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<document id="B2C4262C14C76DA4AB843AE0275780CC" ID-CLB-Dataset="29147" ID-DOI="10.11646/zootaxa.4422.2.5" ID-GBIF-Dataset="58bd0a34-478a-4c5d-b1c6-efb610eba18d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1455595" ID-ZooBank="2C026AEA-280E-444B-A70B-38D493EE2B8E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539185499276" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; Huang, Sunbin" docDate="2018" docId="0E2E934E064BAE1009B349B604CBFCD6" docLanguage="en" docName="zootaxa.4422.2.5.pdf" docOrigin="Zootaxa 4422 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aspidaphaenops dudou Tian &amp; Huang 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="258" masterDocId="F217EB360642AE1E09244C6B004BFFC3" masterDocTitle="Contribution to the knowledge of the cavernicolous beetle genus Aspidaphaenops Uéno from Guizhou (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="258" masterPageNumber="244" pageNumber="253" updateTime="1698573746751" updateUser="ExternalLinkService">
<mods:mods id="74D39B0F5EFEE52CE5D7F549C7640294" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="CB3D2934248F2C31E8471E3B1154F986">Contribution to the knowledge of the cavernicolous beetle genus Aspidaphaenops Uéno from Guizhou (Coleoptera: Carabidae: Trechinae)</mods:title>
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<mods:namePart id="8A1E09196254ED1E7DC9915C9234D942">Tian, Mingyi</mods:namePart>
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<mods:namePart id="8C3B4EDEC6CFF507404E388B86F7586E">Huang, Sunbin</mods:namePart>
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<mods:date id="3463AB84A4074CF10F9F4041D541E797">2018</mods:date>
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<mods:number id="E9DF4717F6E8B0E3F682307F2ED59344">4422</mods:number>
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<mods:classification id="8CC6640126A5C2C85473452AF3DE4F3A">journal article</mods:classification>
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<mods:identifier id="C8A7EA02C5D897782EAEA20FA056E46B" type="DOI">10.11646/zootaxa.4422.2.5</mods:identifier>
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<treatment id="0E2E934E064BAE1009B349B604CBFCD6" ID-DOI="http://doi.org/10.5281/zenodo.5984756" ID-GBIF-Taxon="148671586" ID-Zenodo-Dep="5984756" LSID="urn:lsid:plazi:treatment:0E2E934E064BAE1009B349B604CBFCD6" httpUri="http://treatment.plazi.org/id/0E2E934E064BAE1009B349B604CBFCD6" lastPageId="14" lastPageNumber="258" pageId="9" pageNumber="253">
<subSubSection id="CE9D71D3064BAE1709B349B60262F9DB" pageId="9" pageNumber="253" type="nomenclature">
<paragraph id="86382258064BAE1709B349B601B0FA34" blockId="9.[151,553,1501,1561]" box="[151,507,1501,1527]" pageId="9" pageNumber="253">
<heading id="DD709534064BAE1709B349B601B0FA34" bold="true" box="[151,507,1501,1527]" fontSize="11" level="1" pageId="9" pageNumber="253" reason="1">
<emphasis id="B4F3FE4A064BAE1709B349B601B0FA34" bold="true" box="[151,507,1501,1527]" pageId="9" pageNumber="253">
<taxonomicName id="418759DB064BAE1709B349B601F9FA34" ID-CoL="8M6TK" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[151,434,1501,1527]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="dudou" status="sp. nov.">
<emphasis id="B4F3FE4A064BAE1709B349B601F9FA34" bold="true" box="[151,434,1501,1527]" italics="true" pageId="9" pageNumber="253">Aspidaphaenops dudou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC04331064BAE17089D49B601B0FA34" box="[441,507,1501,1527]" pageId="9" pageNumber="253" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="86382258064BAE1709B34A6B0262F9DB" blockId="9.[151,553,1501,1561]" box="[151,553,1536,1561]" pageId="9" pageNumber="253">
<figureCitation id="1EBC3EDD064BAE1709B34A6B00BAF9DB" box="[151,241,1536,1561]" captionStart="FIGURE 1" captionStartId="2.[151,250,1507,1529]" captionTargetBox="[216,1364,259,1473]" captionTargetId="figure@2.[207,1375,257,1479]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Aspidaphaenops species (a. A. dudou n. sp.; b. A. volatidraconis; c. A. masakii; d. A. xiongda n. sp.; e. A. reflexus)." httpUri="https://zenodo.org/record/1455597/files/figure.png" pageId="9" pageNumber="253">Figs. 1a</figureCitation>
,
<figureCitation id="1EBC3EDD064BAE1709D94A6B016BF9DB" box="[253,288,1536,1560]" captionStart="FIGURE 2" captionStartId="3.[151,250,1515,1537]" captionTargetBox="[302,1297,213,1492]" captionTargetId="figure@3.[285,1309,195,1504]" captionTargetPageId="3" captionText="FIGURE 2. Heads of Aspidaphaenops species, dorsal view (A. A. masakii; B. A. volatidraconis; C. A. dudou n. sp.; D. A. reflexus; E. A. xiongda n. sp.)." httpUri="https://zenodo.org/record/1455601/files/figure.png" pageId="9" pageNumber="253">2C</figureCitation>
