387 lines
50 KiB
XML
387 lines
50 KiB
XML
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<mods:title id="1936CDD93B2FFBEC183BAE8B3856907C">Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae)</mods:title>
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<mods:namePart id="7DD3B2BCA38FE6B7166484CBD2698782">Fritz, Uwe</mods:namePart>
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<mods:affiliation id="76E79FB83EC497029E906C8AC3B65602">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:nameIdentifier id="4B27EEA18FF94A7CF99D5413E321485D" type="email">uwe.fritz@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="E29F6F5433BCCE84A4EC625A29CB6F9A">Petzold, Alice</mods:namePart>
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<mods:affiliation id="AF45A9E750E6C11D4161F1B50D17F4AD">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="049B433879CAC536B58D056F1C4860FF">Kehlmaier, Christian</mods:namePart>
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<mods:affiliation id="A0D0B3E635956EC8798D97287D9E276E">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="6E865FD0261B727B09D07C19952EFDFC">Kindler, Carolin</mods:namePart>
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<mods:affiliation id="E0CE266EF35D99E45BC4A07FA565A34F">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="F74FFD28B802BE10A045C156729D3CBB">Campbell, Patrick</mods:namePart>
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<mods:affiliation id="4D3AC051F2FA586F4A50E7D47B108C6F">Department of Zoology, Darwin Centre (DC 1), Natural History Museum, Cromwell Road, South Kensington, London SW 7 5 BD, England</mods:affiliation>
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<mods:namePart id="8F6B05B224596CA9AA3044CA14715B80">Hofmeyr, Margaretha D.</mods:namePart>
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<mods:affiliation id="91D908E2BC7FBBA93297D143C32ECDC1">Chelonian Biodiversity and Conservation, Department of Biodiversity and Conservation Biology, University of the Western Cape, Private Bag X 17, Bellville 7535, South Africa</mods:affiliation>
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<mods:namePart id="F8AED52AC9943B1BEDD6158F5CC2A813">Branch, William R.</mods:namePart>
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<mods:affiliation id="AC49D41696307A32739B796B3DE44C1A">Department of Herpetology, Port Elizabeth Museum, P. O. Box 13147, Humewood 6013, South Africa & Department of Zoology, P. O. Box 77000, Nelson Mandela Metropolitan University, Port Elizabeth 6031, South Africa Corresponding author. E-mail: uwe. fritz @ senckenberg. de</mods:affiliation>
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<mods:title id="1C5E2CF1DFF68DF9BE3988E549C88371">Zootaxa</mods:title>
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<mods:date id="F8DCCF6B344C9CD4DE4372853EA265D8">2014</mods:date>
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<mods:number id="5F117DA3C5EE0F430E7DC8630C1447D3">2014-05-15</mods:number>
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<mods:number id="56FA223235761197C166E00FEA7651A3">3795</mods:number>
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<treatment id="03B68780CF56102418C3EC51FDFFF842" ID-DOI="http://doi.org/10.5281/zenodo.5082879" ID-GBIF-Taxon="183403825" ID-Zenodo-Dep="5082879" LSID="urn:lsid:plazi:treatment:03B68780CF56102418C3EC51FDFFF842" httpUri="http://treatment.plazi.org/id/03B68780CF56102418C3EC51FDFFF842" lastPageId="8" lastPageNumber="509" pageId="6" pageNumber="507">
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<subSubSection id="C305651DCF56102A18C3EC51FD6FF821" box="[151,529,1896,1922]" pageId="6" pageNumber="507" type="nomenclature">
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<paragraph id="8BA03696CF56102A18C3EC51FD6FF821" blockId="6.[151,529,1896,1922]" box="[151,529,1896,1922]" pageId="6" pageNumber="507">
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<heading id="D0E881FACF56102A18C3EC51FD6FF821" bold="true" box="[151,529,1896,1922]" fontSize="11" level="1" pageId="6" pageNumber="507" reason="1">
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<taxonomicName id="4C1F4D15CF56102A18C3EC51FD6FF821" authority="Schoepff, 1792" authorityName="Schoepff" authorityYear="1792" box="[151,529,1896,1922]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="507" phylum="Chordata" rank="species" species="galeata">
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<emphasis id="B96BEA84CF56102A18C3EC51FD6FF821" bold="true" box="[151,529,1896,1922]" pageId="6" pageNumber="507">
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<emphasis id="B96BEA84CF56102A18C3EC51FE2DF821" bold="true" box="[151,339,1896,1922]" italics="true" pageId="6" pageNumber="507">Testudo galeata</emphasis>
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<bibRefCitation id="EF8E4B67CF56102A190EEC51FD6FF821" author="Schoepff, I. D." box="[346,529,1896,1922]" pageId="6" pageNumber="507" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff, 1792</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C305651DCF56102B18C3EC9EFE2AFBFB" lastPageId="7" lastPageNumber="508" pageId="6" pageNumber="507" type="description">
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<paragraph id="8BA03696CF56102B18C3EC9EFCDEFEBE" blockId="6.[151,1436,1959,2021]" lastBlockId="7.[151,1437,151,1977]" lastPageId="7" lastPageNumber="508" pageId="6" pageNumber="507">
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The name
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<taxonomicName id="4C1F4D15CF56102A1947EC9EFEB9F863" authorityName="Schoepff" authorityYear="1792" box="[275,455,1959,1984]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="507" phylum="Chordata" rank="species" species="galeata">
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<emphasis id="B96BEA84CF56102A1947EC9EFEB9F863" box="[275,455,1959,1984]" italics="true" pageId="6" pageNumber="507">Testudo galeata</emphasis>
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</taxonomicName>
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was erected by
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<bibRefCitation id="EF8E4B67CF56102A1AD3EC9EFC42F863" author="Schoepff, I. D." box="[647,828,1959,1984]" pageId="6" pageNumber="507" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff (1792</bibRefCitation>
