="India" pageId="4" pageNumber="21">Hind</collectingCountry>
wing with three basal and five apical hamuli.
</paragraph>
</subSubSection>
<subSubSection id="CE8CFFE815513D4DC4C3C86FD0C2FE05" pageId="4" pageNumber="21" type="diagnosis">
<paragraph id="8629AC6315513D4DC4C3C86FD0C2FE05" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3C86FD26AFE9E" bold="true" box="[197,319,259,284]" pageId="4" pageNumber="21">Diagnosis.</emphasis>
MALE. Posterior ocellus distant from vertex crest 0.3 × 
<collectingCountry id="FE81ECF315513D4DC7D8C868D74BFE9E" box="[990,1054,260,284]" name="Iceland" pageId="4" pageNumber="21">DAO</collectingCountry>
. Mesoscutum and scutellum polished. Mesotibia without spines. Protarsomeres without spines. 
<collectingCountry id="FE81ECF315513D4DC779C84BD0E2FEBD" box="[895,951,295,319]" name="India" pageId="4" pageNumber="21">Hind</collectingCountry>
wing with five medial hamuli. Paramere strongly arched ventrad, narrow and almost 3.0 × longer than basiparamere. Aedeagus inner margin slightly expanded subapically. Apodeme base projected laterally, not dilated.
</paragraph>
</subSubSection>
<subSubSection id="CE8CFFE815513D4DC4C3C8FFD19DF85E" pageId="4" pageNumber="21" type="description">
<paragraph id="8629AC6315513D4DC4C3C8FFD750FE2E" blockId="4.[151,1437,151,2012]" box="[197,1029,403,428]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3C8FFD226FE2E" bold="true" box="[197,371,403,428]" pageId="4" pageNumber="21">Redescription.</emphasis>
<typeStatus id="592D12C115513D4DC57FC8F8D132FE2E" box="[377,615,404,428]" pageId="4" pageNumber="21" type="paralectotype">PARALECTOTYPE</typeStatus>
. Body length 4.0 mm. LFW 
<quantity id="416E018615513D4DC7AEC8F8D754FE2E" box="[936,1025,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="21" unit="mm" value="2.2">2.2 mm</quantity>
.
</paragraph>
<paragraph id="8629AC6315513D4DC4C3C8D4D0A0FD95" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3C8D4D251FE52" box="[197,260,440,464]" italics="true" pageId="4" pageNumber="21">Color</emphasis>
. Head, 
<taxonomicName id="4196D7E015513D4DC55DC8D4D284FE4D" box="[347,465,440,463]" class="Crustacea" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="30" phylum="Arthropoda" rank="genus">mesosoma</taxonomicName>
and coxae dark castaneous nearly black. Metasoma dark castaneous nearly black with apex castaneous. Wing venation, trochanters and femora dark castaneous. Scape, pedicel, flagellum, mandible, tegula and tibiae castaneous. Palpi and tarsi light castaneous. Wings subhyaline.
</paragraph>
<paragraph id="8629AC6315513D4DC4C3CB48D14FFC26" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3CB48D257FDBE" box="[197,258,548,572]" italics="true" pageId="4" pageNumber="21">Head</emphasis>
(
<figureCitation id="1EADB0E615513D4DC514CB48D20CFDBE" box="[274,345,548,572]" captionStart="FIGURES 1 – 6" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[151,1436,195,1882]" captionTargetId="figure@6.[151,1436,195,1883]" captionTargetPageId="6" captionText="FIGURES 1 – 6. Head. 1. Epyris afer Ƥ, lectotype, dorsal view. 2 – 3. E. enerterus sp. nov. Ƥ, holotype. 2. Dorsal view; 3. Ventral view. 4. E. afer 3, paralectotype, dorsal view. 5 – 6. E. penatii sp. nov. 3, holotype. 5. Dorsal view; 6. Ventral view. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204625/files/figure.png" pageId="4" pageNumber="21">Fig. 4</figureCitation>
). Mandible (
<figureCitation id="1EADB0E615513D4DC5ECCB48D127FDBE" box="[490,626,548,572]" captionStart="FIGURES 7 – 18" captionStartId="7.[151,264,1085,1108]" captionTargetBox="[184,1407,198,1061]" captionTargetId="figure@7.[178,1409,193,1064]" captionTargetPageId="7" captionText="FIGURES 7 – 18. Mandible. 7 – 9. Epyris afer Ƥ, lectotype. 7. Frontal view; 8. Ventral view; 9. Dorsal view. 10 – 12. E. afer 3, paralectotype. 10. Frontal view; 11. Ventral view; 12. Dorsal view. 13 – 15. E. enerterus sp. nov. Ƥ, holotype. 13. Frontal view; 14. Ventral view; 15. Dorsal view. 16 – 18. E. penatii sp. nov. 3, holotype. 16. Frontal view; 17. Ventral view; 18. Dorsal view. (Scale bar = 200 µm)." httpUri="https://zenodo.org/record/204626/files/figure.png" pageId="4" pageNumber="21">Figs. 10–12</figureCitation>
) apex as wide as base, with five rounded apical teeth, two lower wider than others, pre-apical lower tooth present; sensillae chaetica in lower margin of mandible present. Clypeus shorter than wide, median lobe angulate; median carina present; apex of clypeus with long setae; lateral lobe reduced. Transversal section of scape elliptical; first four antennal segments ratio 18:5:4:7, segment III 0.6 × longer than thick; antennal sensillae visible in ventral surface of antenna. Antennal scrobe projected forward, not dorsally carinate. Toruli distant from each other 1.5 × their diameter. Eye almost reaching upper mandibular condyle, prominent, glabrous. Frons coriaceous; punctures small and shallow, separated from each other by 3.0 × their diameter. WH 0.9 × LH; WF 1.5 × HE; WF 0.7 × WH; OOL 1.9 × WOT; VOL 1.0 × HE; distance of posterior ocellus to vertex crest 0.3 × 
<collectingCountry id="FE81ECF315513D4DC540CA28D2D3FCDE" box="[326,390,836,860]" name="Iceland" pageId="4" pageNumber="21">DAO</collectingCountry>
. Ocelli small, frontal angle of ocellar triangle acute. Temple parallel in dorsal view. Vertex straight in dorsal view. Crest without setae. Occipital carina present dorsally and ventrally. Gena flattened. Hypostomal carina straight and polished.
