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<document id="06B0889B2277C929126B8EDE71B91507" ID-CLB-Dataset="29147" ID-DOI="10.11646/zootaxa.4422.2.5" ID-GBIF-Dataset="58bd0a34-478a-4c5d-b1c6-efb610eba18d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1455595" ID-ZooBank="2C026AEA-280E-444B-A70B-38D493EE2B8E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539185499276" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; Huang, Sunbin" docDate="2018" docId="0E2E934E0641AE1709B34A1B03C6FD6B" docLanguage="en" docName="zootaxa.4422.2.5.pdf" docOrigin="Zootaxa 4422 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aspidaphaenops xiongda Tian &amp; Huang 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="253" masterDocId="F217EB360642AE1E09244C6B004BFFC3" masterDocTitle="Contribution to the knowledge of the cavernicolous beetle genus Aspidaphaenops Uéno from Guizhou (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="258" masterPageNumber="244" pageNumber="247" updateTime="1698573746751" updateUser="ExternalLinkService">
<mods:mods id="0B58D4E6E87AEA1C8BBACE381A9B6E11" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F910FEDAEBDB8BDC95BECA0C179874C0">Contribution to the knowledge of the cavernicolous beetle genus Aspidaphaenops Uéno from Guizhou (Coleoptera: Carabidae: Trechinae)</mods:title>
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<mods:name id="8BF8046D21667F7C5087DBE3D602B17E" type="personal">
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<mods:namePart id="E50EB92DA4E28EC390B588F6570C3C21">Tian, Mingyi</mods:namePart>
</mods:name>
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<mods:namePart id="8778ADD7F4280B6FCD808E93E6640C7D">Huang, Sunbin</mods:namePart>
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<mods:title id="4CB79284CF55EBCD1E3800AD21AA1827">Zootaxa</mods:title>
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<mods:part id="16DE67594BF74D343CA97ABA4CBAC743">
<mods:date id="47F85FB5A0B48E7CF1D5E722EA0EE10A">2018</mods:date>
<mods:detail id="805572611DCEB1A220183CB2975A0E06" type="volume">
<mods:number id="804708B8D0F7A26B47DCC70795996E3F">4422</mods:number>
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<mods:classification id="C08F4C07E1D213C803134146BA06CB8A">journal article</mods:classification>
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<mods:identifier id="171C26AA480DA66F0FB086012D3FE3B9" type="DOI">10.11646/zootaxa.4422.2.5</mods:identifier>
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<treatment id="0E2E934E0641AE1709B34A1B03C6FD6B" ID-DOI="http://doi.org/10.5281/zenodo.5984754" ID-GBIF-Taxon="148671588" ID-Zenodo-Dep="5984754" LSID="urn:lsid:plazi:treatment:0E2E934E0641AE1709B34A1B03C6FD6B" httpUri="http://treatment.plazi.org/id/0E2E934E0641AE1709B34A1B03C6FD6B" lastPageId="9" lastPageNumber="253" pageId="3" pageNumber="247">
<subSubSection id="CE9D71D30641AE1D09B34A1B01B8F96F" pageId="3" pageNumber="247" type="nomenclature">
<paragraph id="863822580641AE1D09B34A1B025BF948" blockId="3.[151,528,1648,1708]" box="[151,528,1648,1675]" pageId="3" pageNumber="247">
<heading id="DD7095340641AE1D09B34A1B025BF948" bold="true" box="[151,528,1648,1675]" fontSize="11" level="1" pageId="3" pageNumber="247" reason="1">
<emphasis id="B4F3FE4A0641AE1D09B34A1B025BF948" bold="true" box="[151,528,1648,1675]" pageId="3" pageNumber="247">
<taxonomicName id="418759DB0641AE1D09B34A1B018CF949" ID-CoL="8MH7G" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[151,455,1648,1674]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="247" phylum="Arthropoda" rank="species" species="xiongda" status="sp. nov.">
<emphasis id="B4F3FE4A0641AE1D09B34A1B018CF949" bold="true" box="[151,455,1648,1674]" italics="true" pageId="3" pageNumber="247">Aspidaphaenops xiongda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC043310641AE1D08EA4A1A025BF948" box="[462,528,1649,1675]" pageId="3" pageNumber="247" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="863822580641AE1D09B34AF801B8F96F" blockId="3.[151,528,1648,1708]" box="[151,499,1683,1708]" pageId="3" pageNumber="247">
<figureCitation id="1EBC3EDD0641AE1D09B34AF800B9F96F" box="[151,242,1683,1708]" captionStart="FIGURE 1" captionStartId="2.[151,250,1507,1529]" captionTargetBox="[216,1364,259,1473]" captionTargetId="figure@2.[207,1375,257,1479]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Aspidaphaenops species (a. A. dudou n. sp.; b. A. volatidraconis; c. A. masakii; d. A. xiongda n. sp.; e. A. reflexus)." httpUri="https://zenodo.org/record/1455597/files/figure.png" pageId="3" pageNumber="247">Figs. 1d</figureCitation>