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: p. 12) in his famous “
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<emphasis id="B96BEA84CF56102A1C18EC91FF80F846" italics="true" pageId="6" pageNumber="507">Historia Testudinum iconibus illustrata</emphasis>
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”. For a long time
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<taxonomicName id="4C1F4D15CF56102A1986ECF5FDFAF846" authorityName="Schoepff" authorityYear="1792" box="[466,644,1996,2021]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="6" pageNumber="507" phylum="Chordata" rank="species" species="galeata">
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<emphasis id="B96BEA84CF56102A1986ECF5FDFAF846" box="[466,644,1996,2021]" italics="true" pageId="6" pageNumber="507">Testudo galeata</emphasis>
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</taxonomicName>
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was used for helmeted terrapins, until
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<bibRefCitation id="EF8E4B67CF56102A1C15ECF5FB8EF846" author="Mertens, R." box="[1089,1264,1996,2021]" pageId="6" pageNumber="507" pagination="139 - 142" refId="ref12650" refString="Mertens, R. (1937) Bemerkungen uber die Rassen von Pelomedusa subrufa (La Cepede). Zoologischer Anzeiger, 117, 139 - 142." type="journal article" year="1937">Mertens (1937)</bibRefCitation>
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and
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<bibRefCitation id="EF8E4B67CF56102B1D7DECF5FF9FFF13" author="Loveridge, A." lastPageId="7" lastPageNumber="508" pageId="6" pageNumber="507" pagination="467 - 524" refId="ref12616" refString="Loveridge, A. (1941) Revision of the African terrapin [sic] of the family Pelomedusidae. Bulletin of the Museum of Comparative Zoology, 88, 467 - 524." type="journal article" year="1941">Loveridge (1941)</bibRefCitation>
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pointed out that the older name
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<taxonomicName id="4C1F4D15CF57102B1A31EBAEFC62FF13" authorityName="Bonnaterre" authorityYear="1789" box="[613,796,151,176]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="subrufa">
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<emphasis id="B96BEA84CF57102B1A31EBAEFC62FF13" box="[613,796,151,176]" italics="true" pageId="7" pageNumber="508">Testudo subrufa</emphasis>
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</taxonomicName>
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should be applied. Schoepff’s original description was based on detailed unpublished notes received from two Swedish naturalists. Originally,
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<bibRefCitation id="EF8E4B67CF57102B1C3CEB85FA66FF76" author="Schoepff, I. D." box="[1128,1304,188,213]" pageId="7" pageNumber="508" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff (1792</bibRefCitation>
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: pp. 12–16) intended to assemble information about another terrapin species,
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<taxonomicName id="4C1F4D15CF57102B1BD7EBE6FB97FF5B" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[899,1257,223,248]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="scabra">
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<emphasis id="B96BEA84CF57102B1BD7EBE6FB51FF5B" box="[899,1071,223,248]" italics="true" pageId="7" pageNumber="508">Testudo scabra</emphasis>
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<bibRefCitation id="EF8E4B67CF57102B1C6CEBE6FB97FF5B" author="Linnaeus, C." box="[1080,1257,223,248]" pageId="7" pageNumber="508" pagination="1" refId="ref12541" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Regnum animale. Editio decima, reformata. Laurentius Salvius, Stockholm, (3) + 823 + (1) pp. http: // dx. doi. org / 10.5962 / bhl. title. 542" type="journal article" year="1758">Linnaeus, 1758</bibRefCitation>
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</taxonomicName>
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, a taxon whose identity puzzled scientists for about 250 years (
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<bibRefCitation id="EF8E4B67CF57102B1AF3EA3DFCEAFEBE" author="Rhodin, A. G. J. & Carr, J. L." box="[679,916,260,285]" pageId="7" pageNumber="508" pagination="1 - 18" refId="ref12821" refString="Rhodin, A. G. J. & Carr, J. L. (2009) A quarter millenium of uses and misuses of the turtle name Testudo scabra: identification of the type specimens of T. scabra Linnaeus 1758 (= Rhinoclemmys punctularia) and T. scripta Thunberg in Schoepff 1792 (= Trachemys scripta scripta). Zootaxa, 2226, 1 - 18." type="journal article" year="2009">Rhodin & Carr 2009</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BA03696CF57102B1893EA1EFBADFDFC" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="508">
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Schoepff received from Anders Jahan Retzius (1742–1821) of Lund a detailed description and drawings of a “
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<taxonomicName id="4C1F4D15CF57102B18F7EA75FE32FEC6" authorityName="Linnaeus" authorityYear="1758" box="[163,332,332,357]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="scabra">
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<emphasis id="B96BEA84CF57102B18F7EA75FE32FEC6" box="[163,332,332,357]" italics="true" pageId="7" pageNumber="508">Testudo scabra</emphasis>
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</taxonomicName>
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” from “
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<collectingCountry id="F3087606CF57102B19FCEA75FE9AFEC6" box="[424,484,332,357]" name="India" pageId="7" pageNumber="508">India</collectingCountry>
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orientalis” [East
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<collectingCountry id="F3087606CF57102B1AF3EA75FD98FEC6" box="[679,742,332,357]" name="India" pageId="7" pageNumber="508">India</collectingCountry>
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], which Retzius had kept alive for two years.
