treatments-xml/data/D9/67/0F/D9670F02E3A85A71953F42B11CFD350C.xml
2024-06-21 12:53:51 +02:00

284 lines
22 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1137.97068" ID-Pensoft-Pub="1313-2970-1137-133" ID-Pensoft-UUID="818ADEB99A7A5D0C851421411FA5B421" ID-ZooBank="DFA4D0BDCBA649D487ACC741E6C216E9" ModsDocID="1313-2970-1137-133" checkinTime="1671892165935" checkinUser="pensoft" docAuthor="Caterino, Michael S." docDate="2022" docId="D9670F02E3A85A71953F42B11CFD350C" docLanguage="en" docName="ZooKeys 1137: 133-175" docOrigin="ZooKeys 1137" docPubDate="2022-12-22" docSource="http://dx.doi.org/10.3897/zookeys.1137.97068" docTitle="Euconnus vexillus Caterino 2022, sp. nov." docType="treatment" docUuid="47ED180B-4423-46AC-8CA4-3BB1FEDCDB19" docUuidSource="ZooBank" docVersion="1" id="818ADEB99A7A5D0C851421411FA5B421" lastPageNumber="133" masterDocId="818ADEB99A7A5D0C851421411FA5B421" masterDocTitle="First report of the Euconnus Thomson subgenus Cladoconnus Reitter in the New World, represented by thirteen new Appalachian species (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="175" masterPageNumber="133" pageNumber="133" updateTime="1671892165935" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>First report of the Euconnus Thomson subgenus Cladoconnus Reitter in the New World, represented by thirteen new Appalachian species (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caterino, Michael S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2597-5707</mods:nameIdentifier>
<mods:affiliation>Dept of Plant &amp; Environmental Sciences, Clemson University, Clemson, SC 29634, USA</mods:affiliation>
<mods:nameIdentifier type="email">mcateri@clemson.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1137</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>133</mods:start>
<mods:end>175</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1137.97068</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1137.97068</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1137-133</mods:identifier>
<mods:identifier type="ZooBank">DFA4D0BDCBA649D487ACC741E6C216E9</mods:identifier>
<mods:identifier type="Pensoft-UUID">818ADEB99A7A5D0C851421411FA5B421</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:47ED180B-4423-46AC-8CA4-3BB1FEDCDB19" httpUri="http://treatment.plazi.org/id/D9670F02E3A85A71953F42B11CFD350C" lastPageNumber="133" pageId="0" pageNumber="133">
<subSubSection pageId="0" pageNumber="133" type="nomenclature">
<paragraph pageId="0" pageNumber="133">
<taxonomicName LSID="https://zoobank.org/47ED180B-4423-46AC-8CA4-3BB1FEDCDB19" authority="Caterino, 2022" authorityName="Caterino" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euconnus vexillus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="vexillus" status="sp. nov.">Euconnus vexillus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="133">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="description">
<paragraph pageId="0" pageNumber="133">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus A Euconnus megalops B Euconnus vexillus C Euconnus cumberlandus D Euconnus vetustus." figureDoi="10.3897/zookeys.1137.97068.figure2" httpUri="https://binary.pensoft.net/fig/786413" pageId="0" pageNumber="133">Figs 2B</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus and character photos of Euconnus vexillus A male, dorsal view B female, dorsal view C SEM of mouthparts, ventral view D SEM of female head, lateral E SEM of male antenna, lateral F SEM of elytra and pronotum G SEM of meta- and mesoventrites." figureDoi="10.3897/zookeys.1137.97068.figure4" httpUri="https://binary.pensoft.net/fig/786415" pageId="0" pageNumber="133">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Map of collecting records for Euconnus (Cladoconnus) vexillus (squares), E. (C.) vetustus (stars), and E. (C.) adversus (circles)." figureDoi="10.3897/zookeys.1137.97068.figure5" httpUri="https://binary.pensoft.net/fig/786416" pageId="0" pageNumber="133">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="type material">
<paragraph pageId="0" pageNumber="133">Type material.</paragraph>
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" italics="true" pageId="0" pageNumber="133">Holotype</emphasis>
♂, deposited in FMNH: &quot;USA:SC: Greenville Co.
<geoCoordinate degrees="35.1523" direction="north" orientation="latitude" precision="5" value="35.1523">35.1523°N</geoCoordinate>
,
<geoCoordinate degrees="82.2814" direction="west" orientation="longitude" precision="5" value="-82.2814">82.2814°W</geoCoordinate>
, Chestnut Ridge Heritage Preserve, vi.05.2015, S. Myers, Hardwood litter&quot; / &quot;[QR code] CLEMSON ENT CUAC000026944&quot;.