,
<figureCitation id="1EBC3EDD064BAE1708084A6A0104F9DB" box="[300,335,1537,1560]" captionStart="FIGURE 7" captionStartId="8.[151,250,1774,1796]" captionTargetBox="[192,1394,201,1753]" captionTargetId="figure@8.[188,1398,193,1753]" captionTargetPageId="8" captionText="FIGURE 7. Left elytron of Aspidaphaenops species (A. A. xiongda n. sp.; B. A. dudou n. sp.)." httpUri="https://zenodo.org/record/1455607/files/figure.png" pageId="9" pageNumber="253">7B</figureCitation>
,
<figureCitation id="1EBC3EDD064BAE17087F4A6B0132F9DB" box="[347,377,1536,1560]" captionStart="FIGURE 10" captionStartId="11.[151,250,1799,1821]" captionText="FIGURE 10. Habitus of Aspidaphaenops dudou n. sp., female, paratype." pageId="9" pageNumber="253">10</figureCitation>
,
<figureCitation id="1EBC3EDD064BAE1708A14A6B019CF9DB" box="[389,471,1536,1560]" captionStart="FIGURE 11" captionStartId="12.[151,250,1013,1035]" captionTargetBox="[340,1242,193,1000]" captionTargetId="figure@12.[336,1244,193,1005]" captionTargetPageId="12" captionText="FIGURE 11. Male genitalia of Aspidaphaenops species, median lobe and parameres (lateral view), and apex (dorsal view) (A, B. A. dudou n. sp. holotype; C, D. A. masakii)." httpUri="https://zenodo.org/record/1455613/files/figure.png" pageId="9" pageNumber="253">11A, B</figureCitation>
,
<figureCitation id="1EBC3EDD064BAE1708C74A6B024AF9DB" box="[483,513,1536,1560]" captionStart="FIGURE 12" captionStartId="12.[151,250,1982,2004]" captionTargetBox="[179,1419,1124,1966]" captionTargetId="figure@12.[168,1432,1116,1971]" captionTargetPageId="12" captionText="FIGURE 12. Map of cave Xiaozi Dong, type locality of Aspidaphaenops dudou n. sp. (courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society, triangles indicating where the beetles were collected)." httpUri="https://zenodo.org/record/1455615/files/figure.png" pageId="9" pageNumber="253">12</figureCitation>
,
<figureCitation id="1EBC3EDD064BAE170B294A6B0262F9DB" box="[525,553,1536,1560]" captionStart="FIGURE 13" captionStartId="13.[151,250,1966,1988]" captionTargetBox="[250,1336,296,1951]" captionTargetId="figure@13.[247,1339,293,1956]" captionTargetPageId="13" captionText="FIGURE 13. Cave Xiaozi Dong, type locality of Aspidaphaenops dudou n. sp. (A. entrance; B. main passage; C, D. active beetles in the cave)." httpUri="https://zenodo.org/record/1455617/files/figure.png" pageId="9" pageNumber="253">13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CE9D71D3064BAE1009B34A2C0249FEA7" lastPageId="14" lastPageNumber="258" pageId="9" pageNumber="253" type="materials_examined">
<paragraph id="86382258064BAE1709B34A2C02C6F96B" blockId="9.[151,1437,1607,2029]" pageId="9" pageNumber="253">
<materialsCitation id="36EF2805064BAE1709B34A2C00AEF96B" ID-GBIF-Occurrence="1915498333" collectingDate="2017-05-14" collectionCode="SCAU" collectorName="Mingyi Tian &amp; Weixin Liu &amp; Xiaozhu Luo &amp; Pingjing Yang &amp; Yanyi Pu" county="Xiaozi Dong" elevation="1764" latitude="25.112106" location="Sege" longLatPrecision="1" longitude="104.77572" municipality="Houshan" pageId="9" pageNumber="253" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="B4F3FE4A064BAE1709B34A2C014DF9A3" bold="true" box="[151,262,1607,1632]" pageId="9" pageNumber="253">Material.</emphasis>
<typeStatus id="593C9CFA064BAE17082A4A23013DF9A2" box="[270,374,1608,1633]" pageId="9" pageNumber="253" type="holotype">Holotype</typeStatus>
male,
<collectingCounty id="6F595AD4064BAE1708E74A23021CF9A2" box="[451,599,1608,1633]" pageId="9" pageNumber="253">Xiaozi Dong</collectingCounty>
,
<collectingMunicipality id="665CB822064BAE170B474A230282F9A2" box="[611,713,1608,1633]" pageId="9" pageNumber="253">Houshan</collectingMunicipality>
,
<location id="83587483064BAE170BF24A230344F9A3" LSID="urn:lsid:plazi:treatment:0E2E934E064BAE1009B349B604CBFCD6:83587483064BAE170BF24A230344F9A3" box="[726,783,1608,1632]" county="Xiaozi Dong" latitude="25.112106" longLatPrecision="1" longitude="104.77572" municipality="Houshan" name="Sege" pageId="9" pageNumber="253" stateProvince="Guizhou">Sege</location>
,
<location id="83587483064BAE170A3F4A230332F9A2" LSID="urn:lsid:plazi:treatment:0E2E934E064BAE1009B349B604CBFCD6:83587483064BAE170A3F4A230332F9A2" box="[795,889,1608,1633]" county="Xiaozi Dong" latitude="25.112106" longLatPrecision="1" longitude="104.77572" municipality="Houshan" name="Wushan" pageId="9" pageNumber="253" stateProvince="Guizhou">Wushan</location>
,