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<bibRefCitation id="EF8E4B67CF57102B1CB2EA75FAE2FEC6" author="Schoepff, I. D." box="[1254,1436,332,357]" pageId="7" pageNumber="508" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff (1792)</bibRefCitation>
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published Retzius’ description (pp. 13–14) and a coloured etching of the terrapin as figure
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<quantity id="4CE79B73CF57102B1CF9EA56FBA4FE2B" box="[1197,1242,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="508" unit="in" value="1.0">1 in</quantity>
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plate III. Shown under the name “
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<taxonomicName id="4C1F4D15CF57102B1901EAACFD61FE0E" authority="Retz." authorityName="Retz." box="[341,543,404,429]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="scabra">
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<emphasis id="B96BEA84CF57102B1901EAACFEA4FE0E" box="[341,474,404,429]" italics="true" pageId="7" pageNumber="508">Test. scabra</emphasis>
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Retz.
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</taxonomicName>
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”, it is clearly a juvenile
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<taxonomicName id="4C1F4D15CF57102B1B7EEAADFB71FE0E" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[810,1039,404,429]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="subrufa">
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<emphasis id="B96BEA84CF57102B1B7EEAADFB71FE0E" box="[810,1039,404,429]" italics="true" pageId="7" pageNumber="508">Pelomedusa subrufa</emphasis>
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</taxonomicName>
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(
|
||
<figureCitation id="13242A13CF57102B1C4AEAADFB1FFE0E" box="[1054,1121,404,429]" captionStart="FIGURE 3" captionStartId="8.[151,250,1597,1619]" captionTargetBox="[224,1354,222,1560]" captionTargetId="figure-260@8.[209,1384,195,1578]" captionTargetPageId="8" captionText="FIGURE 3. Top, left: Folio from Schoepff’s legacy (Museum für Naturkunde zu Berlin, Historische Bild- und Schriftgutsammlungen, SI, Nachl. Schoepff II) consisting of the drawings provided by Retzius and an imprint of the resultant etching. Retzius’ hand-written German text reads:" figureDoi="http://doi.org/10.5281/zenodo.4915070" httpUri="https://zenodo.org/record/4915070/files/figure.png" pageId="7" pageNumber="508">Fig. 3</figureCitation>
|
||
, top), and “Prof. D. Swartz” (probably to be identified with Olof Peter Swartz, 1760–1818) of Stockholm wrote to Schoepff that a very similar specimen in spirit is present in the Cabinet of the Royal Academy of Sciences at Stockholm.
|
||
<bibRefCitation id="EF8E4B67CF57102B1CF5EAE5FA2FFE56" author="Schoepff, I. D." box="[1185,1361,476,501]" pageId="7" pageNumber="508" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff (1792</bibRefCitation>
|
||
: p. 16) concluded that these two terrapins could represent a new species, for which he proposed the name
|
||
<taxonomicName id="4C1F4D15CF57102B1CB0EAC6FAEBFDBB" authorityName="Schoepff" authorityYear="1792" box="[1252,1429,511,536]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF57102B1CB0EAC6FAEBFDBB" box="[1252,1429,511,536]" italics="true" pageId="7" pageNumber="508">Testudo galeata</emphasis>
|
||
</taxonomicName>
|
||
. The eponymous character (
|
||
<taxonomicName id="4C1F4D15CF57102B199CE91DFD1BFD9E" authority=", Latin" authorityName="Latin" box="[456,613,548,573]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF57102B199CE91DFD62FD9E" box="[456,540,548,573]" italics="true" pageId="7" pageNumber="508">galeata</emphasis>
|
||
, Latin
|
||
</taxonomicName>
|
||
= helmeted) of this putative new species is its helmet-like head scalation. The two terrapins from Lund and Stockholm have to be regarded as
|
||
<typeStatus id="54A48834CF57102B1BC7E971FC8BFDC3" box="[915,1013,584,608]" pageId="7" pageNumber="508" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1F4D15CF57102B1C4FE97EFBB2FDC3" box="[1051,1228,583,608]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF57102B1C4FE97EFBB2FDC3" box="[1051,1228,583,608]" italics="true" pageId="7" pageNumber="508">Testudo galeata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA03696CF57102B1893E955FE2AFBFB" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="508">
|
||
In the collection of the Biological Museum of Lund University there is still extant a single alcohol-preserved
|
||
<taxonomicName id="4C1F4D15CF57102B18C3E9B6FE5EFD0B" box="[151,288,655,680]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA84CF57102B18C3E9B6FE5EFD0B" box="[151,288,655,680]" italics="true" pageId="7" pageNumber="508">Pelomedusa</emphasis>
|
||
</taxonomicName>
|
||
specimen, bearing the catalogue number ZMUL 6481 (
|
||
<figureCitation id="13242A13CF57102B1BDEE9B6FCB0FD0B" box="[906,974,655,680]" captionStart="FIGURE 3" captionStartId="8.[151,250,1597,1619]" captionTargetBox="[224,1354,222,1560]" captionTargetId="figure-260@8.[209,1384,195,1578]" captionTargetPageId="8" captionText="FIGURE 3. Top, left: Folio from Schoepff’s legacy (Museum für Naturkunde zu Berlin, Historische Bild- und Schriftgutsammlungen, SI, Nachl. Schoepff II) consisting of the drawings provided by Retzius and an imprint of the resultant etching. Retzius’ hand-written German text reads:" figureDoi="http://doi.org/10.5281/zenodo.4915070" httpUri="https://zenodo.org/record/4915070/files/figure.png" pageId="7" pageNumber="508">Fig. 3</figureCitation>