</paragraph>
<paragraph id="8629AC6315513D4DC4C3CADDD13DFA67" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="21">
<taxonomicName id="4196D7E015513D4DC4C3CADDD268FC4A" box="[197,317,945,968]" class="Crustacea" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115513D4DC4C3CADDD268FC4A" box="[197,317,945,968]" italics="true" pageId="4" pageNumber="21">Mesosoma</emphasis>
</taxonomicName>
(
<figureCitation id="1EADB0E615513D4DC54DCAC3D2CAFC45" box="[331,415,943,967]" captionStart="FIGURES 23 – 26" captionStartId="10.[151,264,1865,1888]" captionTargetBox="[151,1436,193,1844]" captionTargetId="figure@10.[151,1436,193,1844]" captionTargetPageId="10" captionText="FIGURES 23 – 26. Mesosoma in dorsal view and mesopleuron. 23 – 24. Epyris afer 3, paralectotype. 23. Mesosoma; 24. Mesopleuron. 25 – 26. E. penatii sp. nov. 3, holotype. 25. Mesosoma; 26. Mesopleuron. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204628/files/figure.png" pageId="4" pageNumber="21">Fig. 23</figureCitation>
). Pronotal collar polished, without transversal groove. Pronotal disc 0.6 × longer than wide, coriaceous, trapezoidal, sparsely punctate, transversal carina in anterior margin present. Mesoscutum polished, lateral slightly elevate posteriorly. Notaulus touching posterior margin of mesoscutum, convergent posteriorly, gradual anteroposterior enlargement. Parapsidal furrow convergent posteriorly, shorter and as deep as notaulus, not reaching posterior margin of mesoscutum. Scutellum polished, scutellar groove absent, scutellar pit circular. Propodeal disc 0.5 × as long as wide; anterior carina medially narrower than laterally; anterior corner groove-shaped; five discal carinae present, median carina complete, inner discal carina incomplete, space between median and inner discal carina without sculpture, second pair of discal carina complete, space between inner and second pair of discal carina without ridge; lateral margin sinuous; sublateral carina present and weak, lateral carina present, space between sublateral and lateral carina with series of transversal ridges; posterior margin somewhat concave; posterior corner with suboval fovea; lateral of propodeum strigate; declivity of propodeum areolate, with median carina. Mesopleuron (
<figureCitation id="1EADB0E615513D4DC53BCC50D2C4FAD6" box="[317,401,1340,1364]" captionStart="FIGURES 23 – 26" captionStartId="10.[151,264,1865,1888]" captionTargetBox="[151,1436,193,1844]" captionTargetId="figure@10.[151,1436,193,1844]" captionTargetPageId="10" captionText="FIGURES 23 – 26. Mesosoma in dorsal view and mesopleuron. 23 – 24. Epyris afer 3, paralectotype. 23. Mesosoma; 24. Mesopleuron. 25 – 26. E. penatii sp. nov. 3, holotype. 25. Mesosoma; 26. Mesopleuron. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204628/files/figure.png" pageId="4" pageNumber="21">Fig. 24</figureCitation>
) coriaceous; subtegular groove elongate, large anteriorly and narrow posteriorly; anterior fovea closed and large; sub-anterior fovea closed and large; mesopleural fovea open; mesopleural pit present; mesopleural elevation present; lower fovea open, groove of inferior margin of lower fovea present only in anterior portion of margin; episternal furrow with inner margin striate. Epicnemium lateromedially enlarged. Pleurosternum with lozenge-shaped longitudinal groove.
</paragraph>
<paragraph id="8629AC6315513D4DC4C3CC9DD04AF9D2" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3CC9DD3AFF98A" box="[197,250,1521,1544]" italics="true" pageId="4" pageNumber="21">Legs</emphasis>
. Profemur 2.8 × longer than wide. Mesotibia without spines. Metatibia with sparse series of short setae in posterior face. Protarsomeres without spines. Meso and metatarsomeres distally spinose, spines long and flattened. Tarsal claws bidentate, inner tooth as long as apical tooth.