,
<figureCitation id="1EBC3EDD0641AE1D09DB4AFF016AF96F" box="[255,289,1684,1708]" captionStart="FIGURE 2" captionStartId="3.[151,250,1515,1537]" captionTargetBox="[302,1297,213,1492]" captionTargetId="figure@3.[285,1309,195,1504]" captionTargetPageId="3" captionText="FIGURE 2. Heads of Aspidaphaenops species, dorsal view (A. A. masakii; B. A. volatidraconis; C. A. dudou n. sp.; D. A. reflexus; E. A. xiongda n. sp.)." httpUri="https://zenodo.org/record/1455601/files/figure.png" pageId="3" pageNumber="247">2E</figureCitation>
,
<figureCitation id="1EBC3EDD0641AE1D08084AFF0139F968" box="[300,370,1684,1708]" captionStart="FIGURE 4" captionStartId="5.[151,250,1400,1422]" captionTargetBox="[163,1411,196,1349]" captionTargetId="figure@5.[163,1427,193,1387]" captionTargetPageId="5" captionText="FIGURE 4. Male genitalia of Aspidaphaenops species, median lobe and parameres (lateral view), and apex (dorsal view) (A, B. A. reflexus; C, D. A. xiongda n. sp. holotype)." httpUri="https://zenodo.org/record/1455603/files/figure.png" pageId="3" pageNumber="247">4C, D</figureCitation>
,
<figureCitation id="1EBC3EDD0641AE1D08594AF801C6F96F" box="[381,397,1683,1708]" captionStart="FIGURE 6" captionStartId="7.[151,250,2014,2036]" captionText="FIGURE 6. Habitus of Aspidaphaenops xiongda n. sp., male, paratype." pageId="3" pageNumber="247">6</figureCitation>
,
<figureCitation id="1EBC3EDD0641AE1D08BD4AFF01F5F96F" box="[409,446,1684,1708]" captionStart="FIGURE 7" captionStartId="8.[151,250,1774,1796]" captionTargetBox="[192,1394,201,1753]" captionTargetId="figure@8.[188,1398,193,1753]" captionTargetPageId="8" captionText="FIGURE 7. Left elytron of Aspidaphaenops species (A. A. xiongda n. sp.; B. A. dudou n. sp.)." httpUri="https://zenodo.org/record/1455607/files/figure.png" pageId="3" pageNumber="247">7A</figureCitation>
,
<figureCitation id="1EBC3EDD0641AE1D08ED4AFF0193F96F" box="[457,472,1684,1708]" captionStart="FIGURE 8" captionStartId="9.[151,250,1379,1401]" captionTargetBox="[352,1235,744,1348]" captionTargetId="figure@9.[340,1247,735,1356]" captionTargetPageId="9" captionText="FIGURE 8. Map of cave Fengyan Dong, type locality of Aspidaphaenops xiongda n. sp. (courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society, triangles indicating where the beetles were collected)." httpUri="https://zenodo.org/record/1455609/files/figure.png" pageId="3" pageNumber="247">8</figureCitation>
,
<figureCitation id="1EBC3EDD0641AE1D08C14AFF01B8F96F" box="[485,499,1684,1708]" captionStart="FIGURE 9" captionStartId="10.[151,250,1884,1906]" captionTargetBox="[280,1313,437,1864]" captionTargetId="figure@10.[278,1315,434,1866]" captionTargetPageId="10" captionText="FIGURE 9. Cave Fengyan Dong, type locality of Aspidaphaenops xiongda n. sp. (A. entrance; B. main passage; C, D. active beetles in the cave).Prothorax not dilated, propleura invisible from above. Pronotum barrel-like, longer than wide, PnL/PnW = 1.19–1.20, widest near the middle of pronotum, strongly recurved before hind angles which are not well-defined, fore angles obtuse; base as wide as front; latero-margins gradually widened and reflexed from middle to base, fore latero-marginal setae at about 1/5 anterior of pronotum. Scutellum fairly large." httpUri="https://zenodo.org/record/1455611/files/figure.png" pageId="3" pageNumber="247">9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CE9D71D30641AE1709B34AB001D5FF3B" lastPageId="9" lastPageNumber="253" pageId="3" pageNumber="247" type="materials_examined">
<paragraph id="863822580641AE1D09B34AB002DFF8FF" blockId="3.[151,1436,1755,2032]" pageId="3" pageNumber="247">
<bibRef id="5F27B6AE0641AE1D09B34AB002DFF8FF" pageId="3" pageNumber="247" refId="ref1740" type="journal article">
<materialsCitation id="36EF28050641AE1D09B34AB00149F8FF" ID-GBIF-Occurrence="1915498334" collectingDate="2017-05-12" collectionCode="SCAU" collectorName="Mingyi Tian &amp; Weixin Liu &amp; Xiaozhu Luo &amp; Pingjing Yang &amp; Yanyi Pu" county="Fengyan Dong" elevation="1436" latitude="25.168236" location="Longguang" longLatPrecision="1" longitude="105.23075" municipality="Renjia Zhai" pageId="3" pageNumber="247" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="B4F3FE4A0641AE1D09B34AB0014DF937" bold="true" box="[151,262,1755,1780]" pageId="3" pageNumber="247">
<title id="F483C96E0641AE1D09B34AB00149F937" box="[151,258,1755,1780]" jopScore="0.1423801" pageId="3" pageNumber="247">Material</title>
.