<location id="83587483064BAE170AA14A23039DF9A2" LSID="urn:lsid:plazi:treatment:0E2E934E064BAE1009B349B604CBFCD6:83587483064BAE170AA14A23039DF9A2" box="[901,982,1608,1633]" county="Xiaozi Dong" latitude="25.112106" longLatPrecision="1" longitude="104.77572" municipality="Houshan" name="Xingyi" pageId="9" pageNumber="253" stateProvince="Guizhou">Xingyi</location>
,
<location id="83587483064BAE170AC54A230412F9A2" LSID="urn:lsid:plazi:treatment:0E2E934E064BAE1009B349B604CBFCD6:83587483064BAE170AC54A230412F9A2" box="[993,1113,1608,1633]" county="Xiaozi Dong" latitude="25.112106" longLatPrecision="1" longitude="104.77572" municipality="Houshan" name="Qianxinan" pageId="9" pageNumber="253" stateProvince="Guizhou">Qianxinan</location>
,
<collectingRegion id="4443ECBA064BAE170D414A230482F9A2" box="[1125,1225,1608,1633]" country="China" name="Guizhou" pageId="9" pageNumber="253">Guizhou</collectingRegion>
,
<geoCoordinate id="E3B3449F064BAE170DF04A2305C7F9A3" box="[1236,1420,1608,1633]" direction="north" orientation="latitude" pageId="9" pageNumber="253" precision="1" value="25.112106">25°06’43.58’’ N</geoCoordinate>
/
<geoCoordinate id="E3B3449F064BAE1709B34A070110F947" box="[151,347,1644,1669]" direction="east" orientation="longitude" pageId="9" pageNumber="253" precision="1" value="104.77572">104°46’32.58’’ E</geoCoordinate>
,
<elevation id="0DAAC56B064BAE17084C4A0701F4F947" box="[360,447,1644,1669]" metricMagnitude="3" metricUnit="m" metricValue="1.764" pageId="9" pageNumber="253" unit="m" value="1764.0">
<quantity id="417F8FBD064BAE17084C4A0701F4F947" box="[360,447,1644,1669]" metricMagnitude="3" metricUnit="m" metricValue="1.764" pageId="9" pageNumber="253" unit="m" value="1764.0">1764 m</quantity>
</elevation>
,
<collectingDate id="E27DFD70064BAE1708E84A07020CF947" box="[460,583,1644,1668]" pageId="9" pageNumber="253" value="2017-05-14">
<date id="F2390498064BAE1708E84A07020CF947" box="[460,583,1644,1668]" pageId="9" pageNumber="253" value="2017-05-14">14-V-2017</date>
</collectingDate>
,
<collectorName id="2B72478E064BAE170B764A0702AAF946" box="[594,737,1644,1669]" pageId="9" pageNumber="253">Mingyi Tian</collectorName>
,
<collectorName id="2B72478E064BAE170BC94A070325F946" box="[749,878,1644,1669]" pageId="9" pageNumber="253">Weixin Liu</collectorName>
,
<collectorName id="2B72478E064BAE170A5E4A07045BF947" box="[890,1040,1644,1669]" pageId="9" pageNumber="253">Xiaozhu Luo</collectorName>
,
<collectorName id="2B72478E064BAE170D384A0704F7F946" box="[1052,1212,1644,1669]" pageId="9" pageNumber="253">Pingjing Yang</collectorName>
&amp;
<collectorName id="2B72478E064BAE170DC44A07050CF947" box="[1248,1351,1644,1669]" pageId="9" pageNumber="253">Yanyi Pu</collectorName>
leg., in
<collectionCode id="E096BA9D064BAE1709B34AFB00AEF96B" box="[151,229,1680,1704]" pageId="9" pageNumber="253">SCAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="36EF2805064BAE1709D44AFA02C1F96B" ID-GBIF-Occurrence="1915498335" box="[240,650,1680,1705]" collectingDate="2017-05-14" collectionCode="SCAU" collectorName="Mingyi Tian &amp; Weixin Liu &amp; Xiaozhu Luo &amp; Pingjing Yang &amp; Yanyi Pu" county="Xiaozi Dong" elevation="1764" latitude="25.112106" location="Sege" longLatPrecision="1" longitude="104.77572" municipality="Houshan" pageId="9" pageNumber="253" specimenCount="7" specimenCount-female="7" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="593C9CFA064BAE1709D44AFA012BF96A" box="[240,352,1681,1705]" pageId="9" pageNumber="253" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9081E9D1064BAE17084E4AFA0192F96A" box="[362,473,1680,1705]" pageId="9" pageNumber="253" type="female">7 females</specimenCount>
, ibid, in
<collectionCode id="E096BA9D064BAE170B1F4AFB02C1F96B" box="[571,650,1680,1704]" pageId="9" pageNumber="253">SCAU</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="86382258064BAE1709E34ADF03C6F90F" blockId="9.[151,1437,1607,2029]" box="[199,909,1716,1741]" pageId="9" pageNumber="253">
Length:
<quantity id="417F8FBD064BAE1708014ADF01E4F90F" box="[293,431,1716,1741]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.2" metricValueMin="5.9" pageId="9" pageNumber="253" unit="mm" value="6.05" valueMax="6.2" valueMin="5.9">5.9–6.2 mm</quantity>
; width:
<quantity id="417F8FBD064BAE170B2D4ADF02D8F90F" box="[521,659,1716,1741]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="9" pageNumber="253" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
. Habitus as in
<figureCitation id="1EBC3EDD064BAE170A134ADF03C2F90F" box="[823,905,1716,1741]" captionStart="FIGURE 10" captionStartId="11.[151,250,1799,1821]" captionText="FIGURE 10. Habitus of Aspidaphaenops dudou n. sp., female, paratype." pageId="9" pageNumber="253">Fig. 10</figureCitation>
.