</paragraph>
<paragraph id="8629AC6315513D4DC4C3CF31D62DF91A" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3CF31D3A8F9F6" box="[197,253,1629,1652]" italics="true" pageId="4" pageNumber="21">Wing</emphasis>
. Forewing with metacarpus present, 0.7 × as long as stigma length; radial vein curved forward, 3.0 × longer than basal; basal vein slightly concave; transverse median vein convex. 
<collectingCountry id="FE81ECF315513D4DC7E0CFECD74BF91A" box="[998,1054,1664,1688]" name="India" pageId="4" pageNumber="21">Hind</collectingCountry>
wing with five medial hamuli.
</paragraph>
<paragraph id="8629AC6315513D4DC4C3CFC9D0C7F962" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3CFC9D26CF93E" box="[197,313,1701,1724]" italics="true" pageId="4" pageNumber="21">Metasoma</emphasis>
. Ventral surface of petiole (
<figureCitation id="1EADB0E615513D4DC675CFC8D191F93E" box="[627,708,1700,1724]" captionStart="FIGURES 27 – 30" captionStartId="11.[151,264,1358,1381]" captionTargetBox="[151,1436,195,1336]" captionTargetId="figure@11.[151,1436,195,1336]" captionTargetPageId="11" captionText="FIGURES 27 – 30. Petiole in ventral view. 27. Epyris afer Ƥ, lectotype. 28. E. enerterus sp. nov. Ƥ, holotype. 29. E. afer 3, paralectotype. 30. E. penatii sp. nov. 3, holotype. (Scale bar = 200 µm)" httpUri="https://zenodo.org/record/204629/files/figure.png" pageId="4" pageNumber="21">Fig. 29</figureCitation>
) unsegmented, space between carinae of petiole narrow. Tergum II as long as others; terga III–VI with sparse setae on dorsal surface.
</paragraph>
<paragraph id="8629AC6315513D4DC4C3CF80D19DF85E" blockId="4.[151,1437,151,2012]" pageId="4" pageNumber="21">
<emphasis id="B4E2707115513D4DC4C3CF80D27AF886" box="[197,303,1772,1796]" italics="true" pageId="4" pageNumber="21">Genitalia</emphasis>
(
<figureCitation id="1EADB0E615513D4DC539CF80D296F886" box="[319,451,1772,1796]" captionStart="FIGURES 31 – 36" captionStartId="12.[151,264,1915,1938]" captionTargetBox="[151,1415,208,1890]" captionTargetId="figure@12.[151,1436,193,1894]" captionTargetPageId="12" captionText="FIGURES 31 – 36. Genitalia. 31 – 33. Epyris afer 3, paralectotype. 31. Dorsal view; 32. Ventral view. 33. Lateral view. 34 – 36. E. penatii sp. nov. 3, holotype. 34. Dorsal view; 35. Ventral view. 36. Lateral view. (Scale bar = 250 µm)" httpUri="https://zenodo.org/record/204630/files/figure.png" pageId="4" pageNumber="21">Figs 31–33</figureCitation>
). Paramere positioned dorsally, arched strongly ventrad, narrow, very elongate, almost 3.0 × longer than basiparamere, apical margin convex, membranous expansion large and stout; cuspis elongate, its apex surpassing apical half of paramere length, divided into two arms only at apex, its ventral base with rounded projection, ventral arm slightly longer and narrower than dorsal arm; digitus with apical margin denticulate, its apex not reaching cuspis apex; aedeagus bottle-shaped, short, its apex not reaching cuspis length, apex with concavity inside, inner margin slightly expanded subapically; inner lobe membranous; apodeme not extending beyond genital ring, base projected lateral and not dilated.
</paragraph>
</subSubSection>
<subSubSection id="CE8CFFE815503D4CC4C3C9FBD0FFF870" pageId="5" pageNumber="22" type="discussion">
<paragraph id="8629AC6315503D4CC4C3C9FBD751FEBD" blockId="5.[151,1437,151,2035]" pageId="5" pageNumber="22">
<emphasis id="B4E2707115503D4CC4C3C9FBD21FFF32" bold="true" box="[197,330,151,176]" pageId="5" pageNumber="22">Discussion.</emphasis>
This species does not belong to 
<emphasis id="B4E2707115503D4CC6D7C9F5D069FF32" box="[721,828,153,176]" italics="true" pageId="5" pageNumber="22">Disepyris</emphasis>
because the scutellar pits are not connected by any groove, the setae of the protarsomeres are not stout, the eyes are not large, and the stigma is not large. Besides, it has the pronotal disc ecarinate, wider posteriorly and the anterior corners not acutely angled, the mesoscutum has notauli and the antennae are not spinose. This combination of characteristics corresponds to 
<taxonomicName id="4196D7E015503D4CC0A2C868D7B8FE99" box="[1188,1261,260,283]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC0A2C868D7B8FE99" box="[1188,1261,260,283]" italics="true" pageId="5" pageNumber="22">Epyris</emphasis>
</taxonomicName>
and because of that we revalidated the combination to 
<taxonomicName id="4196D7E015503D4CC64AC844D1C0FEBD" box="[588,661,296,319]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC64AC844D1C0FEBD" box="[588,661,296,319]" italics="true" pageId="5" pageNumber="22">Epyris</emphasis>
</taxonomicName>
as proposed by 
<bibRefCitation id="E207D19215503D4CC74DC84BD754FEBD" author="Magretti" box="[843,1025,295,319]" pageId="5" pageNumber="22" refString="Magretti, P. (1897) Viaggio di Leonardo Fea in Birmania e regioni Vicine 74. Imenotteri. Parte Seconda. Trigonalidi, Betilidi, Crisididi colla descrizione de parecchie nuove specie. Annali de Museo Civico di Storia Naturale d'Genova. Series 2, 17, 308 - 326." type="journal article" year="1897">Magretti (1897)</bibRefCitation>
.