<emphasis bold="true" italics="true" pageId="0" pageNumber="133">Paratypes</emphasis>
(10, CUAC, FMNH) - 2 ♂, 2 ♀: same data as type; 1 ♂, 1 ♀: &quot;USA:SC: Greenville Co.
<geoCoordinate degrees="35.1518" direction="north" orientation="latitude" precision="5" value="35.1518">35.1518°N</geoCoordinate>
,
<geoCoordinate degrees="82.2839" direction="west" orientation="longitude" precision="5" value="-82.2839">82.2839°W</geoCoordinate>
, Chestnut Ridge Heritage Preserve, vi.05.2015, S. Myers, Hardwood litter&quot;; 1 ♀: &quot;USA:SC: Greenville Co.
<geoCoordinate degrees="35.1501" direction="north" orientation="latitude" precision="5" value="35.1501">35.1501°N</geoCoordinate>
,
<geoCoordinate degrees="82.2820" direction="west" orientation="longitude" precision="5" value="-82.282">82.2820°W</geoCoordinate>
, Chestnut Ridge Heritage Preserve, vi.05.2015, S. Myers, Hardwood litter&quot;; 2 ♀: &quot;USA:SC: Greenville Co.
<geoCoordinate degrees="35.1406" direction="north" orientation="latitude" precision="5" value="35.1406">35.1406°N</geoCoordinate>
,
<geoCoordinate degrees="82.2790" direction="west" orientation="longitude" precision="5" value="-82.279">82.2790°W</geoCoordinate>
, Chestnut Ridge Heritage Preserve, vi.05.2015, S. Myers, Hardwood litter&quot;; 1 ♀: &quot;USA:SC: Greenville Co.
<geoCoordinate degrees="35.1506" direction="north" orientation="latitude" precision="5" value="35.1506">35.1506°N</geoCoordinate>
,
<geoCoordinate degrees="82.2799" direction="west" orientation="longitude" precision="5" value="-82.2799">82.2799°W</geoCoordinate>
Chestnut Ridge Heritage Preserve, iv.08.2018, M. Caterino &amp; L.
<normalizedToken originalValue="Vásquez-Vélez">Vasquez-Velez</normalizedToken>
, sifted litter&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="materials_examined">
<paragraph pageId="0" pageNumber="133">Other material.</paragraph>
<paragraph pageId="0" pageNumber="133">
(26)
<materialsCitation collectingDate="1971-07-23" county="Mercer Co." location="Camp Creek State Forest" specimenCount="15" specimenCount-female="11" specimenCount-male="4">
<emphasis bold="true" pageId="0" pageNumber="133">WV</emphasis>
:
<collectingCounty>Mercer Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:D9670F02E3A85A71953F42B11CFD350C:14C625D27B772E4C2EF84296A8F04CF8" county="Mercer Co." name="Camp Creek State Forest">Camp Creek State Forest</location>
,
<collectingDate value="1971-07-23">23-Jul-1971</collectingDate>
, leaf litter (
<specimenCount type="male">4 ♂</specimenCount>
,
<specimenCount type="female">11 ♀</specimenCount>
)
</materialsCitation>
; '
<materialsCitation collectingDate="1901-10" location="Black Mts'" specimenCount="3" specimenCount-male="3">
<location LSID="urn:lsid:plazi:treatment:D9670F02E3A85A71953F42B11CFD350C:EC60ACF9CCA0C41FB6A6A8D35E53B796" name="Black Mts'">
Black
<normalizedToken originalValue="Mts">Mts'</normalizedToken>
</location>
,
<collectingDate value="1901-10">x.1901</collectingDate>
(
<specimenCount type="male">3 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2021-05-17" county="Caldwell Co." elevation="1225" location="Grandfather Mt. State Park" municipality="Nuwati Trail" specimenCount="1" specimenCount-male="1">
<emphasis bold="true" pageId="0" pageNumber="133">NC</emphasis>
:
<collectingCounty>Caldwell Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:D9670F02E3A85A71953F42B11CFD350C:418E8E7AFE688504F1272AB919825232" county="Caldwell Co." municipality="Nuwati Trail" name="Grandfather Mt. State Park">Grandfather Mt. State Park</location>
,
<collectingMunicipality>Nuwati Trail</collectingMunicipality>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.225296" unit="ft" value="4020.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.225296" unit="ft" value="4020.0">4020 ft.</elevation>
</quantity>
,
<collectingDate value="2021-05-17">17-May-2021</collectingDate>
(
<specimenCount type="male">1 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2015-08-25" county="McDowell Co." elevation="1046" location="Pisgah National Forest" municipality="Mountain Trail" specimenCount="2" specimenCount-male="2">
<collectingCounty>McDowell Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:D9670F02E3A85A71953F42B11CFD350C:79274BF63EEEA576A96DF3291C688279" county="McDowell Co." municipality="Mountain Trail" name="Pisgah National Forest">Pisgah National Forest</location>
, Mackey
<collectingMunicipality>Mountain Trail</collectingMunicipality>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0463784" unit="ft" value="3433.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.0463784" unit="ft" value="3433.0">3433 ft.</elevation>
</quantity>
,
<collectingDate value="2015-08-25">25-Aug-2015</collectingDate>
(