</emphasis>
<journalOrPublisher id="5C4628940641AE1D082B4AB000A9F8DB" jopScore="2.566658" pageId="3" pageNumber="247">
<typeStatus id="593C9CFA0641AE1D082B4AB0013CF937" box="[271,375,1755,1780]" pageId="3" pageNumber="247" type="holotype">Holotype</typeStatus>
male,
<collectingCounty id="6F595AD40641AE1D08E34AB70239F937" box="[455,626,1756,1780]" pageId="3" pageNumber="247">Fengyan Dong</collectingCounty>
,
<collectingMunicipality id="665CB8220641AE1D0BA44AB00343F937" box="[640,776,1755,1780]" pageId="3" pageNumber="247">Renjia Zhai</collectingMunicipality>
,
<location id="835874830641AE1D0A324AB703D3F937" LSID="urn:lsid:plazi:treatment:0E2E934E0641AE1709B34A1B03C6FD6B:835874830641AE1D0A324AB703D3F937" box="[790,920,1756,1780]" county="Fengyan Dong" latitude="25.168236" longLatPrecision="1" longitude="105.23075" municipality="Renjia Zhai" name="Longguang" pageId="3" pageNumber="247" stateProvince="Guizhou">Longguang</location>
,
<location id="835874830641AE1D0A824AB003B6F937" LSID="urn:lsid:plazi:treatment:0E2E934E0641AE1709B34A1B03C6FD6B:835874830641AE1D0A824AB003B6F937" box="[934,1021,1755,1780]" county="Fengyan Dong" latitude="25.168236" longLatPrecision="1" longitude="105.23075" municipality="Renjia Zhai" name="Anlong" pageId="3" pageNumber="247" stateProvince="Guizhou">Anlong</location>
,
<location id="835874830641AE1D0D2E4AB004CAF937" LSID="urn:lsid:plazi:treatment:0E2E934E0641AE1709B34A1B03C6FD6B:835874830641AE1D0D2E4AB004CAF937" box="[1034,1153,1755,1780]" county="Fengyan Dong" latitude="25.168236" longLatPrecision="1" longitude="105.23075" municipality="Renjia Zhai" name="Qianxinan" pageId="3" pageNumber="247" stateProvince="Guizhou">Qianxinan</location>
,
<collectingRegion id="4443ECBA0641AE1D0DAB4AB004B9F937" box="[1167,1266,1755,1780]" country="China" name="Guizhou" pageId="3" pageNumber="247">Guizhou</collectingRegion>
,
<geoCoordinate id="E3B3449F0641AE1D0C244AB000E5F8DB" direction="north" orientation="latitude" pageId="3" pageNumber="247" precision="1" value="25.168236">25°10’05.65’’ N</geoCoordinate>
/
<geoCoordinate id="E3B3449F0641AE1D09934A9400A9F8DB" box="[183,226,1791,1816]" direction="east" orientation="longitude" pageId="3" pageNumber="247" precision="1" value="105.23075">105</geoCoordinate>
</journalOrPublisher>
°
<part id="F22388C50641AE1D09C84A94014CF8DB" box="[236,263,1791,1816]" pageId="3" pageNumber="247" type="volume">13</part>
’50.69’’ E,
<elevation id="0DAAC56B0641AE1D08A04A940197F8D4" box="[388,476,1791,1816]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="3" pageNumber="247" unit="m" value="1436.0">
<quantity id="417F8FBD0641AE1D08A04A940197F8D4" box="[388,476,1791,1816]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="3" pageNumber="247" unit="m" value="1436.0">
<pagination id="BF4EDCAC0641AE1D08A04A9401F7F8DB" box="[388,444,1791,1816]" pageId="3" pageNumber="247" type="pagination">1436</pagination>
m
</quantity>
</elevation>
,
<collectingDate id="E27DFD700641AE1D08CC4B6B0228F8DB" box="[488,611,1792,1816]" pageId="3" pageNumber="247" value="2017-05-12">
12-V-
<year id="F22F630B0641AE1D0B0C4B6B0228F8DB" box="[552,611,1792,1816]" pageId="3" pageNumber="247">2017</year>
</collectingDate>
,
<collectorName id="2B72478E0641AE1D0B4A4A9402B5F8DB" box="[622,766,1791,1816]" pageId="3" pageNumber="247">
<author id="5E3ACABD0641AE1D0B4A4A9402B5F8DB" box="[622,766,1791,1816]" fns="N" kam="kam" npo="FL" pageId="3" pageNumber="247">Mingyi Tian</author>
</collectorName>
,
<collectorName id="2B72478E0641AE1D0A2D4A9403C0F8DB" box="[777,907,1791,1816]" pageId="3" pageNumber="247">
<author id="5E3ACABD0641AE1D0A2D4A9403C0F8DB" box="[777,907,1791,1816]" fns="N" kam="kam" npo="FL" pageId="3" pageNumber="247">Weixin Liu</author>
</collectorName>
,
<collectorName id="2B72478E0641AE1D0AB34A940466F8D4" box="[919,1069,1791,1816]" pageId="3" pageNumber="247">
<author id="5E3ACABD0641AE1D0AB34A940466F8D4" box="[919,1069,1791,1816]" fns="N" npo="FL" pageId="3" pageNumber="247">Xiaozhu Luo</author>
</collectorName>
,
<collectorName id="2B72478E0641AE1D0D1C4A940492F8DB" box="[1080,1241,1791,1816]" pageId="3" pageNumber="247">
<author id="5E3ACABD0641AE1D0D1C4A940492F8DB" box="[1080,1241,1791,1816]" fns="N" npo="FL" pageId="3" pageNumber="247">Pingjing Yang</author>
</collectorName>
&amp;
<collectorName id="2B72478E0641AE1D0DDA4A94052FF8D4" box="[1278,1380,1791,1816]" pageId="3" pageNumber="247">
<author id="5E3ACABD0641AE1D0DDA4A94052FF8D4" box="[1278,1380,1791,1816]" fns="N" npo="FL" pageId="3" pageNumber="247">Yanyi Pu</author>
</collectorName>
leg., in
<collectionCode id="E096BA9D0641AE1D09904B4F0149F8FF" box="[180,258,1828,1852]" pageId="3" pageNumber="247">SCAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="36EF28050641AE1D08294B4F02DAF8FF" ID-GBIF-Occurrence="1915498332" box="[269,657,1827,1852]" collectingDate="2017-05-12" collectionCode="SCAU" collectorName="Mingyi Tian &amp; Weixin Liu &amp; Xiaozhu Luo &amp; Pingjing Yang &amp; Yanyi Pu" county="Fengyan Dong" elevation="1436" latitude="25.168236" location="Longguang" longLatPrecision="1" longitude="105.23075" municipality="Renjia Zhai" pageId="3" pageNumber="247" specimenCount="5" specimenCount-male="5" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="593C9CFA0641AE1D08294B4F0136F8FF" box="[269,381,1828,1852]" pageId="3" pageNumber="247" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9081E9D10641AE1D08A34B4801ABF8FF" box="[391,480,1827,1852]" pageId="3" pageNumber="247" type="male">5 males</specimenCount>
, ibid, in
<collectionCode id="E096BA9D0641AE1D0B664B4F02DAF8FF" box="[578,657,1828,1852]" pageId="3" pageNumber="247">SCAU</collectionCode>
</materialsCitation>
.