</paragraph>
<paragraph id="8629AC6315503D4CC4C3C826D097FE04" blockId="5.[151,1437,151,2035]" pageId="5" pageNumber="22">
The 
<typeStatus id="592D12C115503D4CC4F1C826D20BFEE0" box="[247,350,330,354]" pageId="5" pageNumber="22" type="lectotype">lectotype</typeStatus>
designation was necessary because there were three different species in the 
<typeStatus id="592D12C115503D4CC0ADC827D78EFEE1" box="[1195,1243,331,355]" pageId="5" pageNumber="22">type</typeStatus>
series. The specimen selected represents Magretti’s original idea for the name of 
<taxonomicName id="4196D7E015503D4CC776C803D0EEFE04" box="[880,955,366,390]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC776C803D0EEFE04" box="[880,955,366,390]" italics="true" pageId="5" pageNumber="22">E. afer</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8629AC6315503D4CC4C3C8FED1A6FD97" blockId="5.[151,1437,151,2035]" pageId="5" pageNumber="22">
Three species are considered junior subjective synonyms of 
<taxonomicName id="4196D7E015503D4CC774C8FFD0EAFE28" box="[882,959,402,426]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC774C8FFD0EAFE28" box="[882,959,402,426]" italics="true" pageId="5" pageNumber="22">E. afer</emphasis>
</taxonomicName>
, namely 
<taxonomicName id="4196D7E015503D4CC02EC8FFD7F8FE28" box="[1064,1197,402,426]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B4E2707115503D4CC02EC8FFD7F8FE28" box="[1064,1197,402,426]" italics="true" pageId="5" pageNumber="22">E. pilosipes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4196D7E015503D4CC0BAC8FFD63EFE28" box="[1212,1387,402,426]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="plurilineatus">
<emphasis id="B4E2707115503D4CC0BAC8FFD63EFE28" box="[1212,1387,402,426]" italics="true" pageId="5" pageNumber="22">E. plurilineatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4196D7E015503D4CC491C8DBD3A9FE4C" box="[151,252,438,462]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="B4E2707115503D4CC491C8DBD3A9FE4C" box="[151,252,438,462]" italics="true" pageId="5" pageNumber="22">E. analis</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4196D7E015503D4CC519C8DAD2D3FE4C" box="[287,390,438,462]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="secundus">
<emphasis id="B4E2707115503D4CC519C8DAD2D3FE4C" box="[287,390,438,462]" italics="true" pageId="5" pageNumber="22">secundus</emphasis>
</taxonomicName>
) because they have the mandible with five rounded apical teeth, the two lower ones wider than the upper ones; the presence of long setae in the apex of median clypeal lobe; the presence of occipital carina ventrally, and the propodeal anterior corner foveolate.
</paragraph>
<paragraph id="8629AC6315503D4CC4C3CB4DD39CFCF8" blockId="5.[151,1437,151,2035]" pageId="5" pageNumber="22">
Another species, 
<taxonomicName id="4196D7E015503D4CC588CB4DD1C4FDBB" box="[398,657,545,569]" class="Insecta" family="Bethylidae" genus="Isobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B4E2707115503D4CC588CB4DD1C4FDBB" box="[398,657,545,569]" italics="true" pageId="5" pageNumber="22">Isobrachium rugicollis</emphasis>
</taxonomicName>
is also considered as junior subjective synonym of 
<taxonomicName id="4196D7E015503D4CC0F2CB4ED616FDBB" box="[1268,1347,545,569]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC0F2CB4ED616FDBB" box="[1268,1347,545,569]" italics="true" pageId="5" pageNumber="22">E. afer</emphasis>
</taxonomicName>
. It was described in 
<taxonomicName id="4196D7E015503D4CC52ECB2AD224FDDF" box="[296,369,582,605]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC52ECB2AD224FDDF" box="[296,369,582,605]" italics="true" pageId="5" pageNumber="22">Epyris</emphasis>
</taxonomicName>
by 
<bibRefCitation id="E207D19215503D4CC59BCB28D103FDDE" author="Cameron" box="[413,598,580,604]" pageId="5" pageNumber="22" refString="Cameron, P. (1906) On the Tenthredinidae &amp; parasitic Hymenoptera collected in Baluchistan by Major C. G. Nurse. Journal of Bombay Natural History Society, 17 (2), 89 - 288." type="journal article" year="1906">
<collectingCountry id="FE81ECF315503D4CC59BCB28D151FDDE" box="[413,516,580,604]" name="Cameroon" pageId="5" pageNumber="22">Cameron</collectingCountry>
(1906)
</bibRefCitation>
based on one single female. However 
<bibRefCitation id="E207D19215503D4CC009CB28D7E7FDDE" author="Kieffer" box="[1039,1202,580,604]" pageId="5" pageNumber="22" refString="Kieffer, J. J. (1908) Hymenoptera. Fam. Bethylidae. In: Wytsman, P., Genera Insectorum. Fasc. 76. V. Verteneuil &amp; L. Desmet Imprimeurs-Editeurs, Bruxelles, 50 pp. + 3 pls." type="book" year="1908">Kieffer (1908)</bibRefCitation>
transferred it to 
<taxonomicName id="4196D7E015503D4CC176CB2AD2CAFD02" authority="Forster, 1856" authorityName="Forster" authorityYear="1856" class="Insecta" family="Bethylidae" genus="Isobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC176CB2AD256FD02" italics="true" pageId="5" pageNumber="22">Isobrachium</emphasis>
Förster, 1856
</taxonomicName>
, although he did not explain the reason for this nomenclatural act. 