</paragraph>
<paragraph id="86382258064BAE1709E34AB302F0F89F" blockId="9.[151,1437,1607,2029]" pageId="9" pageNumber="253">Body yellowish brown to brown, but pale on mouthparts palps and tarsi; moderately shiny and whole body including abdominal ventrites glabrous, legs pubescent; microsculptural engraved meshes moderately transverse on head, strongly transverse on pronotum and elytra; fore body (with mandibles) slightly shorter than elytra, (HmL+PrL) / EL = 0.97, (HlL+PrL) / EL = 0.79.</paragraph>
<paragraph id="86382258064BAE1409E34B030095FE48" blockId="9.[151,1437,1607,2029]" lastBlockId="10.[151,1436,151,395]" lastPageId="10" lastPageNumber="254" pageId="9" pageNumber="253">
Head elongated quadrate, much longer than wide, HLm/HW = 2.10–2.34, HLl/HW = 1.58–1.59; widest just behind sockets of antennae, at about 1/4 of head from labrum, genae gently and straightly narrowed posteriad; frons and vertex moderately convex, frontal furrows well-marked, deep and slightly divergent posteriad, ended at about middle of head from clypeus; clypeus transverse, quadrisetose; labrum transverse, frontal margin slightly emarginated, 6-setose (but 7-setose in a male
<typeStatus id="593C9CFA0648AE140BB64CF302B3FF73" box="[658,760,152,176]" pageId="10" pageNumber="254" type="paratype">paratype</typeStatus>
); anterior and posterior supraorbital setiferous pores at about middle and basal 2/11 of head from labrum respectively; mentum and submentum completely fused, mentum bisetose on either side of tooth which is simple and blunt at apex, basal fovea widely concave; submentum 8- setose; both maxillary and labial palps thin and subcylindrical, glabrous except for the 2nd labial palpomere which is bisetose on inner margin; the penultimate labial and maxillary palpomere about 1.3 times longer than the last one respectively; suborbital pore closer to base than to submentum. Antennae thin and long, extending over apices of elytra.
</paragraph>
<caption id="D2F872D00648AE1409B34B3702F4F82F" httpUri="https://zenodo.org/record/1455611/files/figure.png" pageId="10" pageNumber="254" startId="10.[151,250,1884,1906]" targetBox="[280,1313,437,1864]" targetPageId="10">
<paragraph id="863822580648AE1409B34B3702F4F82F" blockId="10.[151,1436,1884,2028]" pageId="10" pageNumber="254">
<emphasis id="B4F3FE4A0648AE1409B34B370158F8B2" bold="true" box="[151,275,1884,1906]" pageId="10" pageNumber="254">FIGURE 9.</emphasis>
Cave Fengyan Dong, type locality of
<taxonomicName id="418759DB0648AE140BBE4B3703D1F8B2" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[666,922,1884,1905]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="254" phylum="Arthropoda" rank="species" species="xiongda" status="sp. nov.">
<emphasis id="B4F3FE4A0648AE140BBE4B3703D1F8B2" box="[666,922,1884,1905]" italics="true" pageId="10" pageNumber="254">Aspidaphaenops xiongda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC043310648AE140A854B370390F8B2" box="[929,987,1884,1905]" pageId="10" pageNumber="254" rank="species">
<emphasis id="B4F3FE4A0648AE140A854B370390F8B2" bold="true" box="[929,987,1884,1905]" pageId="10" pageNumber="254">n. sp.</emphasis>
</taxonomicNameLabel>
(A. entrance; B. main passage; C, D. active beetles in the cave).Prothorax not dilated, propleura invisible from above. Pronotum barrel-like, longer than wide, PnL/PnW = 1.19–1.20, widest near the middle of pronotum, strongly recurved before hind angles which are not well-defined, fore angles obtuse; base as wide as front; latero-margins gradually widened and reflexed from middle to base, fore latero-marginal setae at about 1/5 anterior of pronotum. Scutellum fairly large.