</bibRef>
</paragraph>
<paragraph id="863822580641AE1D09E34B2C0334F8A3" blockId="3.[151,1436,1755,2032]" box="[199,895,1863,1888]" pageId="3" pageNumber="247">
Length:
<quantity id="417F8FBD0641AE1D08014B2C01E4F89C" box="[293,431,1863,1888]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="7.5" metricValueMin="7.3" pageId="3" pageNumber="247" unit="mm" value="7.4" valueMax="7.5" valueMin="7.3">7.3–7.5 mm</quantity>
; width: 1.9–2.0 mm. Habitus as in
<figureCitation id="1EBC3EDD0641AE1D0A134B2C0330F8A3" box="[823,891,1863,1888]" captionStart="FIGURE 6" captionStartId="7.[151,250,2014,2036]" captionText="FIGURE 6. Habitus of Aspidaphaenops xiongda n. sp., male, paratype." pageId="3" pageNumber="247">Fig. 6</figureCitation>
.
</paragraph>
<paragraph id="863822580641AE1D09E34B000393F833" blockId="3.[151,1436,1755,2032]" pageId="3" pageNumber="247">Body yellowish brown to brown, but pale on mouthparts palps, tarsi and antennomeres 9–11; moderately shiny and whole body including abdominal ventrites glabrous, legs pubescent; microsculptural engraved meshes strongly transverse on head, pronotum and elytra; fore body (with mandibles) slightly longer than elytra, (HmL+PrL) / EL = 1.07–1.10, or shorter (excluding mandibles), (HlL+PrL) / EL = 0.90–0.91.</paragraph>
<caption id="D2F872D00646AE1A09B34BAE0280F818" box="[151,715,1989,2011]" pageId="4" pageNumber="248">
<paragraph id="863822580646AE1A09B34BAE0280F818" blockId="4.[151,715,1989,2011]" box="[151,715,1989,2011]" pageId="4" pageNumber="248">
<emphasis id="B4F3FE4A0646AE1A09B34BAE0159F819" bold="true" box="[151,274,1989,2011]" pageId="4" pageNumber="248">FIGURE 3.</emphasis>
Habitus of
<taxonomicName id="418759DB0646AE1A08AD4BAE02CDF819" authorityName="Ueno" authorityYear="2006" box="[393,646,1989,2010]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="248" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A0646AE1A08AD4BAE02CDF819" box="[393,646,1989,2010]" italics="true" pageId="4" pageNumber="248">Aspidaphaenops reflexus</emphasis>
</taxonomicName>
, male.
</paragraph>
</caption>
<caption id="D2F872D00647AE1B09B3491302CEFA6F" httpUri="https://zenodo.org/record/1455603/files/figure.png" pageId="5" pageNumber="249" startId="5.[151,250,1400,1422]" targetBox="[163,1411,196,1349]" targetPageId="5">
<paragraph id="863822580647AE1B09B3491302CEFA6F" blockId="5.[151,1435,1400,1452]" pageId="5" pageNumber="249">
<emphasis id="B4F3FE4A0647AE1B09B349130158FA4E" bold="true" box="[151,275,1400,1422]" pageId="5" pageNumber="249">FIGURE 4.</emphasis>
Male genitalia of
<taxonomicName id="418759DB0647AE1B08F44913023CFA4E" authorityName="Ueno" authorityYear="2006" box="[464,631,1400,1421]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B4F3FE4A0647AE1B08F44913023CFA4E" box="[464,631,1400,1421]" italics="true" pageId="5" pageNumber="249">Aspidaphaenops</emphasis>
</taxonomicName>
species, median lobe and parameres (lateral view), and apex (dorsal view) (A, B.
<taxonomicName id="418759DB0647AE1B099049FC0154FA6F" authorityName="Ueno" authorityYear="2006" box="[180,287,1431,1452]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="249" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A0647AE1B099049FC0154FA6F" box="[180,287,1431,1452]" italics="true" pageId="5" pageNumber="249">A. reflexus</emphasis>
</taxonomicName>
; C, D.