|
||
, bottom). Retzius donated this terrapin as “
|
||
<taxonomicName id="4C1F4D15CF57102B18F7E98DFE33FD6E" box="[163,333,692,717]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="scabra">
|
||
<emphasis id="B96BEA84CF57102B18F7E98DFE33FD6E" box="[163,333,692,717]" italics="true" pageId="7" pageNumber="508">Testudo scabra</emphasis>
|
||
</taxonomicName>
|
||
” in 1811 to the Lund Museum, together with a large collection of other specimens (L. Lundqvist, pers. comm.). We conclude that ZMUL 6481, which originates evidently from Retzius, has to be identified with the
|
||
<typeStatus id="54A48834CF57102B18C3E9C5FF90FCB7" box="[151,238,764,788]" pageId="7" pageNumber="508" type="syntype">syntype</typeStatus>
|
||
described in detail in
|
||
<bibRefCitation id="EF8E4B67CF57102B19B2E9C5FDE9FCB6" author="Schoepff, I. D." box="[486,663,764,789]" pageId="7" pageNumber="508" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff (1792</bibRefCitation>
|
||
: pp. 13–14). The size of ZMUL 6481 (straight carapacial length
|
||
<quantity id="4CE79B73CF57102B1D3FE9C5FFC2FC94" metricMagnitude="-2" metricUnit="m" metricValue="5.84" pageId="7" pageNumber="508" unit="cm" value="5.84">5.84 cm</quantity>
|
||
, width
|
||
<quantity id="4CE79B73CF57102B1940E826FE0AFC9B" box="[276,372,799,824]" metricMagnitude="-2" metricUnit="m" metricValue="4.97" pageId="7" pageNumber="508" unit="cm" value="4.97">4.97 cm</quantity>
|
||
) approximately matches the measurements given in
|
||
<bibRefCitation id="EF8E4B67CF57102B1B83E826FBF5FC9B" author="Schoepff, I. D." box="[983,1163,799,824]" pageId="7" pageNumber="508" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff (1792</bibRefCitation>
|
||
: p. 13) with 2 ½ and 2 Prussian inches (1 Prussian inch =
|
||
<quantity id="4CE79B73CF57102B1A73E87DFDEBFCFE" box="[551,661,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.615" pageId="7" pageNumber="508" unit="cm" value="2.615">2.615 cm</quantity>
|
||
), especially when it is considered that Retzius could have measured over the curve of the carapace. However, the shell shape of ZMUL 6481 is a bit too oval compared to Schoepff’s figure (
|
||
<figureCitation id="13242A13CF57102B18B9E8B5FE4BFC06" box="[237,309,908,933]" captionStart="FIGURE 3" captionStartId="8.[151,250,1597,1619]" captionTargetBox="[224,1354,222,1560]" captionTargetId="figure-260@8.[209,1384,195,1578]" captionTargetPageId="8" captionText="FIGURE 3. Top, left: Folio from Schoepff’s legacy (Museum für Naturkunde zu Berlin, Historische Bild- und Schriftgutsammlungen, SI, Nachl. Schoepff II) consisting of the drawings provided by Retzius and an imprint of the resultant etching. Retzius’ hand-written German text reads:" figureDoi="http://doi.org/10.5281/zenodo.4915070" httpUri="https://zenodo.org/record/4915070/files/figure.png" pageId="7" pageNumber="508">Fig. 3</figureCitation>
|
||
, top). Retzius’ two original drawings of the terrapin and part of his letter have survived among Schoepff’s legacy, and are now kept in the Museum für Naturkunde,
|
||
<collectingRegion id="49DBF874CF57102B1BCFE896FC9FFC6B" box="[923,993,943,968]" country="Germany" name="Berlin" pageId="7" pageNumber="508">Berlin</collectingRegion>
|
||
(
|
||
<figureCitation id="13242A13CF57102B1BA4E896FB4BFC6B" box="[1008,1077,943,968]" captionStart="FIGURE 3" captionStartId="8.[151,250,1597,1619]" captionTargetBox="[224,1354,222,1560]" captionTargetId="figure-260@8.[209,1384,195,1578]" captionTargetPageId="8" captionText="FIGURE 3. Top, left: Folio from Schoepff’s legacy (Museum für Naturkunde zu Berlin, Historische Bild- und Schriftgutsammlungen, SI, Nachl. Schoepff II) consisting of the drawings provided by Retzius and an imprint of the resultant etching. Retzius’ hand-written German text reads:" figureDoi="http://doi.org/10.5281/zenodo.4915070" httpUri="https://zenodo.org/record/4915070/files/figure.png" pageId="7" pageNumber="508">Fig. 3</figureCitation>
|
||
, top). The original drawings are very rough, but they illustrate a shell shape that agrees better with ZMUL 6481 than with
|
||
<bibRefCitation id="EF8E4B67CF57102B1C99E8EDFAE2FC4E" author="Schoepff, I. D." box="[1229,1436,980,1005]" pageId="7" pageNumber="508" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff’s (1792)</bibRefCitation>
|
||
published figure. In comparison it appears that Schoepff’s figure is somewhat idealized. The coloration of the
|
||
<typeStatus id="54A48834CF57102B18C3EF25FF90FB97" box="[151,238,1052,1076]" pageId="7" pageNumber="508" type="syntype">syntype</typeStatus>
|
||
is much faded due to the age of the specimen and allows no direct comparison with the description in
|
||
<bibRefCitation id="EF8E4B67CF57102B18C3EF06FE2EFBFB" author="Schoepff, I. D." box="[151,336,1087,1112]" pageId="7" pageNumber="508" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff (1792)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305651DCF5710241893EF5DFE12F826" lastPageId="8" lastPageNumber="509" pageId="7" pageNumber="508" type="discussion">
|
||
<paragraph id="8BA03696CF57102B1893EF5DFA20FAF6" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="508">
|
||
In the Swedish Museum of Natural History, Stockholm, there are two old
|
||
<taxonomicName id="4C1F4D15CF57102B1BAFEF5DFBFAFBDE" box="[1019,1156,1124,1149]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA84CF57102B1BAFEF5DFBFAFBDE" box="[1019,1156,1124,1149]" italics="true" pageId="7" pageNumber="508">Pelomedusa</emphasis>
|
||
</taxonomicName>
|
||