<specimenCount type="male">2 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2015-08-25" county="McDowell Co." elevation="609" location="Pisgah National Forest" municipality="Snooks Nose Trail" specimenCount="4" specimenCount-female="3" specimenCount-male="1">
<collectingCounty>McDowell Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:D9670F02E3A85A71953F42B11CFD350C:4A3F0D1D43EB22A73DFE12D0B46AC914" county="McDowell Co." municipality="Snooks Nose Trail" name="Pisgah National Forest">Pisgah National Forest</location>
,
<collectingMunicipality>Snooks Nose Trail</collectingMunicipality>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.089904" unit="ft" value="1998.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.089904" unit="ft" value="1998.0">1998 ft.</elevation>
</quantity>
,
<collectingDate value="2015-08-25">25-Aug-2015</collectingDate>
(
<specimenCount type="female">3 ♀</specimenCount>
,
<specimenCount type="male">1 ♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation collectingDate="2018-03-18" county="Polk Co." elevation="530" location="Green River" specimenCount="1" specimenCount-male="1">
<collectingCounty>Polk Co.</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:D9670F02E3A85A71953F42B11CFD350C:9AA07F8F84093642AD080A59ACE463EE" county="Polk Co." name="Green River">Green River</location>
Game Lands,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.30352" unit="ft" value="1740.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.30352" unit="ft" value="1740.0">1740 ft.</elevation>
</quantity>
,
<collectingDate value="2018-03-18">18-Mar-2018</collectingDate>
(
<specimenCount type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="description">
<paragraph pageId="0" pageNumber="133">Diagnostic description.</paragraph>
<paragraph pageId="0" pageNumber="133">
This species is very similar to the preceding species and can best be distinguished by male genitalic and antennal characters. Like the preceding, males are winged, while females appear not to be. A few noteworthy external differences can also be cited: males with distinct carinae on antennomeres VIII and IX (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus and character photos of Euconnus vexillus A male, dorsal view B female, dorsal view C SEM of mouthparts, ventral view D SEM of female head, lateral E SEM of male antenna, lateral F SEM of elytra and pronotum G SEM of meta- and mesoventrites." figureDoi="10.3897/zookeys.1137.97068.figure4" httpUri="https://binary.pensoft.net/fig/786415" pageId="0" pageNumber="133">4E</figureCitation>
); female antennae less distinctly tetramerous, with antennomere VIII intermediate in size between antennomeres VII and IX (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus and character photos of Euconnus vexillus A male, dorsal view B female, dorsal view C SEM of mouthparts, ventral view D SEM of female head, lateral E SEM of male antenna, lateral F SEM of elytra and pronotum G SEM of meta- and mesoventrites." figureDoi="10.3897/zookeys.1137.97068.figure4" httpUri="https://binary.pensoft.net/fig/786415" pageId="0" pageNumber="133">4D</figureCitation>
); body darker, stout, with only faintly rufescent elytral humeri, most distinctly lighter along posterior half of elytral suture; male eyes smaller, less protuberant, with only ~ 25 ommatidia; median basal carina of pronotum slightly better developed to base (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus and character photos of Euconnus vexillus A male, dorsal view B female, dorsal view C SEM of mouthparts, ventral view D SEM of female head, lateral E SEM of male antenna, lateral F SEM of elytra and pronotum G SEM of meta- and mesoventrites." figureDoi="10.3897/zookeys.1137.97068.figure4" httpUri="https://binary.pensoft.net/fig/786415" pageId="0" pageNumber="133">4F</figureCitation>
); aedeagus (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus A Euconnus megalops B Euconnus vexillus C Euconnus cumberlandus D Euconnus vetustus." figureDoi="10.3897/zookeys.1137.97068.figure2" httpUri="https://binary.pensoft.net/fig/786413" pageId="0" pageNumber="133">2B</figureCitation>
) with median lobe broadly truncate at apex; parameres short, tapered, bearing three apical setae; compressor plate strongly asymmetrical, rather short; endophallic armature with strong pair of upper processes: the left long and strongly hooked apically, with a thin inner laminar blade, its inner edge concave; the right shorter, more strongly curved toward middle of longer process; lower endophallic armature consisting of three hook-like processes of varying lengths, the lateral-most rather short and strongly curved, the medial-most of intermediate length, the one between them the longest, its tip just visible between bases of upper processes (in
<normalizedToken originalValue="dorsal">'dorsal'</normalizedToken>
view).