<taxonomicName id="418759DB0647AE1B084E49FC019CFA6F" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[362,471,1431,1452]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="249" phylum="Arthropoda" rank="species" species="xiongda" status="sp. nov.">
<emphasis id="B4F3FE4A0647AE1B084E49FC019CFA6F" box="[362,471,1431,1452]" italics="true" pageId="5" pageNumber="249">A. xiongda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC043310647AE1B08F949FC025CFA6F" box="[477,535,1431,1452]" pageId="5" pageNumber="249" rank="species">
<emphasis id="B4F3FE4A0647AE1B08F949FC025CFA6F" bold="true" box="[477,535,1431,1452]" pageId="5" pageNumber="249">n. sp.</emphasis>
</taxonomicNameLabel>
holotype).
</paragraph>
</caption>
<paragraph id="863822580647AE1B09E349B203D6F8A1" blockId="5.[151,1437,1497,2034]" pageId="5" pageNumber="249">Head elongated quadrate, much longer than wide, HLm/HW = 2.24–2.30, HLl/HW = 1.71–1.74; widest just behind sockets of antennae, at about 1/3 of head from labrum, genae gently and straightly narrowed posteriad, not parallel-sided; frons and vertex moderately convex, frontal furrows well-marked, deep and distinctly divergent posteriad, ended just behind middle head from labrum; clypeus transverse, quadrisetose; labrum transverse, frontal margin slightly emarginated, 6-setose; anterior and posterior frontal setiferous pores at apical 1/3 and basal 1/3 of head from labrum and clypeus respectively; mentum and submentum completely fused, mentum bisetose on either side of tooth which is simple and blunt at apex, basal fovea widely concave; submentum 11 to 12-setose; both maxillary and labial palps thin and subcylindrical, glabrous except for the 2nd labial palpomere which is bisetose on inner margin and one or two additional setae on outer margin; the 2nd labial palpomere nearly 1.4 times as long as 3rd; the 3rd maxillary palpomere nearly 1.3 times as long as 4th; suborbital pore closer to base of head than to submentum. Antennae thin and long, extending over apices of elytra.</paragraph>
<paragraph id="863822580647AE1B09E34B0603B8F831" blockId="5.[151,1437,1497,2034]" pageId="5" pageNumber="249">Prothorax not dilated, propleura invisible from above. Pronotum campanulate, longer than wide, PnL/PnW = 1.20–1.22, widest at about 1/3 of pronotum from base, recurved before hind angles which are not well-defined, fore angles protruded though obtuse; base as wide as front; latero-margins strongly widened and reflexed near base, fore latero-marginal setae at about apical 1/4 of pronotum. Scutellum fairly large.</paragraph>
<caption id="D2F872D00644AE1809B34B610208F8FD" httpUri="https://zenodo.org/record/1455605/files/figure.png" pageId="6" pageNumber="250" startId="6.[151,250,1802,1824]" targetBox="[173,1411,198,1774]" targetPageId="6">
<paragraph id="863822580644AE1809B34B610208F8FD" blockId="6.[151,1436,1802,1854]" pageId="6" pageNumber="250">
<emphasis id="B4F3FE4A0644AE1809B34B61015FF8DC" bold="true" box="[151,276,1802,1824]" pageId="6" pageNumber="250">FIGURE 5.</emphasis>
Cave Da Dong, locality of
<taxonomicName id="418759DB0644AE180B174B610294F8E3" authority="Ueno" authorityName="Ueno" authorityYear="2006" box="[563,735,1802,1824]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="250" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A0644AE180B174B6102EBF8DC" box="[563,672,1802,1823]" italics="true" pageId="6" pageNumber="250">A. reflexus</emphasis>
Uéno
</taxonomicName>
(A. entrance; B. cave chamber; C. a living exemplar of
<taxonomicName id="418759DB0644AE180C014B610080F8FD" authorityName="Ueno" authorityYear="2006" class="Insecta" family="Carabidae" genus="Satotrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="250" phylum="Arthropoda" rank="species" species="rieae">
<emphasis id="B4F3FE4A0644AE180C014B610080F8FD" italics="true" pageId="6" pageNumber="250">Satotrechus rieae</emphasis>
</taxonomicName>
; D. a living exemplar of
<taxonomicName id="418759DB0644AE1808EE4B42027EF8FD" authorityName="Ueno" authorityYear="2006" box="[458,565,1833,1854]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="250" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A0644AE1808EE4B42027EF8FD" box="[458,565,1833,1854]" italics="true" pageId="6" pageNumber="250">A. reflexus</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="863822580644AE1909E34B0003CBFF19" blockId="6.[151,1436,1899,2035]" lastBlockId="7.[151,1436,151,329]" lastPageId="7" lastPageNumber="251" pageId="6" pageNumber="250">
Elytra ovate (
<figureCitation id="1EBC3EDD0644AE1808454B0001F0F840" box="[353,443,1899,1924]" captionStart="FIGURE 7" captionStartId="8.[151,250,1774,1796]" captionTargetBox="[192,1394,201,1753]" captionTargetId="figure@8.[188,1398,193,1753]" captionTargetPageId="8" captionText="FIGURE 7. Left elytron of Aspidaphaenops species (A. A. xiongda n. sp.; B. A. dudou n. sp.)." httpUri="https://zenodo.org/record/1455607/files/figure.png" pageId="6" pageNumber="250">Fig. 7A</figureCitation>
), much longer than wide, EL/EW = 1.43–1.45; twice as wide as pronotum; widest near the middle, prehumeral borders strongly serrate, marginal gutters well-marked; punctures of striae isolated one another; sites of anterior dorsal pores variable, at the portions about 1/4 of elytra from base; middle pores at about middle of elytra; preapical pores much closer to elytral suture than to elytral apices; the 1st to 3rd pores of the marginal umbilicate series not equidistant, 2nd pore closer to 3rd than to 1st; distance between the 4th and 3rd subequal to that between 1
<taxonomicName id="418759DB0645AE1908734CD601DAFF19" box="[343,401,189,218]" pageId="7" pageNumber="251" rank="subSpecies" subSpecies="and">st and</taxonomicName>
second, 5th and 6th pores close to each other.
</paragraph>
<paragraph id="863822580645AE1909E34C8D01D3FE8A" blockId="7.[151,1436,151,329]" pageId="7" pageNumber="251">
Legs thin and long, bearing short pubescence; fore tibia smooth, presence of a faint longitudinal furrow or sulcus; the 1
<taxonomicName id="418759DB0645AE1908074D6301EEFEE7" box="[291,421,264,292]" pageId="7" pageNumber="251" rank="subSpecies" subSpecies="tarsomere">st tarsomere</taxonomicName>
, shorter, as long as and slightly longer than 2nd –4th tarsomeres together in fore, middle and hind legs, respectively.
</paragraph>
<caption id="D2F872D00645AE1909B34BB5033AF837" box="[151,881,2014,2036]" pageId="7" pageNumber="251">
<paragraph id="863822580645AE1909B34BB5033AF837" blockId="7.[151,881,2014,2036]" box="[151,881,2014,2036]" pageId="7" pageNumber="251">
<emphasis id="B4F3FE4A0645AE1909B34BB50159F830" bold="true" box="[151,274,2014,2036]" pageId="7" pageNumber="251">FIGURE 6.</emphasis>
Habitus of
<taxonomicName id="418759DB0645AE1908AD4BB502C2F830" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[393,649,2014,2035]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="251" phylum="Arthropoda" rank="species" species="xiongda" status="sp. nov.">
<emphasis id="B4F3FE4A0645AE1908AD4BB502C2F830" box="[393,649,2014,2035]" italics="true" pageId="7" pageNumber="251">Aspidaphaenops xiongda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC043310645AE190BAB4BB40283F837" box="[655,712,2015,2036]" pageId="7" pageNumber="251" rank="species">
<emphasis id="B4F3FE4A0645AE190BAB4BB40283F837" bold="true" box="[655,712,2015,2036]" pageId="7" pageNumber="251">n. sp.</emphasis>
</taxonomicNameLabel>
, male, paratype.
</paragraph>
</caption>
<caption id="D2F872D0064AAE1609B34A850413F8C7" box="[151,1112,1774,1796]" httpUri="https://zenodo.org/record/1455607/files/figure.png" pageId="8" pageNumber="252" startId="8.[151,250,1774,1796]" targetBox="[192,1394,201,1753]" targetPageId="8">
<paragraph id="86382258064AAE1609B34A850413F8C7" blockId="8.[151,1112,1774,1796]" box="[151,1112,1774,1796]" pageId="8" pageNumber="252">
<emphasis id="B4F3FE4A064AAE1609B34A850159F8C7" bold="true" box="[151,274,1774,1796]" pageId="8" pageNumber="252">FIGURE 7.</emphasis>
Left elytron of
<taxonomicName id="418759DB064AAE1608974A850211F8C0" authorityName="Ueno" authorityYear="2006" box="[435,602,1774,1795]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="B4F3FE4A064AAE1608974A850211F8C0" box="[435,602,1774,1795]" italics="true" pageId="8" pageNumber="252">Aspidaphaenops</emphasis>
</taxonomicName>
species (A.
<taxonomicName id="418759DB064AAE160BF34A84030EF8C0" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[727,837,1774,1796]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="252" phylum="Arthropoda" rank="species" species="xiongda" status="sp. nov.">
<emphasis id="B4F3FE4A064AAE160BF34A84030EF8C0" box="[727,837,1774,1796]" italics="true" pageId="8" pageNumber="252">A. xiongda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC04331064AAE160A6F4A8403CFF8C7" box="[843,900,1775,1796]" pageId="8" pageNumber="252" rank="species">
<emphasis id="B4F3FE4A064AAE160A6F4A8403CFF8C7" bold="true" box="[843,900,1775,1796]" pageId="8" pageNumber="252">n. sp.</emphasis>
</taxonomicNameLabel>
; B.
<taxonomicName id="418759DB064AAE160A8B4A840441F8C0" authority="Tian &amp; Huang, 2018" authorityName="Tian &amp; Huang" authorityYear="2018" box="[943,1034,1774,1796]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="252" phylum="Arthropoda" rank="species" species="dudou" status="sp. nov.">
<emphasis id="B4F3FE4A064AAE160A8B4A840441F8C0" box="[943,1034,1774,1796]" italics="true" pageId="8" pageNumber="252">A. dudou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFC04331064AAE160D354A840401F8C7" box="[1041,1098,1775,1796]" pageId="8" pageNumber="252" rank="species">
<emphasis id="B4F3FE4A064AAE160D354A840401F8C7" bold="true" box="[1041,1098,1775,1796]" pageId="8" pageNumber="252">n. sp.</emphasis>
</taxonomicNameLabel>
).
</paragraph>
</caption>
<paragraph id="86382258064AAE1609E34B400422F887" blockId="8.[151,1436,1835,2004]" box="[199,1129,1835,1860]" pageId="8" pageNumber="252">Ventrites IV–VI each with a pair of paramedian setae; ventrite VII bisetose in male</paragraph>
<paragraph id="86382258064AAE1709E34B240446FF17" blockId="8.[151,1436,1835,2004]" lastBlockId="9.[151,1437,151,680]" lastPageId="9" lastPageNumber="253" pageId="8" pageNumber="252">
Male genitalia (
<figureCitation id="1EBC3EDD064AAE1608A44B240244F8AB" box="[384,527,1871,1896]" captionStart="FIGURE 4" captionStartId="5.[151,250,1400,1422]" captionTargetBox="[163,1411,196,1349]" captionTargetId="figure@5.[163,1427,193,1387]" captionTargetPageId="5" captionText="FIGURE 4. Male genitalia of Aspidaphaenops species, median lobe and parameres (lateral view), and apex (dorsal view) (A, B. A. reflexus; C, D. A. xiongda n. sp. holotype)." httpUri="https://zenodo.org/record/1455603/files/figure.png" pageId="8" pageNumber="252">Figs. 4C, D</figureCitation>
): Moderately sclerotized, thin and slender, more elongate than in
<taxonomicName id="418759DB064AAE160C3E4B3B05DEF8AB" authorityName="Ueno" authorityYear="2006" box="[1306,1429,1871,1896]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="252" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A064AAE160C3E4B3B05DEF8AB" box="[1306,1429,1871,1896]" italics="true" pageId="8" pageNumber="252">A. reflexus</emphasis>
</taxonomicName>
, almost rectangularly arcuate ventrally at about basal 1/4, apical lobe shorter than in
<taxonomicName id="418759DB064AAE160D6B4B1F048CF84F" authorityName="Ueno" authorityYear="2006" box="[1103,1223,1907,1932]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="252" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A064AAE160D6B4B1F048CF84F" box="[1103,1223,1907,1932]" italics="true" pageId="8" pageNumber="252">A. reflexus</emphasis>
</taxonomicName>
, strongly recurved and reflexed at apex; base wide, sagittal aileron large and thick; inner sac of the median lobe with an anisotopic and short copulatory piece which is about 1/6 as long as the median lobe; in dorsal view, apex more or less tube-like, thin and nearly parallel-sided, broad at apex which is reflexed; parameres well-developed but much thinner than in
<taxonomicName id="418759DB064BAE1709B34CD70145FF16" authorityName="Ueno" authorityYear="2006" box="[151,270,188,213]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A064BAE1709B34CD70145FF16" box="[151,270,188,213]" italics="true" pageId="9" pageNumber="253">A. reflexus</emphasis>
</taxonomicName>
, about 1/3 as long as median lobe, each with three long apical setae.
</paragraph>
<paragraph id="86382258064BAE1709E34CB401D5FF3B" blockId="9.[151,1437,151,680]" box="[199,414,223,248]" pageId="9" pageNumber="253">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="CE9D71D3064BAE1709E34D6F02ADFE44" pageId="9" pageNumber="253" type="discussion">
<paragraph id="86382258064BAE1709E34D6F02ADFE44" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="253">
<emphasis id="B4F3FE4A064BAE1709E34D6F0170FEDF" bold="true" box="[199,315,260,284]" pageId="9" pageNumber="253">Remarks.</emphasis>
Similar to
<taxonomicName id="418759DB064BAE1708934D6F0310FEDE" authority="Ueno, 2006" authorityName="Ueno" authorityYear="2006" box="[439,859,260,285]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A064BAE1708934D6F0284FEDE" box="[439,719,260,285]" italics="true" pageId="9" pageNumber="253">Aspidaphaenops reflexus</emphasis>
Uéno, 2006
</taxonomicName>
, but differs from it by: (1) head gradually narrowed posteriad, instead of nearly parallel-sided in
<taxonomicName id="418759DB064BAE170BAE4D43034AFE83" authorityName="Ueno" authorityYear="2006" box="[650,769,295,320]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A064BAE170BAE4D43034AFE83" box="[650,769,295,320]" italics="true" pageId="9" pageNumber="253">A. reflexus</emphasis>
</taxonomicName>
; (2) fore angles of pronotum strongly protruded (vs. hardly protruded in
<taxonomicName id="418759DB064BAE1708094D2701ECFEA6" authorityName="Ueno" authorityYear="2006" box="[301,423,332,357]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A064BAE1708094D2701ECFEA6" box="[301,423,332,357]" italics="true" pageId="9" pageNumber="253">A. reflexus</emphasis>
</taxonomicName>
); (3) hind lobes of pronotum more reflexed; and (4) fore latero-marginal setae at about frontal 1/4 of pronotum, instead of 1/
<quantity id="417F8FBD064BAE170B124D040229FE4B" box="[566,610,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="9" pageNumber="253" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="418759DB064BAE170B4D4D1B02ABFE4B" authorityName="Ueno" authorityYear="2006" box="[617,736,367,392]" class="Insecta" family="Carabidae" genus="Aspidaphaenops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B4F3FE4A064BAE170B4D4D1B02ABFE4B" box="[617,736,367,392]" italics="true" pageId="9" pageNumber="253">A. reflexus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CE9D71D3064BAE1709E34DFF0477FE13" pageId="9" pageNumber="253" type="etymology">
<paragraph id="86382258064BAE1709E34DFF0477FE13" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="253">
<emphasis id="B4F3FE4A064BAE1709E34DFF0105FE6F" bold="true" box="[199,334,404,428]" pageId="9" pageNumber="253">Etymology.</emphasis>
“Xiongda” is a nickname of Dr. Philip John Rowsell, a famous British speleologist (Hong Meigui Cave Exploration Society), to thank his valuable support to our field trip in Xingyi.
</paragraph>
</subSubSection>
<subSubSection id="CE9D71D3064BAE1709E34DB003C6FD6B" pageId="9" pageNumber="253" type="distribution">
<paragraph id="86382258064BAE1709E34DB004C2FE37" blockId="9.[151,1437,151,680]" box="[199,1161,475,501]" pageId="9" pageNumber="253">
<emphasis id="B4F3FE4A064BAE1709E34DB0012BFE37" bold="true" box="[199,352,475,500]" pageId="9" pageNumber="253">Distribution.</emphasis>
<collectingRegion id="4443ECBA064BAE1708434DB7018CFE36" box="[359,455,476,501]" country="China" name="Guizhou" pageId="9" pageNumber="253">Guizhou</collectingRegion>
(Anlong). Known only from cave Fengyan Dong (
<figureCitation id="1EBC3EDD064BAE170D264DB70414FE36" box="[1026,1119,476,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1507,1529]" captionTargetBox="[216,1364,259,1473]" captionTargetId="figure@2.[207,1375,257,1479]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of Aspidaphaenops species (a. A. dudou n. sp.; b. A. volatidraconis; c. A. masakii; d. A. xiongda n. sp.; e. A. reflexus)." httpUri="https://zenodo.org/record/1455597/files/figure.png" pageId="9" pageNumber="253">Figs. 1d</figureCitation>
,
<figureCitation id="1EBC3EDD064BAE170D4F4DB70430FE37" box="[1131,1147,476,500]" captionStart="FIGURE 8" captionStartId="9.[151,250,1379,1401]" captionTargetBox="[352,1235,744,1348]" captionTargetId="figure@9.[340,1247,735,1356]" captionTargetPageId="9" captionText="FIGURE 8. Map of cave Fengyan Dong, type locality of Aspidaphaenops xiongda n. sp. (courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society, triangles indicating where the beetles were collected)." httpUri="https://zenodo.org/record/1455609/files/figure.png" pageId="9" pageNumber="253">8</figureCitation>
).
</paragraph>
<paragraph id="86382258064BAE1709E34E6B03C6FD6B" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="253">
Fengyan Dong opens at the right side of a road to the village Renjia Zhai, with a large entrance and a small stream throughout the main passage (
<figureCitation id="1EBC3EDD064BAE170B194E4F0288FDFF" box="[573,707,548,573]" captionStart="FIGURE 9" captionStartId="10.[151,250,1884,1906]" captionTargetBox="[280,1313,437,1864]" captionTargetId="figure@10.[278,1315,434,1866]" captionTargetPageId="10" captionText="FIGURE 9. Cave Fengyan Dong, type locality of Aspidaphaenops xiongda n. sp. (A. entrance; B. main passage; C, D. active beetles in the cave).Prothorax not dilated, propleura invisible from above. Pronotum barrel-like, longer than wide, PnL/PnW = 1.19–1.20, widest near the middle of pronotum, strongly recurved before hind angles which are not well-defined, fore angles obtuse; base as wide as front; latero-margins gradually widened and reflexed from middle to base, fore latero-marginal setae at about 1/5 anterior of pronotum. Scutellum fairly large." httpUri="https://zenodo.org/record/1455611/files/figure.png" pageId="9" pageNumber="253">Figs. 9A, B</figureCitation>
). It is about
<quantity id="417F8FBD064BAE170A764E4F03ECFDFF" box="[850,935,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="253" unit="m" value="1100.0">1100 m</quantity>
long, with two branches at both sides of the main passage. Both are longer than the main passage. There are several small portions which are muddy and moist enough and good for cave beetles (
<figureCitation id="1EBC3EDD064BAE170B194E070286FD47" box="[573,717,620,645]" captionStart="FIGURE 9" captionStartId="10.[151,250,1884,1906]" captionTargetBox="[280,1313,437,1864]" captionTargetId="figure@10.[278,1315,434,1866]" captionTargetPageId="10" captionText="FIGURE 9. Cave Fengyan Dong, type locality of Aspidaphaenops xiongda n. sp. (A. entrance; B. main passage; C, D. active beetles in the cave).Prothorax not dilated, propleura invisible from above. Pronotum barrel-like, longer than wide, PnL/PnW = 1.19–1.20, widest near the middle of pronotum, strongly recurved before hind angles which are not well-defined, fore angles obtuse; base as wide as front; latero-margins gradually widened and reflexed from middle to base, fore latero-marginal setae at about 1/5 anterior of pronotum. Scutellum fairly large." httpUri="https://zenodo.org/record/1455611/files/figure.png" pageId="9" pageNumber="253">Figs. 9C, D</figureCitation>
). This new species is sympatric with another trechine beetle
<taxonomicName id="418759DB064BAE1709B34EE40140FD6B" authorityName="Jeannel" authorityYear="1954" box="[151,267,655,680]" class="Insecta" family="Carabidae" genus="Trechiotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B4F3FE4A064BAE1709B34EE40140FD6B" box="[151,267,655,680]" italics="true" pageId="9" pageNumber="253">Trechiotes</emphasis>
</taxonomicName>
sp., millipedes, pseudoscorpions, spiders and springtails.
</paragraph>
</subSubSection>
</treatment>
</document>
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