specimens catalogued as one lot (NRM 7043), which were once part of the collection of Charles de Geer (1720–1778). These specimens were transferred by his widow to the collection of the Royal Swedish Academy of Sciences in 1778. The two alcohol-preserved terrapins are in quite bad condition with many shell scutes missing. They are the only helmeted terrapins which were present in the collection when Swartz corresponded with Schoepff, so that one of these specimens is most likely the second
|
||
<typeStatus id="54A48834CF57102B1A66EE21FDF7FA93" box="[562,649,1304,1328]" pageId="7" pageNumber="508" type="syntype">syntype</typeStatus>
|
||
(E. Åhlander, pers. comm.). However, as it cannot be unambiguously identified, we choose to designate hereby the Lund specimen as
|
||
<typeStatus id="54A48834CF57102B1B3CEE05FCB0FAF6" box="[872,974,1340,1365]" pageId="7" pageNumber="508" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1F4D15CF57102B1BA0EE05FA24FAF6" authority="Schoepff, 1792" authorityName="Schoepff" authorityYear="1792" box="[1012,1370,1340,1365]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF57102B1BA0EE05FBDBFAF6" box="[1012,1189,1340,1365]" italics="true" pageId="7" pageNumber="508">Testudo galeata</emphasis>
|
||
<bibRefCitation id="EF8E4B67CF57102B1CF8EE05FA24FAF6" author="Schoepff, I. D." box="[1196,1370,1340,1365]" pageId="7" pageNumber="508" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff, 1792</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA03696CF57102B1893EE66FE9DF88B" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="508">
|
||
<bibRefCitation id="EF8E4B67CF57102B1893EE66FE1EFADB" author="Hewitt, J." box="[199,352,1375,1400]" pageId="7" pageNumber="508" pagination="283 - 357" refId="ref12349" refString="Hewitt, J. (1935) Some new forms of batrachians and reptiles from South Africa. Records of the Albany Museum, 4, 283 - 357, plates XXVII - XXXVI." type="journal article" year="1935">Hewitt (1935</bibRefCitation>
|
||
: p. 326) corrected the
|
||
<typeStatus id="54A48834CF57102B1A09EE59FDF3FADB" box="[605,653,1376,1400]" pageId="7" pageNumber="508">type</typeStatus>
|
||
locality “
|
||
<collectingCountry id="F3087606CF57102B1AAFEE66FC49FADB" box="[763,823,1375,1400]" name="India" pageId="7" pageNumber="508">India</collectingCountry>
|
||
orientalis” of
|
||
<taxonomicName id="4C1F4D15CF57102B1B8EEE66FBF2FADB" authorityName="Schoepff" authorityYear="1792" box="[986,1164,1375,1400]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF57102B1B8EEE66FBF2FADB" box="[986,1164,1375,1400]" italics="true" pageId="7" pageNumber="508">Testudo galeata</emphasis>
|
||
</taxonomicName>
|
||
to the environs of Cape Town (and not Cape Flats as thought by
|
||
<bibRefCitation id="EF8E4B67CF57102B1A25EEBDFC6CFA3E" author="Mertens, R." box="[625,786,1412,1437]" pageId="7" pageNumber="508" pagination="139 - 142" refId="ref12650" refString="Mertens, R. (1937) Bemerkungen uber die Rassen von Pelomedusa subrufa (La Cepede). Zoologischer Anzeiger, 117, 139 - 142." type="journal article" year="1937">Mertens 1937</bibRefCitation>
|
||
: p. 139, later repeated by
|
||
<bibRefCitation id="EF8E4B67CF57102B1C16EEBDFB85FA3E" author="Loveridge, A." box="[1090,1275,1412,1437]" pageId="7" pageNumber="508" pagination="467 - 524" refId="ref12616" refString="Loveridge, A. (1941) Revision of the African terrapin [sic] of the family Pelomedusidae. Bulletin of the Museum of Comparative Zoology, 88, 467 - 524." type="journal article" year="1941">Loveridge 1941</bibRefCitation>
|
||
: p. 470). This makes sense as the Dutch East
|
||
<collectingCountry id="F3087606CF57102B1A5FEE9EFD3BFA63" box="[523,581,1447,1472]" name="India" pageId="7" pageNumber="508">India</collectingCountry>
|
||
Company (Vereenigde Oost-Indische Compagnie) in 1652 established an outpost near the Cape of Good Hope for provisioning their ships coming from East
|
||
<collectingCountry id="F3087606CF57102B1C28EEF5FBC7FA46" box="[1148,1209,1484,1509]" name="India" pageId="7" pageNumber="508">India</collectingCountry>
|
||
. This outpost, later becoming the Cape Colony and finally Cape Town, was a supplier of early natural history specimens for Europe (
|
||
<bibRefCitation id="EF8E4B67CF57102B18CAED2DFEFFF98E" author="Wren-Sargent, M." box="[158,385,1556,1581]" pageId="7" pageNumber="508" pagination="26 - 31" refId="ref14026" refString="Wren-Sargent, M. (1999) Fieldguides, past and present. Part 2: South African publications. Bird Numbers, 8, 26 - 31." type="journal article" year="1999">Wren-Sargent 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B67CF57102B19C5ED2DFDDFF98E" author="Bauer, A. M. & Gunther, R." box="[401,673,1556,1581]" pageId="7" pageNumber="508" pagination="167 - 185" refId="ref11033" refString="Bauer, A. M. & Gunther, R. (2013) Origin and identity of the von Borcke collection of amphibians and reptiles in the Museum fur Naturkunde in Berlin: a cache of Seba specimens? Zoosystematics and Evolution, 89, 167 - 185. http: // dx. doi. org / 10.1002 / zoos. 201300005" type="journal article" year="2013">Bauer & Günther 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B67CF57102B1AE5ED2DFCADF98E" author="Bauer, A. M. & Wahlgren, R." box="[689,979,1556,1581]" pageId="7" pageNumber="508" pagination="220 - 252" refId="ref11097" refString="Bauer, A. M. & Wahlgren, R. (2013) On the Linck collection and specimens of snakes figured by Johann Jakob Scheuchzer (1735) - the oldest fluid-preserved herpetological collection in the world? Bonn zoological Bulletin, 62, 220 - 252." type="journal article" year="2013">Bauer & Wahlgren 2013</bibRefCitation>
|
||
), and such specimens might have been mixed up or confused with materials coming from East
|
||
<collectingCountry id="F3087606CF57102B1B59ED0EFC34F9F3" box="[781,842,1591,1616]" name="India" pageId="7" pageNumber="508">India</collectingCountry>
|
||
.