</paragraph>
</caption>
<caption id="D2F872D00649AE1509B34B6C0334F8DE" box="[151,895,1799,1821]" pageId="11" pageNumber="255">
<paragraph id="863822580649AE1509B34B6C0334F8DE" blockId="11.[151,895,1799,1821]" box="[151,895,1799,1821]" pageId="11" pageNumber="255">
<emphasis id="B4F3FE4A0649AE1509B34B6C0154F8DF" bold="true" box="[151,287,1799,1821]" pageId="11" pageNumber="255">FIGURE 10.</emphasis>
Habitus of
<taxonomicName id="418759DB0649AE1508B24B6C02C8F8DF" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[406,643,1799,1820]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="255" phylum="Arthropoda" rank="species" species="dudou" status="sp. nov.">
<emphasis id="B4F3FE4A0649AE1508B24B6C02C8F8DF" box="[406,643,1799,1820]" italics="true" pageId="11" pageNumber="255">Aspidaphaenops dudou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC043310649AE150BAD4B6C0289F8DF" box="[649,706,1799,1820]" pageId="11" pageNumber="255" rank="species">
<emphasis id="B4F3FE4A0649AE150BAD4B6C0289F8DF" bold="true" box="[649,706,1799,1820]" pageId="11" pageNumber="255">n. sp.</emphasis>
</taxonomicNameLabel>
, female, paratype.
</paragraph>
</caption>
<paragraph id="863822580649AE1509E34B2F053EF830" blockId="11.[151,1436,1860,2035]" pageId="11" pageNumber="255">
Elytra elongated ovate (
<figureCitation id="1EBC3EDD0649AE1508FF4B2F027EF89E" box="[475,565,1860,1885]" captionStart="FIGURE 7" captionStartId="8.[151,250,1774,1796]" captionTargetBox="[192,1394,201,1753]" captionTargetId="figure@8.[188,1398,193,1753]" captionTargetPageId="8" captionText="FIGURE 7. Left elytron of Aspidaphaenops species (A. A. xiongda n. sp.; B. A. dudou n. sp.)." httpUri="https://zenodo.org/record/1455607/files/figure.png" pageId="11" pageNumber="255">Fig. 7B</figureCitation>
), much wider than wide, EL/EW = 1.55–1.58; much wider than pronotum, EW/PnW = 1.88–1.91; widest near the middle, lateral sides finely ciliate throughout, marginal gutters wellmarked; punctures of striae visible; sites of anterior dorsal pores variable, at the portions from basal 1/3 of elytra to middle; middle pores at basal 4/7 of elytra; preapical pores closer to elytral suture than to elytral apices; the 1st to 3rd pores of the marginal umbilicate series equidistant, 4th backwardly isolated, 5th and 6th pores close to each other.
</paragraph>
<caption id="D2F872D0064EAE1209B34F9E0239FBE9" httpUri="https://zenodo.org/record/1455613/files/figure.png" pageId="12" pageNumber="256" startId="12.[151,250,1013,1035]" targetBox="[340,1242,193,1000]" targetPageId="12">
<paragraph id="86382258064EAE1209B34F9E0239FBE9" blockId="12.[151,1435,1013,1066]" pageId="12" pageNumber="256">
<emphasis id="B4F3FE4A064EAE1209B34F9E0155FBC9" bold="true" box="[151,286,1013,1035]" pageId="12" pageNumber="256">FIGURE 11.</emphasis>
Male genitalia of
<taxonomicName id="418759DB064EAE1208FC4F9E0234FBC9" authorityName="Ueno" authorityYear="2006" box="[472,639,1013,1034]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="B4F3FE4A064EAE1208FC4F9E0234FBC9" box="[472,639,1013,1034]" italics="true" pageId="12" pageNumber="256">Aspidaphaenops</emphasis>
</taxonomicName>
species, median lobe and parameres (lateral view), and apex (dorsal view) (A, B.
<taxonomicName id="418759DB064EAE120990487F015BFBEA" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[180,272,1044,1065]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="256" phylum="Arthropoda" rank="species" species="dudou" status="sp. nov.">
<emphasis id="B4F3FE4A064EAE120990487F015BFBEA" box="[180,272,1044,1065]" italics="true" pageId="12" pageNumber="256">A. dudou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC04331064EAE120832487F0104FBEA" box="[278,335,1044,1065]" pageId="12" pageNumber="256" rank="species">
<emphasis id="B4F3FE4A064EAE120832487F0104FBEA" bold="true" box="[278,335,1044,1065]" pageId="12" pageNumber="256">n. sp.</emphasis>
</taxonomicNameLabel>
holotype; C, D.
<taxonomicName id="418759DB064EAE1208DC487F0228FBEA" authorityName="Ueno" authorityYear="2006" box="[504,611,1044,1065]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="256" phylum="Arthropoda" rank="species" species="masakii">
<emphasis id="B4F3FE4A064EAE1208DC487F0228FBEA" box="[504,611,1044,1065]" italics="true" pageId="12" pageNumber="256">A. masakii</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<caption id="D2F872D0064EAE1209B34BD5041DF830" httpUri="https://zenodo.org/record/1455615/files/figure.png" pageId="12" pageNumber="256" startId="12.[151,250,1982,2004]" targetBox="[179,1419,1124,1966]" targetPageId="12">
<paragraph id="86382258064EAE1209B34BD5041DF830" blockId="12.[151,1436,1982,2035]" pageId="12" pageNumber="256">
<emphasis id="B4F3FE4A064EAE1209B34BD5016BF817" bold="true" box="[151,288,1982,2004]" pageId="12" pageNumber="256">FIGURE 12.</emphasis>
Map of cave Xiaozi Dong, type locality of
<taxonomicName id="418759DB064EAE120BC14BD4039FF817" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[741,980,1983,2004]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="256" phylum="Arthropoda" rank="species" species="dudou" status="sp. nov.">
<emphasis id="B4F3FE4A064EAE120BC14BD4039FF817" box="[741,980,1983,2004]" italics="true" pageId="12" pageNumber="256">Aspidaphaenops dudou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC04331064EAE120AFF4BD4045DF817" box="[987,1046,1983,2004]" pageId="12" pageNumber="256" rank="species">
<emphasis id="B4F3FE4A064EAE120AFF4BD4045DF817" bold="true" box="[987,1046,1983,2004]" pageId="12" pageNumber="256">n. sp.</emphasis>
</taxonomicNameLabel>
(courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society, triangles indicating where the beetles were collected).
</paragraph>
</caption>
<paragraph id="86382258064FAE1309E34CF305D7FF73" blockId="13.[151,1436,151,251]" box="[199,1436,151,176]" pageId="13" pageNumber="257">Legs thin and long, bearing short pubescence; fore tibia smooth, absence of a longitudinal furrow or sulcus; the</paragraph>
<paragraph id="86382258064FAE1309B34CD40495FF14" blockId="13.[151,1436,151,251]" box="[151,1246,187,215]" pageId="13" pageNumber="257">
1
<taxonomicName id="418759DB064FAE1309814CD0016FFF14" box="[165,292,187,215]" pageId="13" pageNumber="257" rank="subSpecies" subSpecies="tarsomere">st tarsomere</taxonomicName>
as long as the 2nd –4th tarsomeres together in fore, middle and hind legs, respectively.
</paragraph>
<paragraph id="86382258064FAE1309E34C890531FF38" blockId="13.[151,1436,151,251]" box="[199,1402,226,251]" pageId="13" pageNumber="257">Ventrites IV–VI each with a pair of paramedian setae; ventrite VII bisetose in male, quadrisetose in female.</paragraph>
<caption id="D2F872D0064FAE1309B34BC5012AF821" httpUri="https://zenodo.org/record/1455617/files/figure.png" pageId="13" pageNumber="257" startId="13.[151,250,1966,1988]" targetBox="[250,1336,296,1951]" targetPageId="13">
<paragraph id="86382258064FAE1309B34BC5012AF821" blockId="13.[151,1435,1966,2018]" pageId="13" pageNumber="257">
<emphasis id="B4F3FE4A064FAE1309B34BC5016AF800" bold="true" box="[151,289,1966,1988]" pageId="13" pageNumber="257">FIGURE 13.</emphasis>
Cave Xiaozi Dong, type locality of
<taxonomicName id="418759DB064FAE130BBA4BC503C7F800" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[670,908,1966,1987]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="257" phylum="Arthropoda" rank="species" species="dudou" status="sp. nov.">
<emphasis id="B4F3FE4A064FAE130BBA4BC503C7F800" box="[670,908,1966,1987]" italics="true" pageId="13" pageNumber="257">Aspidaphaenops dudou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC04331064FAE130AB14BC5039BF800" box="[917,976,1966,1987]" pageId="13" pageNumber="257" rank="species">
<emphasis id="B4F3FE4A064FAE130AB14BC5039BF800" bold="true" box="[917,976,1966,1987]" pageId="13" pageNumber="257">n. sp.</emphasis>
</taxonomicNameLabel>
(A. entrance; B. main passage; C, D. active beetles in the cave).
</paragraph>
</caption>
<paragraph id="86382258064CAE1009E34CFC0249FEA7" blockId="14.[151,1437,151,789]" pageId="14" pageNumber="258">
Male genitalia (
<figureCitation id="1EBC3EDD064CAE1008524CFC024CFF73" box="[374,519,151,176]" captionStart="FIGURE 11" captionStartId="12.[151,250,1013,1035]" captionTargetBox="[340,1242,193,1000]" captionTargetId="figure@12.[336,1244,193,1005]" captionTargetPageId="12" captionText="FIGURE 11. Male genitalia of Aspidaphaenops species, median lobe and parameres (lateral view), and apex (dorsal view) (A, B. A. dudou n. sp. holotype; C, D. A. masakii)." httpUri="https://zenodo.org/record/1455613/files/figure.png" pageId="14" pageNumber="258">Figs. 11A, B</figureCitation>
): Moderately sclerotized, thin and slender, similar in
<taxonomicName id="418759DB064CAE100D734CF3051EFF73" authority="Ueno, 2006" authorityName="Ueno" authorityYear="2006" box="[1111,1365,151,176]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="masakii">
<emphasis id="B4F3FE4A064CAE100D734CF30487FF73" box="[1111,1228,151,176]" italics="true" pageId="14" pageNumber="258">A. masakii</emphasis>
Uéno, 2006
</taxonomicName>
(
<figureCitation id="1EBC3EDD064CAE100C404CFC00A4FF17" captionStart="FIGURE 11" captionStartId="12.[151,250,1013,1035]" captionTargetBox="[340,1242,193,1000]" captionTargetId="figure@12.[336,1244,193,1005]" captionTargetPageId="12" captionText="FIGURE 11. Male genitalia of Aspidaphaenops species, median lobe and parameres (lateral view), and apex (dorsal view) (A, B. A. dudou n. sp. holotype; C, D. A. masakii)." httpUri="https://zenodo.org/record/1455613/files/figure.png" pageId="14" pageNumber="258">Figs. 11C, D</figureCitation>
). Median lobe moderately arcuate ventrally at about basal 1/4, apical lobe thinner and a little longer, slightly recurved and reflexed at apex; base wide, with a very large sagittal aileron; inner sac of the median lobe with an anisotopic and long copulatory piece which is about 1/4 as long as the median lobe; in dorsal view, apex broad, slightly expanded at sides, roundly blunt at apex; parameters fairly large, about 1/3 as long as median lobe, each with four long apical setae.
</paragraph>
</subSubSection>
<subSubSection id="CE9D71D3064CAE1009E34D1B04AAFDDB" pageId="14" pageNumber="258" type="discussion">
<paragraph id="86382258064CAE1009E34D1B04AAFDDB" blockId="14.[151,1437,151,789]" pageId="14" pageNumber="258">
<emphasis id="B4F3FE4A064CAE1009E34D1B0170FE4B" bold="true" box="[199,315,368,392]" pageId="14" pageNumber="258">Remarks.</emphasis>
<taxonomicName id="418759DB064CAE1008614D040205FE4B" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[325,590,367,392]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="dudou" status="sp. nov.">
<emphasis id="B4F3FE4A064CAE1008614D040205FE4B" box="[325,590,367,392]" italics="true" pageId="14" pageNumber="258">Aspidaphaenops dudou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC04331064CAE100B7C4D1B02D0FE4B" box="[600,667,368,392]" pageId="14" pageNumber="258" rank="species">
<emphasis id="B4F3FE4A064CAE100B7C4D1B02D0FE4B" bold="true" box="[600,667,368,392]" pageId="14" pageNumber="258">n. sp.</emphasis>
</taxonomicNameLabel>
is close to
<taxonomicName id="418759DB064CAE100A024D040550FE4B" authority="Ueno, 2006" authorityName="Ueno" authorityYear="2006" box="[806,1307,367,392]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="volatidraconis">
<emphasis id="B4F3FE4A064CAE100A024D0404C0FE4B" box="[806,1163,367,392]" italics="true" pageId="14" pageNumber="258">Aspidaphaenops volatidraconis</emphasis>
Uéno, 2006
</taxonomicName>
but differs from it by: (1) head thinner, sides straightly narrowed posteriad (vs. thicker, sides slightly dilated in
<taxonomicName id="418759DB064CAE100CA04DFF0172FE13" authorityName="Ueno" authorityYear="2006" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="volatidraconis">
<emphasis id="B4F3FE4A064CAE100CA04DFF0172FE13" italics="true" pageId="14" pageNumber="258">A. volatidraconis</emphasis>
</taxonomicName>
); (2) fore angles of pronotum sharp (vs. obtuse in
<taxonomicName id="418759DB064CAE100A584DD30474FE13" authorityName="Ueno" authorityYear="2006" box="[892,1087,439,464]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="volatidraconis">
<emphasis id="B4F3FE4A064CAE100A584DD30474FE13" box="[892,1087,439,464]" italics="true" pageId="14" pageNumber="258">A. volatidraconis</emphasis>
</taxonomicName>
); and (3) median lobe of male genitalia slenderer, with a larger sagittal aileron and longer apical lobe in lateral view (vs. median lobe shorter, with a smaller sagittal aileron and shorter apical lobe in
<taxonomicName id="418759DB064CAE100BF64E6B03D8FDDB" authorityName="Ueno" authorityYear="2006" box="[722,915,511,536]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="volatidraconis">
<emphasis id="B4F3FE4A064CAE100BF64E6B03D8FDDB" box="[722,915,511,536]" italics="true" pageId="14" pageNumber="258">A. volatidraconis</emphasis>
</taxonomicName>
) (Uéno, 2006, see
<figureCitation id="1EBC3EDD064CAE100D4C4D94049FFDDB" box="[1128,1236,511,536]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="8.[151,250,1774,1796]" captionStartId-1="9.[151,250,1379,1401]" captionTargetBox-0="[192,1394,201,1753]" captionTargetBox-1="[352,1235,744,1348]" captionTargetId-0="figure@8.[188,1398,193,1753]" captionTargetId-1="figure@9.[340,1247,735,1356]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 7. Left elytron of Aspidaphaenops species (A. A. xiongda n. sp.; B. A. dudou n. sp.)." captionText-1="FIGURE 8. Map of cave Fengyan Dong, type locality of Aspidaphaenops xiongda n. sp. (courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society, triangles indicating where the beetles were collected)." httpUri-0="https://zenodo.org/record/1455607/files/figure.png" httpUri-1="https://zenodo.org/record/1455609/files/figure.png" pageId="14" pageNumber="258">Figs. 7–8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CE9D71D3064CAE1009E34E4F02F6FDA3" pageId="14" pageNumber="258" type="etymology">
<paragraph id="86382258064CAE1009E34E4F02F6FDA3" blockId="14.[151,1437,151,789]" pageId="14" pageNumber="258">
<emphasis id="B4F3FE4A064CAE1009E34E4F0105FDFF" bold="true" box="[199,334,548,572]" pageId="14" pageNumber="258">Etymology.</emphasis>
“Dudou” is a nickname of Mr. Hongbo Yang, the head of Jinzhou Cave Exploration Club in Xingyi, to thank his assistance during the survey.
</paragraph>
</subSubSection>
<subSubSection id="CE9D71D3064CAE1009E34E0004CBFCD6" pageId="14" pageNumber="258" type="distribution">
<paragraph id="86382258064CAE1009E34E0004E8FD47" blockId="14.[151,1437,151,789]" box="[199,1187,619,645]" pageId="14" pageNumber="258">
<emphasis id="B4F3FE4A064CAE1009E34E00012BFD47" bold="true" box="[199,352,619,644]" pageId="14" pageNumber="258">Distribution.</emphasis>
<collectingRegion id="4443ECBA064CAE1008434E07018CFD46" box="[359,455,620,645]" country="China" name="Guizhou" pageId="14" pageNumber="258">Guizhou</collectingRegion>
(Xingyi). Known only from the cave Xiaozi Dong (
<figureCitation id="1EBC3EDD064CAE100D2B4E070421FD47" box="[1039,1130,620,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1507,1529]" captionTargetBox="[216,1364,259,1473]" captionTargetId="figure@2.[207,1375,257,1479]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Aspidaphaenops species (a. A. dudou n. sp.; b. A. volatidraconis; c. A. masakii; d. A. xiongda n. sp.; e. A. reflexus)." httpUri="https://zenodo.org/record/1455597/files/figure.png" pageId="14" pageNumber="258">Figs. 1a</figureCitation>
,
<figureCitation id="1EBC3EDD064CAE100D534E0704DDFD47" box="[1143,1174,620,644]" captionStart="FIGURE 12" captionStartId="12.[151,250,1982,2004]" captionTargetBox="[179,1419,1124,1966]" captionTargetId="figure@12.[168,1432,1116,1971]" captionTargetPageId="12" captionText="FIGURE 12. Map of cave Xiaozi Dong, type locality of Aspidaphaenops dudou n. sp. (courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society, triangles indicating where the beetles were collected)." httpUri="https://zenodo.org/record/1455615/files/figure.png" pageId="14" pageNumber="258">12</figureCitation>
).
</paragraph>
<paragraph id="86382258064CAE1009E34EE404CBFCD6" blockId="14.[151,1437,151,789]" pageId="14" pageNumber="258">
Xiaozi Dong (
<figureCitation id="1EBC3EDD064CAE1008434EE401B3FD6B" box="[359,504,655,680]" captionStart="FIGURE 13" captionStartId="13.[151,250,1966,1988]" captionTargetBox="[250,1336,296,1951]" captionTargetId="figure@13.[247,1339,293,1956]" captionTargetPageId="13" captionText="FIGURE 13. Cave Xiaozi Dong, type locality of Aspidaphaenops dudou n. sp. (A. entrance; B. main passage; C, D. active beetles in the cave)." httpUri="https://zenodo.org/record/1455617/files/figure.png" pageId="14" pageNumber="258">Figs. 13A, B</figureCitation>
) opens at mid hill near village Sege, with a small entrance which is surrounded by bushes. It is about
<quantity id="417F8FBD064CAE1008564EDF01F7FD0F" box="[370,444,692,716]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="258" unit="m" value="200.0">200 m</quantity>
long, with a unique passage. There are several small chambers inside the cave, and a small pool at the end point. The blind beetles (
<figureCitation id="1EBC3EDD064CAE100BBA4EBC0364FD33" box="[670,815,727,752]" captionStart="FIGURE 13" captionStartId="13.[151,250,1966,1988]" captionTargetBox="[250,1336,296,1951]" captionTargetId="figure@13.[247,1339,293,1956]" captionTargetPageId="13" captionText="FIGURE 13. Cave Xiaozi Dong, type locality of Aspidaphaenops dudou n. sp. (A. entrance; B. main passage; C, D. active beetles in the cave)." httpUri="https://zenodo.org/record/1455617/files/figure.png" pageId="14" pageNumber="258">Figs. 13C, D</figureCitation>
) were collected in the wet area between the second and third chambers, sympatric with ant-loving beetles, rove beetles, millipedes and diplurans.
</paragraph>
</subSubSection>
</treatment>
</document>
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