<taxonomicName id="4196D7E015503D4CC080CB04D646FD02" box="[1158,1299,616,640]" class="Insecta" family="Bethylidae" genus="Isobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC080CB04D646FD02" box="[1158,1299,616,640]" italics="true" pageId="5" pageNumber="22">Isobrachium</emphasis>
</taxonomicName>
has as main diagnostic character the lack of notauli, which is absent in this species, according to the original description. Cameron’s description may have led Kieffer to propose the transference to 
<taxonomicName id="4196D7E015503D4CC7CDCBDCD70DFD4A" box="[971,1112,688,712]" class="Insecta" family="Bethylidae" genus="Isobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC7CDCBDCD70DFD4A" box="[971,1112,688,712]" italics="true" pageId="5" pageNumber="22">Isobrachium</emphasis>
</taxonomicName>
. However, our observations showed the presence of notauli in the 
<typeStatus id="592D12C115503D4CC646CBBFD1F0FD69" box="[576,677,723,747]" pageId="5" pageNumber="22" type="holotype">holotype</typeStatus>
, so, the inclusion of this species in 
<taxonomicName id="4196D7E015503D4CC035CBB8D794FD6E" box="[1075,1217,724,748]" class="Insecta" family="Bethylidae" genus="Isobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC035CBB8D794FD6E" box="[1075,1217,724,748]" italics="true" pageId="5" pageNumber="22">Isobrachium</emphasis>
</taxonomicName>
became unjustified and we revalidate the name established by 
<bibRefCitation id="E207D19215503D4CC675CB9BD07FFC8D" author="Cameron" box="[627,810,759,783]" pageId="5" pageNumber="22" refString="Cameron, P. (1906) On the Tenthredinidae &amp; parasitic Hymenoptera collected in Baluchistan by Major C. G. Nurse. Journal of Bombay Natural History Society, 17 (2), 89 - 288." type="journal article" year="1906">
<collectingCountry id="FE81ECF315503D4CC675CB9BD18FFC8D" box="[627,730,759,783]" name="Cameroon" pageId="5" pageNumber="22">Cameron</collectingCountry>
(1906)
</bibRefCitation>
and propose it as an 
<taxonomicName id="4196D7E015503D4CC015CB94D709FC8D" box="[1043,1116,760,783]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC015CB94D709FC8D" box="[1043,1116,760,783]" italics="true" pageId="5" pageNumber="22">Epyris</emphasis>
</taxonomicName>
species. Nevertheless, when we transferred the species to 
<taxonomicName id="4196D7E015503D4CC5E6CA70D17CFCB1" box="[480,553,796,819]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC5E6CA70D17CFCB1" box="[480,553,796,819]" italics="true" pageId="5" pageNumber="22">Epyris</emphasis>
</taxonomicName>
, we realized that it is too similar to 
<taxonomicName id="4196D7E015503D4CC7C4CA70D75BFCB1" box="[962,1038,795,819]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC7C4CA70D75BFCB1" box="[962,1038,795,819]" italics="true" pageId="5" pageNumber="22">E. afer</emphasis>
</taxonomicName>
exactly by same reasons explained above to 
<taxonomicName id="4196D7E015503D4CC4FBCA2CD2D5FCD5" box="[253,384,831,855]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B4E2707115503D4CC4FBCA2CD2D5FCD5" box="[253,384,831,855]" italics="true" pageId="5" pageNumber="22">E. pilosipes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4196D7E015503D4CC588CA2CD141FCD5" box="[398,532,831,855]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="secundus">
<emphasis id="B4E2707115503D4CC588CA2CD141FCD5" box="[398,532,831,855]" italics="true" pageId="5" pageNumber="22">E. secundus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4196D7E015503D4CC64CCA2CD1ADFCD5" box="[586,760,831,855]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="plurilineatus">
<emphasis id="B4E2707115503D4CC64CCA2CD1ADFCD5" box="[586,760,831,855]" italics="true" pageId="5" pageNumber="22">E. plurilineatus</emphasis>
</taxonomicName>
. Thus we considered it as a junior subjective synonym of 
<taxonomicName id="4196D7E015503D4CC182CA2CD396FCF9" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC182CA2CD396FCF9" italics="true" pageId="5" pageNumber="22">E. afer</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8629AC6315503D4CC4C3CAEBD727FBD3" blockId="5.[151,1437,151,2035]" pageId="5" pageNumber="22">
<taxonomicName id="4196D7E015503D4CC4C3CAEBD217FC1C" box="[197,322,902,926]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC4C3CAEBD217FC1C" box="[197,322,902,926]" italics="true" pageId="5" pageNumber="22">Epyris afer</emphasis>
</taxonomicName>
has sexual dimorphism. The main differences between both sexes are: the females have VOL 0.6 × HE, the distance of posterior ocellus to vertex crest 1.0 × 
<collectingCountry id="FE81ECF315503D4CC724CAC6D034FC40" box="[802,865,938,962]" name="Iceland" pageId="5" pageNumber="22">DAO</collectingCountry>
, the scutellar pits oval, the propodeal disc 0.9 × as long as wide with anterior corner foveolate, the mesotibiae spinose and the forewings with metacarpus 0.2 × as long as stigma length, whereas the male has VOL 1.0 × HE, the distance of posterior ocellus to vertex crest 0.3 × 
<collectingCountry id="FE81ECF315503D4CC491CD79D382FBAF" box="[151,215,1045,1069]" name="Iceland" pageId="5" pageNumber="22">DAO</collectingCountry>
, the scutellar pits circular, the propodeal disc 0.5 × as long as wide with anterior corner groove-shaped, the mesotibiae without spines, the forewing with metacarpus 0.7 × as long as stigma length.
</paragraph>
<paragraph id="8629AC6315503D4CC4C3CD30D2DBFAF2" blockId="5.[151,1437,151,2035]" pageId="5" pageNumber="22">
The females of 
<taxonomicName id="4196D7E015503D4CC572CD32D2A4FBF7" box="[372,497,1117,1141]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC572CD32D2A4FBF7" box="[372,497,1117,1141]" italics="true" pageId="5" pageNumber="22">Epyris afer</emphasis>
</taxonomicName>
are similar to 
<taxonomicName id="4196D7E015503D4CC695CD32D08DFBF6" authority="Cameron,1909" authorityName="Cameron" authorityYear="1909" box="[659,984,1116,1141]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="hirtipennis">
<emphasis id="B4E2707115503D4CC695CD32D07EFBF7" box="[659,811,1117,1141]" italics="true" pageId="5" pageNumber="22">E. hirtipennis</emphasis>
<bibRefCitation id="E207D19215503D4CC734CD30D08DFBF6" author="Cameron" box="[818,984,1116,1140]" pageId="5" pageNumber="22" refString="Cameron, P. (1909) On an undescribed genus and species of Oxyura (Proctotrypidae) from Kuching, Borneo (Hym.). Deutsche Entomologische Zeitschrift, 377 - 378." type="book chapter" year="1909">
<collectingCountry id="FE81ECF315503D4CC734CD30D0C8FBF6" box="[818,925,1116,1140]" name="Cameroon" pageId="5" pageNumber="22">Cameron</collectingCountry>
,1909
</bibRefCitation>
</taxonomicName>
because both species have the mandible with five rounded apical teeth, the two lower ones wider than the upper ones, the pre-apical lower tooth present and the sensillae chaetica in lower margin, the distance of posterior ocellus to vertex crest 1.0 × 
<collectingCountry id="FE81ECF315503D4CC0B3CDC8D7A0FB3E" box="[1205,1269,1188,1212]" name="Iceland" pageId="5" pageNumber="22">DAO</collectingCountry>
, the propodeal disc 0.9 × as long as wide and the protarsomeres with spines short and cylindrical. However, 
<taxonomicName id="4196D7E015503D4CC0AFCDA5D7A0FB62" box="[1193,1269,1224,1248]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC0AFCDA5D7A0FB62" box="[1193,1269,1224,1248]" italics="true" pageId="5" pageNumber="22">E. afer</emphasis>
</taxonomicName>
has the scutellar pits oval, the space between inner and second pair of propodeal discal carina with a structure resembling a fovea and the pleurosternum with straight longitudinal groove, whereas 
<taxonomicName id="4196D7E015503D4CC782CC7DD748FAA5" box="[900,1053,1295,1320]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="hirtipennis">
<emphasis id="B4E2707115503D4CC782CC7DD748FAA5" box="[900,1053,1295,1320]" italics="true" pageId="5" pageNumber="22">E. hirtipennis</emphasis>
</taxonomicName>
has the scutellar pits circular, the space between inner and second pair of propodeal discal carina without fovea-like structure, and the pleurosternum with elliptical groove.
</paragraph>
<paragraph id="8629AC6315503D4CC4C3CC16D23AF9EB" blockId="5.[151,1437,151,2035]" pageId="5" pageNumber="22">
On the other hand, the male of 
<taxonomicName id="4196D7E015503D4CC618CC10D1CEFA11" box="[542,667,1403,1427]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC618CC10D1CEFA11" box="[542,667,1403,1427]" italics="true" pageId="5" pageNumber="22">Epyris afer</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4196D7E015503D4CC72ACC10D0CDFA11" box="[812,920,1404,1427]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="penatii" status="sp. nov.">
<emphasis id="B4E2707115503D4CC72ACC10D0CDFA11" box="[812,920,1404,1427]" italics="true" pageId="5" pageNumber="22">E. penatii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFD1CD0A15503D4CC798CC17D0A2FA11" box="[926,1015,1403,1427]" pageId="5" pageNumber="22" rank="species">
<emphasis id="B4E2707115503D4CC798CC17D0A2FA11" bold="true" box="[926,1015,1403,1427]" pageId="5" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
because both species have the scutellar pits circular, the propodeal disc ranging 0.5–0.6 × as long as wide, the anterior fovea of mesopleuron closed and large and the parameres positioned dorsally and elongate. However, 
<taxonomicName id="4196D7E015503D4CC79DCCAFD0B2FA58" box="[923,999,1474,1498]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC79DCCAFD0B2FA58" box="[923,999,1474,1498]" italics="true" pageId="5" pageNumber="22">E. afer</emphasis>
</taxonomicName>
has mandibular pre-apical lower tooth and the sensillae chaetica present, the eyes glabrous, VOL 1.0 × HE, the protarsomeres without spines and the inner lobe of aedeagus without setae, whereas 
<taxonomicName id="4196D7E015503D4CC667CF66D19AF9A3" box="[609,719,1546,1569]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="penatii" status="sp. nov.">
<emphasis id="B4E2707115503D4CC667CF66D19AF9A3" box="[609,719,1546,1569]" italics="true" pageId="5" pageNumber="22">E. penatii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFD1CD0A15503D4CC6D0CF66D065F9A0" box="[726,816,1546,1570]" pageId="5" pageNumber="22" rank="species">
<emphasis id="B4E2707115503D4CC6D0CF66D065F9A0" bold="true" box="[726,816,1546,1570]" pageId="5" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
has the mandibular pre-apical lower tooth and the sensillae chaetica absent, the eyes weakly pilose, VOL 0.5 × HE, the protarsomeres distally spinose and the inner lobe of aedeagus setose.
</paragraph>
<paragraph id="8629AC6315503D4CC4C3CF1AD0FFF870" blockId="5.[151,1437,151,2035]" pageId="5" pageNumber="22">
<taxonomicName id="4196D7E015503D4CC4C3CF1AD217F90F" box="[197,322,1653,1677]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC4C3CF1AD217F90F" box="[197,322,1653,1677]" italics="true" pageId="5" pageNumber="22">Epyris afer</emphasis>
</taxonomicName>
was previously known from 
<collectingCountry id="FE81ECF315503D4CC68CCF18D19EF90E" box="[650,715,1652,1676]" name="Israel" pageId="5" pageNumber="22">Israel</collectingCountry>
, 
<collectingCountry id="FE81ECF315503D4CC6D1CF18D06CF90E" box="[727,825,1652,1676]" name="Ethiopia" pageId="5" pageNumber="22">Ethiopia</collectingCountry>
, 
<collectingCountry id="FE81ECF315503D4CC742CF18D0B9F90E" box="[836,1004,1652,1676]" name="Guinea-Bissau" pageId="5" pageNumber="22">Guinea-Bissau</collectingCountry>
, 
<collectingCountry id="FE81ECF315503D4CC7F1CF18D60DF90E" box="[1015,1368,1652,1676]" name="Democratic Republic of the Congo" pageId="5" pageNumber="22">Democratic Republic of Congo</collectingCountry>
, 
<collectingCountry id="FE81ECF315503D4CC162CF18D3B6F932" name="Zimbabwe" pageId="5" pageNumber="22">Zimbabwe</collectingCountry>
, 
<collectingCountry id="FE81ECF315503D4CC4F7CFF4D2D2F932" box="[241,391,1688,1712]" name="South Africa" pageId="5" pageNumber="22">South Africa</collectingCountry>
, 
<collectingCountry id="FE81ECF315503D4CC592CFF4D2A6F932" box="[404,499,1688,1712]" name="Pakistan" pageId="5" pageNumber="22">Pakistan</collectingCountry>
and 
<collectingCountry id="FE81ECF315503D4CC636CFF5D1C8F933" box="[560,669,1689,1713]" name="Myanmar" pageId="5" pageNumber="22">Myanmar</collectingCountry>
based on observations of 
<specimenCount id="909067EA15503D4CC7C8CFF4D73CF932" box="[974,1129,1688,1712]" pageId="5" pageNumber="22" type="generic">26 specimens</specimenCount>
(
<figureCitation id="1EADB0E615503D4CC07DCFF4D785F932" box="[1147,1232,1688,1712]" captionStart="FIGURE 37" captionStartId="13.[151,250,1326,1349]" captionTargetBox="[154,1435,195,1304]" captionTargetId="figure@13.[151,1436,193,1305]" captionTargetPageId="13" captionText="FIGURE 37. Current known distribution of Epyris afer. Red circles represent the previous data and blue squares represent the new data of species." httpUri="https://zenodo.org/record/204631/files/figure.png" pageId="5" pageNumber="22">Fig. 37</figureCitation>
, red circles). The large gap among these sites suggests that this species has a broad distribution in several sites through Afrotropical and Oriental regions, and even through Mediterranean region since it occurs in 
<collectingCountry id="FE81ECF315503D4CC01CCF8CD709F97A" box="[1050,1116,1760,1784]" name="Israel" pageId="5" pageNumber="22">Israel</collectingCountry>
. We have searched for more specimens within the material from regions where collection have been recently carried out, namely 
<collectingCountry id="FE81ECF315503D4CC11ACE68D63FF899" box="[1308,1386,1796,1819]" name="Yemen" pageId="5" pageNumber="22">Yemen</collectingCountry>
and 
<collectingCountry id="FE81ECF315503D4CC491CE44D384F8C2" box="[151,209,1832,1856]" name="United Arab Emirates" pageId="5" pageNumber="22">UAE</collectingCountry>
(projects coordinated by A. van Harten), 
<collectingCountry id="FE81ECF315503D4CC6B5CE4BD06FF8BD" box="[691,826,1831,1855]" name="Madagascar" pageId="5" pageNumber="22">Madagascar</collectingCountry>
(project coordinated by B. Fisher), 
<collectingCountry id="FE81ECF315503D4CC0DECE4BD66EF8BD" box="[1240,1339,1831,1855]" name="Thailand" pageId="5" pageNumber="22">Thailand</collectingCountry>
(project coordinated by M. Sharkey) and SE Asia (project coordinated by C. van K. Achterberg). We were able to find 
<specimenCount id="909067EA15503D4CC186CE27D259F805" pageId="5" pageNumber="22" type="generic">71 specimens</specimenCount>
from 
<collectingCountry id="FE81ECF315503D4CC550CE1CD2F2F805" box="[342,423,1904,1927]" name="Yemen" pageId="5" pageNumber="22">Yemen</collectingCountry>
, 
<collectingCountry id="FE81ECF315503D4CC5B3CE03D2A7F805" box="[437,498,1903,1927]" name="United Arab Emirates" pageId="5" pageNumber="22">UAE</collectingCountry>
, 
<collectingCountry id="FE81ECF315503D4CC606CE03D136F805" box="[512,611,1903,1927]" name="Thailand" pageId="5" pageNumber="22">Thailand</collectingCountry>
and 
<collectingCountry id="FE81ECF315503D4CC6A6CE03D1AAF805" box="[672,767,1903,1927]" name="Vietnam" pageId="5" pageNumber="22">Vietnam</collectingCountry>
(
<figureCitation id="1EADB0E615503D4CC717CE03D033F805" box="[785,870,1903,1927]" captionStart="FIGURE 37" captionStartId="13.[151,250,1326,1349]" captionTargetBox="[154,1435,195,1304]" captionTargetId="figure@13.[151,1436,193,1305]" captionTargetPageId="13" captionText="FIGURE 37. Current known distribution of Epyris afer. Red circles represent the previous data and blue squares represent the new data of species." httpUri="https://zenodo.org/record/204631/files/figure.png" pageId="5" pageNumber="22">Fig. 37</figureCitation>
, blue squares). This confirms that this species is widely distributed throughout the Afrotropical and Oriental regions. However we were not able to find 
<taxonomicName id="4196D7E015503D4CC14ACEF8D6C9F829" box="[1356,1436,1939,1963]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="species" species="afer">
<emphasis id="B4E2707115503D4CC14ACEF8D6C9F829" box="[1356,1436,1939,1963]" italics="true" pageId="5" pageNumber="22">E. afer</emphasis>
</taxonomicName>
among the 2102 specimens of 
<taxonomicName id="4196D7E015503D4CC5FDCED4D110F84D" box="[507,581,1976,1999]" class="Insecta" family="Zygaenidae" genus="Epyris" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B4E2707115503D4CC5FDCED4D110F84D" box="[507,581,1976,1999]" italics="true" pageId="5" pageNumber="22">Epyris</emphasis>
</taxonomicName>
identified by 
<bibRefCitation id="E207D19215503D4CC6EECEDAD768F84C" author="Mugrabi" box="[744,1085,1974,1998]" pageId="5" pageNumber="22" refString="Mugrabi, D. F. &amp; Azevedo, C. O. (2010) Insecta, Hymenoptera, Bethylidae: range extension and filling gaps in Madagascar. Check List, 6, 62 - 63." type="journal article" year="2010">Mugrabi and Azevedo (2010)</bibRefCitation>
from 
<collectingCountry id="FE81ECF315503D4CC081CEDAD645F84C" box="[1159,1296,1974,1998]" name="Madagascar" pageId="5" pageNumber="22">Madagascar</collectingCountry>
. Also, there were not any specimens within material from 
<collectingCountry id="FE81ECF315503D4CC69FCEB6D053F870" box="[665,774,2010,2034]" name="Indonesia" pageId="5" pageNumber="22">Indonesia</collectingCountry>
and 
<collectingCountry id="FE81ECF315503D4CC73ACEB6D0F3F870" box="[828,934,2010,2034]" name="Malaysia" pageId="5" pageNumber="22">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>