|
||
<bibRefCitation id="EF8E4B67CF57102B1B01ED0EFCA4F9F3" author="Gray, J. E." box="[853,986,1591,1616]" pageId="7" pageNumber="508" refId="ref12093" refString="Gray, J. E. (1831) Synopsis Reptilium; or Short Descriptions of the Species of Reptiles. Part I. - Cataphracta. Tortoises, Crocodiles, and Enaliosaurians. Treuttel, Wurtz, and Co., London, viii + 85 pp." type="book" year="1831">Gray (1831</bibRefCitation>
|
||
: p. 40) wrote exactly in this spirit about helmeted terrapins “I have never seen this tortoise from
|
||
<collectingCountry id="F3087606CF57102B1B5CED65FC3AF9D6" box="[776,836,1628,1653]" name="India" pageId="7" pageNumber="508">India</collectingCountry>
|
||
, but being brought from the
|
||
<collectingRegion id="49DBF874CF57102B1CD6ED65FBC2F9D6" box="[1154,1212,1628,1653]" country="South Africa" name="Western Cape" pageId="7" pageNumber="508">Cape</collectingRegion>
|
||
by the Indian ships, they, as well as many other
|
||
<collectingRegion id="49DBF874CF57102B1987ED46FD73F93B" box="[467,525,1663,1688]" country="South Africa" name="Western Cape" pageId="7" pageNumber="508">Cape</collectingRegion>
|
||
animals, are often called Indian.” Also the morphology of the Lund
|
||
<typeStatus id="54A48834CF57102B1D4DEDB9FA32F93B" box="[1305,1356,1664,1688]" pageId="7" pageNumber="508">type</typeStatus>
|
||
, which has pectoral scutes in broad midline contact, is typical for South African terrapins (
|
||
<bibRefCitation id="EF8E4B67CF57102B1C04ED9DFB9DF91E" author="Hewitt, J." box="[1104,1251,1700,1725]" pageId="7" pageNumber="508" pagination="283 - 357" refId="ref12349" refString="Hewitt, J. (1935) Some new forms of batrachians and reptiles from South Africa. Records of the Albany Museum, 4, 283 - 357, plates XXVII - XXXVI." type="journal article" year="1935">Hewitt 1935</bibRefCitation>
|
||
: p. 325; unpubl. observ.), and is thus in agreement with an origin near
|
||
<collectingRegion id="49DBF874CF57102B1B47EDFEFC33F943" box="[787,845,1735,1760]" country="South Africa" name="Western Cape" pageId="7" pageNumber="508">Cape</collectingRegion>
|
||
Town. Therefore, we accept Hewitt’s
|
||
<typeStatus id="54A48834CF57102B1D59EDF1FA43F943" box="[1293,1341,1736,1760]" pageId="7" pageNumber="508">type</typeStatus>
|
||
locality designation and identify
|
||
<taxonomicName id="4C1F4D15CF57102B1997EDD5FD02F8A6" authorityName="Schoepff" authorityYear="1792" box="[451,636,1772,1797]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF57102B1997EDD5FD02F8A6" box="[451,636,1772,1797]" italics="true" pageId="7" pageNumber="508">Testudo galeata</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C1F4D15CF57102B1A9FEDD5FC2AF8A6" box="[715,852,1772,1797]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA84CF57102B1A9FEDD5FC2AF8A6" box="[715,852,1772,1797]" italics="true" pageId="7" pageNumber="508">Pelomedusa</emphasis>
|
||
</taxonomicName>
|
||
lineage IX, distributed throughout
|
||
<collectingCountry id="F3087606CF57102B1D51EDD5FAE2F8A6" box="[1285,1436,1772,1797]" name="South Africa" pageId="7" pageNumber="508">South Africa</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8E4B67CF57102B18CAEC36FEA9F88B" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[158,471,1807,1832]" pageId="7" pageNumber="508" pagination="428 - 440" refId="ref13781" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="B96BEA84CF57102B190FEC28FEEDF88B" box="[347,403,1807,1832]" italics="true" pageId="7" pageNumber="508">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA03696CF57102B1893EC0DFE26F81A" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="508">
|
||
Two fresh
|
||
<taxonomicName id="4C1F4D15CF57102B1969EC0DFEB8F8EE" box="[317,454,1844,1869]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA84CF57102B1969EC0DFEB8F8EE" box="[317,454,1844,1869]" italics="true" pageId="7" pageNumber="508">Pelomedusa</emphasis>
|
||
</taxonomicName>
|
||
samples collected close to the
|
||
<collectingRegion id="49DBF874CF57102B1B4AEC0DFC26F8EE" box="[798,856,1844,1869]" country="South Africa" name="Western Cape" pageId="7" pageNumber="508">Cape</collectingRegion>
|
||
of Good Hope (MTD T 5484, Swellendam District; MTD T 5897, Chelance; Table S1) can be regarded as topotypes of
|
||
<taxonomicName id="4C1F4D15CF57102B1BEFEC6EFB0EF8D3" authorityName="Schoepff" authorityYear="1792" box="[955,1136,1879,1904]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF57102B1BEFEC6EFB0EF8D3" box="[955,1136,1879,1904]" italics="true" pageId="7" pageNumber="508">Testudo galeata</emphasis>