</paragraph>
<caption doi="10.3897/zookeys.1137.97068.figure4" httpUri="https://binary.pensoft.net/fig/786415" pageId="0" pageNumber="133" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" pageId="0" pageNumber="133">Figure 4.</emphasis>
Habitus and character photos of
<taxonomicName authorityName="Caterino" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euconnus vexillus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="vexillus">
<emphasis italics="true" pageId="0" pageNumber="133">Euconnus vexillus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="133">A</emphasis>
male, dorsal view
<emphasis bold="true" pageId="0" pageNumber="133">B</emphasis>
female, dorsal view
<emphasis bold="true" pageId="0" pageNumber="133">C</emphasis>
SEM of mouthparts, ventral view
<emphasis bold="true" pageId="0" pageNumber="133">D</emphasis>
SEM of female head, lateral
<emphasis bold="true" pageId="0" pageNumber="133">E</emphasis>
SEM of male antenna, lateral
<emphasis bold="true" pageId="0" pageNumber="133">F</emphasis>
SEM of elytra and pronotum
<emphasis bold="true" pageId="0" pageNumber="133">G</emphasis>
SEM of meta- and mesoventrites.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1137.97068.figure5" httpUri="https://binary.pensoft.net/fig/786416" pageId="0" pageNumber="133" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="133">
<emphasis bold="true" pageId="0" pageNumber="133">Figure 5.</emphasis>
Map of collecting records for
<taxonomicName authorityName="Caterino" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Euconnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euconnus (Cladoconnus) vexillus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="vexillus" subGenus="Cladoconnus">Euconnus (Cladoconnus) vexillus</taxonomicName>
(squares),
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. (C.) vetustus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="vetustus" subGenus="C.">E. (C.) vetustus</taxonomicName>
(stars), and
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. (C.) adversus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="adversus" subGenus="C.">E. (C.) adversus</taxonomicName>
(circles).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="distribution">
<paragraph pageId="0" pageNumber="133">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="133">This species has been found at several widely scattered localities east and northeast of the Asheville Depression, from the headwaters of the French Broad near the southern end of its range, at Chestnut Ridge, South Carolina, to southern West Virginia in the northeast. Most of its occurrences are at middle elevations, from 1090 to 4020 ft, most below 2000.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="133" type="remarks">
<paragraph pageId="0" pageNumber="133">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="133">
This species is similar and, judging by male genitalia, closely related to
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. megalops" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="megalops">
<emphasis italics="true" pageId="0" pageNumber="133">E. megalops</emphasis>
</taxonomicName>
. Both have similar opposing hooked upper endophallic sclerites, though the shorter right one (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aedeagus A Euconnus megalops B Euconnus vexillus C Euconnus cumberlandus D Euconnus vetustus." figureDoi="10.3897/zookeys.1137.97068.figure2" httpUri="https://binary.pensoft.net/fig/786413" pageId="0" pageNumber="133">2B</figureCitation>
) of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. vexillus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="vexillus">
<emphasis italics="true" pageId="0" pageNumber="133">E. vexillus</emphasis>
</taxonomicName>
is distinctive. Moreover, the lower trio of endophallic sclerites are quite distinct in
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. vexillus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="vexillus">
<emphasis italics="true" pageId="0" pageNumber="133">E. vexillus</emphasis>
</taxonomicName>
, with three hooks projecting distad to varying lengths. In males, the carinae of antennomeres VII and IX will immediately distinguish this species from
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. megalops" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="megalops">
<emphasis italics="true" pageId="0" pageNumber="133">E. megalops</emphasis>
</taxonomicName>
, and the only weakly tetramerous club of females can be used to distinguish most of those, although they are similar to darker females of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="E. adversus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="adversus">
<emphasis italics="true" pageId="0" pageNumber="133">E. adversus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="133">
This species name means 'standard (or flag)-
<normalizedToken originalValue="bearer">bearer'</normalizedToken>
, referring to its possession of
<taxonomicName authorityName="Reitter" authorityYear="1909" class="Insecta" family="Staphylinidae" genus="Cladoconnus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cladoconnus" order="Coleoptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="133">Cladoconnus</emphasis>
</taxonomicName>
-typical carinae, borne proudly on its antennae.
</paragraph>
</subSubSection>
</treatment>
</document>