|
||
</taxonomicName>
|
||
. As expected, their DNA sequences (
|
||
<bibRefCitation id="EF8E4B67CF57102B194DEC45FD28F836" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[281,598,1916,1941]" pageId="7" pageNumber="508" pagination="428 - 440" refId="ref13781" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="B96BEA84CF57102B198CEC44FD6CF836" box="[472,530,1916,1941]" italics="true" pageId="7" pageNumber="508">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
; this study) are assigned with high support to lineage IX in phylogenetic analyses (
|
||
<figureCitation id="13242A13CF57102B1951EC99FE38F81A" box="[261,326,1952,1977]" captionStart="FIGURE 1" captionStartId="5.[151,250,1100,1122]" captionTargetBox="[157,1429,201,1075]" captionTargetId="figure-427@5.[151,1436,195,1080]" captionTargetPageId="5" captionText="FIGURE 1. Maximum Likelihood tree for Pelomedusa, based on 1848 bp of mitochondrial DNA (12S, cyt b, ND4+tRNAs), rooted with Pelusios sinuatus. The tree topology completely matches the one inferred by Bayesian analyses. Numbers along nodes indicate bootstrap support and Bayesian posterior probabilities greater than 50 or 0.95, respectively (not shown for some terminal clades with short branches). Sample codes are voucher numbers; see Table S1 for explanation. Name-bearing types (red) and paralectotypes or topotypic samples (blue) highlighted. On the right, mitochondrial lineages of Vargas-Ramírez et al. (2010) indicated. No topotypic material of Pelomedusa galeata orangensis Hewitt, 1935 could be sequenced. However, the lost holotype is thought to come from the vicinity of Kimberley (South Africa) and all South African samples belong to lineage IX, so it can be concluded that Pelomedusa galeata orangensis represents the same lineage. Inset: Helmeted terrapin from the Addo Elephant National Park, South Africa. Photo: M. Gregorič." figureDoi="http://doi.org/10.5281/zenodo.4915066" httpUri="https://zenodo.org/record/4915066/files/figure.png" pageId="7" pageNumber="508">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF60661ECF58102418C3ED04FDECF933" ID-DOI="http://doi.org/10.5281/zenodo.4915070" ID-Zenodo-Dep="4915070" httpUri="https://zenodo.org/record/4915070/files/figure.png" pageId="8" pageNumber="509" startId="8.[151,250,1597,1619]" targetBox="[224,1354,222,1560]" targetPageId="8">
|
||
<paragraph id="8BA03696CF58102418C3ED04FDECF933" blockId="8.[151,1436,1597,2017]" pageId="8" pageNumber="509">
|
||
<emphasis id="B96BEA84CF58102418C3ED04FE63F9F1" bold="true" box="[151,285,1597,1619]" pageId="8" pageNumber="509">FIGURE 3.</emphasis>
|
||
Top, left: Folio from Schoepff’s legacy (Museum für Naturkunde zu Berlin, Historische Bild- und Schriftgutsammlungen, SI, Nachl. Schoepff II) consisting of the drawings provided by Retzius and an imprint of the resultant etching. Retzius’ hand-written German text reads:
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA03696CF58102418C3EDA0FDE9F8A8" blockId="8.[151,1436,1597,2017]" pageId="8" pageNumber="509">
|
||
“Die Skizze der
|
||
<taxonomicName id="4C1F4D15CF5810241913EDA3FED6F90C" box="[327,424,1689,1711]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="scabra">
|
||
<emphasis id="B96BEA84CF5810241913EDA3FED6F90C" box="[327,424,1689,1711]" italics="true" pageId="8" pageNumber="509">T. scabra</emphasis>
|
||
</taxonomicName>
|
||
welche ein Student gemacht hat, lege ich so fehlerhaft sie auch ist bey, indess aus Nr. 1 forma scutellorum u. aus Nr. 2 die Gestalt des Kopfes und der Hinterfüsse wohl zu ersehen ist. Dass meine
|
||
<taxonomicName id="4C1F4D15CF5810241C2BED8EFBB3F96E" authorityName="Linnaeus" authorityYear="1758" box="[1151,1229,1719,1741]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="509" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA84CF5810241C2BED8EFBB3F96E" box="[1151,1229,1719,1741]" italics="true" pageId="8" pageNumber="509">Testudo</emphasis>
|
||
</taxonomicName>
|
||
die wahre
|
||
<taxonomicName id="4C1F4D15CF5810241D6FED8EFAFEF96E" box="[1339,1408,1719,1741]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="scabra">
|
||
<emphasis id="B96BEA84CF5810241D6FED8EFAFEF96E" box="[1339,1408,1719,1741]" italics="true" pageId="8" pageNumber="509">scabra</emphasis>
|
||
</taxonomicName>
|
||
L. ist, dessen bin ich gewiss, ob er [meant is Linnaeus] gleich sagt [illegible] er hat aber ganz gewiss Nr. 73 u. 74 [illegible] completirt, wie das oft der Fall ist.―Rezius” [sic].
|
||
</paragraph>
|
||
<paragraph id="8BA03696CF58102418C3EC2AFE12F826" blockId="8.[151,1436,1597,2017]" pageId="8" pageNumber="509">
|
||
English translation: “I enclose the drawing of the
|
||
<taxonomicName id="4C1F4D15CF5810241ACFEC2DFD84F88A" box="[667,762,1811,1833]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="scabra">
|
||
<emphasis id="B96BEA84CF5810241ACFEC2DFD84F88A" box="[667,762,1811,1833]" italics="true" pageId="8" pageNumber="509">T. scabra</emphasis>
|
||
</taxonomicName>
|
||
made by a student, as flawed as it is, since no. 1 shows well the shape of the scutes and no. 2 the shape of the head and of the hind legs. I am sure that my
|
||
<taxonomicName id="4C1F4D15CF5810241C6FEC0BFBF7F8EB" authorityName="Linnaeus" authorityYear="1758" box="[1083,1161,1842,1864]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="509" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA84CF5810241C6FEC0BFBF7F8EB" box="[1083,1161,1842,1864]" italics="true" pageId="8" pageNumber="509">Testudo</emphasis>
|
||
</taxonomicName>
|
||
is the true
|
||
<taxonomicName id="4C1F4D15CF5810241CA8EC0BFA3FF8EB" box="[1276,1345,1842,1864]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="scabra">
|
||
<emphasis id="B96BEA84CF5810241CA8EC0BFA3FF8EB" box="[1276,1345,1842,1864]" italics="true" pageId="8" pageNumber="509">scabra</emphasis>
|
||
</taxonomicName>
|
||
L., even though he [meant is Linnaeus] says [illegible] he has surely no. 73 and 74 [illegible] completed, as this is often the case.―Rezius” [sic].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305651DCF58102418C3ECB6FB11F807" box="[151,1135,1934,1956]" pageId="8" pageNumber="509" type="reference_group">
|
||
<paragraph id="8BA03696CF58102418C3ECB6FB11F807" blockId="8.[151,1436,1597,2017]" box="[151,1135,1934,1956]" pageId="8" pageNumber="509">
|
||
Top, right: reproduction of figure 1, plate III from
|
||
<bibRefCitation id="EF8E4B67CF5810241AC1ECB7FC44F807" author="Schoepff, I. D." box="[661,826,1934,1956]" pageId="8" pageNumber="509" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff (1792)</bibRefCitation>
|
||
showing “
|
||
<taxonomicName id="4C1F4D15CF5810241BFCECB6FB1FF807" authority="Retz." authorityName="Retz." box="[936,1121,1934,1956]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="scabra">
|
||
<emphasis id="B96BEA84CF5810241BFCECB6FB5FF807" box="[936,1057,1934,1956]" italics="true" pageId="8" pageNumber="509">Test. scabra</emphasis>
|
||
Retz.
|
||
</taxonomicName>
|
||
”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305651DCF58102418C3EC97FDFFF842" pageId="8" pageNumber="509" type="materials_examined">
|
||
<paragraph id="8BA03696CF58102418C3EC97FDFFF842" blockId="8.[151,1436,1597,2017]" pageId="8" pageNumber="509">
|
||
<materialsCitation id="3B773CCBCF58102418C3EC97FD03F842" ID-GBIF-Occurrence="3319793304" collectionCode="ZMUL" pageId="8" pageNumber="509" specimenCode="ZMUL 6481" specimenCount="1" typeStatus="lectotype">
|
||
Bottom: Dorsal and ventral view of the
|
||
<typeStatus id="54A48834CF5810241A60EC94FDEFF860" box="[564,657,1965,1987]" pageId="8" pageNumber="509" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1F4D15CF5810241AE1EC94FC84F860" authority="Schoepff, 1792" authorityName="Schoepff" authorityYear="1792" box="[693,1018,1965,1987]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF5810241AE1EC94FC29F860" box="[693,855,1965,1987]" italics="true" pageId="8" pageNumber="509">Testudo galeata</emphasis>
|
||
<bibRefCitation id="EF8E4B67CF5810241B0AEC94FC84F860" author="Schoepff, I. D." box="[862,1018,1965,1987]" pageId="8" pageNumber="509" refId="ref13074" refString="Schoepff, I. D. (1792) Historia Testudinum iconibus illustrata. Ioannes Iacobus Palm, Erlangen, XII + 136 pp., 31 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5109" type="book" year="1792">Schoepff, 1792</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<specimenCode id="DBB99EEDCF5810241C5DEC94FBF1F860" box="[1033,1167,1965,1987]" collectionCode="ZMUL" country="Sweden" httpUri="http://grbio.org/cool/wjrm-zrx5" name="Universitetets Lund, Zoologiska Museet" pageId="8" pageNumber="509">ZMUL 6481</specimenCode>
|
||
, straight carapacial length
|
||
<quantity id="4CE79B73CF58102418C3ECF2FF92F842" box="[151,236,1995,2017]" metricMagnitude="-2" metricUnit="m" metricValue="5.84" pageId="8" pageNumber="509" unit="cm" value="5.84">5.84 cm</quantity>
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||
).
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||
<collectorName id="26EA5340CF58102418A9ECF2FE4AF842" box="[253,308,1995,2017]" pageId="8" pageNumber="509">Scale</collectorName>
|
||
bar,
|
||
<quantity id="4CE79B73CF5810241932ECF2FEE5F842" box="[358,411,1995,2017]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="509" unit="cm" value="3.0">3 cm</quantity>
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||
.
|
||
<collectorName id="26EA5340CF58102419F1ECF2FE90F842" box="[421,494,1995,2017]" pageId="8" pageNumber="509">Photos</collectorName>
|
||
:
|
||
<collectorName id="26EA5340CF58102419A3ECF5FD03F842" box="[503,637,1995,2017]" pageId="8" pageNumber="509">L. Lundqvist</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |