176 lines
22 KiB
XML
176 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e86008" ID-Pensoft-Pub="1864-8312-80-541" ID-Pensoft-UUID="92A8EB05D4DB5AD3B9FE4382897F3E60" ID-ZooBank="09338026092F42FB8E2F00DD6B868D49" ModsDocID="1864-8312-80-541" checkinTime="1664443249919" checkinUser="pensoft" docAuthor="Magalhaes, Ivan L. F., Perez-Gonzalez, Abel, Labarque, Facundo M., Carboni, Martin, Hammel, Joerg U., Kunz, Robin, Ramirez, Martin J. & Solorzano-Kraemer, Monica M." docDate="2022" docId="C53376FFCED659C0817D78BC31CBAB26" docLanguage="en" docName="ArthropodSystPhyl 80: 541-559" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-09-28" docSource="http://dx.doi.org/10.3897/asp.80.e86008" docTitle="Drymusa aculicaput new combination" docType="treatment" docVersion="1" id="92A8EB05D4DB5AD3B9FE4382897F3E60" lastPageNumber="541" masterDocId="92A8EB05D4DB5AD3B9FE4382897F3E60" masterDocTitle="Revision of recluse spiders (Araneae: Sicariidae: Loxosceles) preserved in Dominican amber and a total-evidence phylogeny of Scytodoidea reveal the first fossil Drymusidae" masterLastPageNumber="559" masterPageNumber="541" pageNumber="541" updateTime="1664443249919" updateUser="pensoft">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Revision of recluse spiders (Araneae: Sicariidae: Loxosceles) preserved in Dominican amber and a total-evidence phylogeny of Scytodoidea reveal the first fossil Drymusidae</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Magalhaes, Ivan L. F.</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3728-3270</mods:nameIdentifier>
|
|
<mods:affiliation>Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Av. Angel Gallardo 470, C 1405 DJR, Buenos Aires, Argentina</mods:affiliation>
|
|
<mods:nameIdentifier type="email">ivanlfmagalhaes@yahoo.com.br</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Perez-Gonzalez, Abel</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6744-6811</mods:nameIdentifier>
|
|
<mods:affiliation>Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Av. Angel Gallardo 470, C 1405 DJR, Buenos Aires, Argentina</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Labarque, Facundo M.</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9413-1949</mods:nameIdentifier>
|
|
<mods:affiliation>Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de Sao Carlos (UFSCar), campus Sao Carlos, Rodovia Washington Luis, Km 235, CEP 13565 - 905, Sao Carlos, SP, Brazil</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Carboni, Martin</mods:namePart>
|
|
<mods:affiliation>Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Av. Angel Gallardo 470, C 1405 DJR, Buenos Aires, Argentina</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Hammel, Joerg U.</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6744-6811</mods:nameIdentifier>
|
|
<mods:affiliation>Institute of Materials Physics, Helmholtz-Zentrum Hereon, Outstation at DESY, Max-Planck-Str. 1, D- 21502 Geesthacht, Germany</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Kunz, Robin</mods:namePart>
|
|
<mods:affiliation>Department of Palaeontology and Historical Geology, Senckenberg Research Institute and Natural History Museum, D- 60325 Frankfurt am Main, Germany</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Ramirez, Martin J.</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0358-0130</mods:nameIdentifier>
|
|
<mods:affiliation>Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", Consejo Nacional de Investigaciones Cientificas y Tecnicas (CONICET), Av. Angel Gallardo 470, C 1405 DJR, Buenos Aires, Argentina</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Solorzano-Kraemer, Monica M.</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3065-119X</mods:nameIdentifier>
|
|
<mods:affiliation>Department of Palaeontology and Historical Geology, Senckenberg Research Institute and Natural History Museum, D- 60325 Frankfurt am Main, Germany</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2022</mods:date>
|
|
<mods:detail type="pubDate">
|
|
<mods:number>2022-09-28</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="volume">
|
|
<mods:number>80</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>541</mods:start>
|
|
<mods:end>559</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/asp.80.e86008</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e86008</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1864-8312-80-541</mods:identifier>
|
|
<mods:identifier type="ZooBank">09338026092F42FB8E2F00DD6B868D49</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">92A8EB05D4DB5AD3B9FE4382897F3E60</mods:identifier>
|
|
</mods:mods>
|
|
<subSection pageId="0" pageNumber="541" type="5. taxonomy">
|
|
<treatment LSID="urn:lsid:plazi:treatment:C53376FFCED659C0817D78BC31CBAB26" httpUri="http://treatment.plazi.org/id/C53376FFCED659C0817D78BC31CBAB26" lastPageNumber="541" pageId="0" pageNumber="541">
|
|
<subSubSection pageId="0" pageNumber="541" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="541">
|
|
5.2.1.
|
|
<taxonomicName LSID="C53376FF-CED6-59C0-817D-78BC31CBAB26" authority="(Wunderlich, 2004) new combination" authorityName="new combination" baseAuthorityName="Wunderlich" baseAuthorityYear="2004" class="Arachnida" family="Sicariidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa aculicaput" order="Araneae" pageId="0" pageNumber="541" phylum="Arthropoda" rank="species" species="aculicaput">Drymusa aculicaput (Wunderlich, 2004) new combination</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="541" type="description">
|
|
<paragraph pageId="0" pageNumber="541">
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), photographs under light microscopy. A, B habitus, dorsolateral. C right femur I. Arrows to anterior macrosetae. D spinnerets, dorsolateral view. E, F, H right tarsi (E: III-IV, F: II, H: I), prolateral. Arrows to third claw. Note highly articulated podotarsite. G Chelicerae and palp, subdorsal view. Arrow to tip of right embolus. Scale bars: 1 mm (A), 0.5 mm (B), 0.2 mm (C), 0.1 mm (D, G), 0.05 mm (E, F, H)." figureDoi="10.3897/asp.80.e86008.figure3" httpUri="https://binary.pensoft.net/fig/749997" pageId="0" pageNumber="541">Figs 3</figureCitation>
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. A, B habitus, dorsal. C habitus, ventral. Inset showing notched labium. D clypeus, subanterior. Scale bars: 1 mm (A), 0.5 mm (B-D)." figureDoi="10.3897/asp.80.e86008.figure4" httpUri="https://binary.pensoft.net/fig/749998" pageId="0" pageNumber="541">, 4</figureCitation>
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. Spinnerets (A - C), left palp (D - H), and right palp (I, J). A ventral. B apical. C dorsal. D subprolateral. E retrolateral. F prolateral. G bulb and cymbium, apical. H bulb and cymbium, dorsal. I prolateral. J retrolateral. Asterisks are artifacts in the rendering. Scale bars: 0.1 mm (only A-C, F, I to scale)." figureDoi="10.3897/asp.80.e86008.figure5" httpUri="https://binary.pensoft.net/fig/749999" pageId="0" pageNumber="541">, 5</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="541" type="reference_group">
|
|
<paragraph pageId="0" pageNumber="541">
|
|
<taxonomicName authorityName="Wunderlich" authorityYear="2004" class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Loxosceles aculicaput" order="Araneae" pageId="0" pageNumber="541" phylum="Bryophyta" rank="species" species="aculicaput">Loxosceles aculicaput</taxonomicName>
|
|
Wunderlich, 2004: 703, fig. 10b-d, photo 36.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="541" type="type material">
|
|
<paragraph pageId="0" pageNumber="541">Type material.</paragraph>
|
|
<paragraph pageId="0" pageNumber="541">Holotype adult male in Dominican amber, deposited in SMNG 07/36287-422, labeled F933/DB/AR/LOX/CJW, examined. No other specimens are known.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="541" type="preservation">
|
|
<paragraph pageId="0" pageNumber="541">Preservation.</paragraph>
|
|
<paragraph pageId="0" pageNumber="541">The spider is completely and well-preserved in an orange piece of amber. The abdomen is slightly shrunken and shriveled but no other structures are obviously deformed. The amber piece has large fissures in the portion ventral to the spider. The piece has been embedded in artificial resin. There are no syninclusions other than some pieces of unidentifiable plants and detritus.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="541" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="541">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="541">
|
|
<taxonomicName authorityName="new combination" baseAuthorityName="Wunderlich" baseAuthorityYear="2004" class="Arachnida" family="Sicariidae" genus="Drymusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymusa aculicaput" order="Araneae" pageId="0" pageNumber="541" phylum="Arthropoda" rank="species" species="aculicaput">
|
|
<emphasis italics="true" pageId="0" pageNumber="541">Drymusa aculicaput</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="541">comb. nov.</emphasis>
|
|
can be distinguished from other scytodoids preserved in Dominican amber, as well as from extant congeners, by the palp with short, incrassate tibia and by the bulb with a thin, straight, needle-like embolus (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. Spinnerets (A - C), left palp (D - H), and right palp (I, J). A ventral. B apical. C dorsal. D subprolateral. E retrolateral. F prolateral. G bulb and cymbium, apical. H bulb and cymbium, dorsal. I prolateral. J retrolateral. Asterisks are artifacts in the rendering. Scale bars: 0.1 mm (only A-C, F, I to scale)." figureDoi="10.3897/asp.80.e86008.figure5" httpUri="https://binary.pensoft.net/fig/749999" pageId="0" pageNumber="541">5E, G, H, J</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="541" type="description">
|
|
<paragraph pageId="0" pageNumber="541">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="541">
|
|
Male holotype (SMNG 07/36287-422).
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="541">Structure</emphasis>
|
|
: Carapace longer than wide, narrowed and more hirsute in the cephalic region. Six eyes in three dyads separated by 2-3 diameters (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. A, B habitus, dorsal. C habitus, ventral. Inset showing notched labium. D clypeus, subanterior. Scale bars: 1 mm (A), 0.5 mm (B-D)." figureDoi="10.3897/asp.80.e86008.figure4" httpUri="https://binary.pensoft.net/fig/749998" pageId="0" pageNumber="541">4D</figureCitation>
|
|
). Chelicera with basal article robust (Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), photographs under light microscopy. A, B habitus, dorsolateral. C right femur I. Arrows to anterior macrosetae. D spinnerets, dorsolateral view. E, F, H right tarsi (E: III-IV, F: II, H: I), prolateral. Arrows to third claw. Note highly articulated podotarsite. G Chelicerae and palp, subdorsal view. Arrow to tip of right embolus. Scale bars: 1 mm (A), 0.5 mm (B), 0.2 mm (C), 0.1 mm (D, G), 0.05 mm (E, F, H)." figureDoi="10.3897/asp.80.e86008.figure3" httpUri="https://binary.pensoft.net/fig/749997" pageId="0" pageNumber="541">3G</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. A, B habitus, dorsal. C habitus, ventral. Inset showing notched labium. D clypeus, subanterior. Scale bars: 1 mm (A), 0.5 mm (B-D)." figureDoi="10.3897/asp.80.e86008.figure4" httpUri="https://binary.pensoft.net/fig/749998" pageId="0" pageNumber="541">4D</figureCitation>
|
|
); cheliceral lamina present, with broad, triangular apex; at least one promarginal tooth present. Endites not converging in front of labium; anterior margin of labium notched (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. A, B habitus, dorsal. C habitus, ventral. Inset showing notched labium. D clypeus, subanterior. Scale bars: 1 mm (A), 0.5 mm (B-D)." figureDoi="10.3897/asp.80.e86008.figure4" httpUri="https://binary.pensoft.net/fig/749998" pageId="0" pageNumber="541">4C</figureCitation>
|
|
). Sternum slightly longer than wider. Legs long and thin (Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), photographs under light microscopy. A, B habitus, dorsolateral. C right femur I. Arrows to anterior macrosetae. D spinnerets, dorsolateral view. E, F, H right tarsi (E: III-IV, F: II, H: I), prolateral. Arrows to third claw. Note highly articulated podotarsite. G Chelicerae and palp, subdorsal view. Arrow to tip of right embolus. Scale bars: 1 mm (A), 0.5 mm (B), 0.2 mm (C), 0.1 mm (D, G), 0.05 mm (E, F, H)." figureDoi="10.3897/asp.80.e86008.figure3" httpUri="https://binary.pensoft.net/fig/749997" pageId="0" pageNumber="541">3A</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. A, B habitus, dorsal. C habitus, ventral. Inset showing notched labium. D clypeus, subanterior. Scale bars: 1 mm (A), 0.5 mm (B-D)." figureDoi="10.3897/asp.80.e86008.figure4" httpUri="https://binary.pensoft.net/fig/749998" pageId="0" pageNumber="541">4A</figureCitation>
|
|
) with a few macrosetae on the prolateral face of the first femur (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), photographs under light microscopy. A, B habitus, dorsolateral. C right femur I. Arrows to anterior macrosetae. D spinnerets, dorsolateral view. E, F, H right tarsi (E: III-IV, F: II, H: I), prolateral. Arrows to third claw. Note highly articulated podotarsite. G Chelicerae and palp, subdorsal view. Arrow to tip of right embolus. Scale bars: 1 mm (A), 0.5 mm (B), 0.2 mm (C), 0.1 mm (D, G), 0.05 mm (E, F, H)." figureDoi="10.3897/asp.80.e86008.figure3" httpUri="https://binary.pensoft.net/fig/749997" pageId="0" pageNumber="541">3C</figureCitation>
|
|
) and on the ventral faces of all tibiae and metatarsi. Well-developed podotarsite; third claw present (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), photographs under light microscopy. A, B habitus, dorsolateral. C right femur I. Arrows to anterior macrosetae. D spinnerets, dorsolateral view. E, F, H right tarsi (E: III-IV, F: II, H: I), prolateral. Arrows to third claw. Note highly articulated podotarsite. G Chelicerae and palp, subdorsal view. Arrow to tip of right embolus. Scale bars: 1 mm (A), 0.5 mm (B), 0.2 mm (C), 0.1 mm (D, G), 0.05 mm (E, F, H)." figureDoi="10.3897/asp.80.e86008.figure3" httpUri="https://binary.pensoft.net/fig/749997" pageId="0" pageNumber="541">3F</figureCitation>
|
|
). Spinnerets in a compact group (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. Spinnerets (A - C), left palp (D - H), and right palp (I, J). A ventral. B apical. C dorsal. D subprolateral. E retrolateral. F prolateral. G bulb and cymbium, apical. H bulb and cymbium, dorsal. I prolateral. J retrolateral. Asterisks are artifacts in the rendering. Scale bars: 0.1 mm (only A-C, F, I to scale)." figureDoi="10.3897/asp.80.e86008.figure5" httpUri="https://binary.pensoft.net/fig/749999" pageId="0" pageNumber="541">5B</figureCitation>
|
|
); colulus broad (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. Spinnerets (A - C), left palp (D - H), and right palp (I, J). A ventral. B apical. C dorsal. D subprolateral. E retrolateral. F prolateral. G bulb and cymbium, apical. H bulb and cymbium, dorsal. I prolateral. J retrolateral. Asterisks are artifacts in the rendering. Scale bars: 0.1 mm (only A-C, F, I to scale)." figureDoi="10.3897/asp.80.e86008.figure5" httpUri="https://binary.pensoft.net/fig/749999" pageId="0" pageNumber="541">5A</figureCitation>
|
|
); ALS short and broad, barely longer than PLS (Figs
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. A, B habitus, dorsal. C habitus, ventral. Inset showing notched labium. D clypeus, subanterior. Scale bars: 1 mm (A), 0.5 mm (B-D)." figureDoi="10.3897/asp.80.e86008.figure4" httpUri="https://binary.pensoft.net/fig/749998" pageId="0" pageNumber="541">4D</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. Spinnerets (A - C), left palp (D - H), and right palp (I, J). A ventral. B apical. C dorsal. D subprolateral. E retrolateral. F prolateral. G bulb and cymbium, apical. H bulb and cymbium, dorsal. I prolateral. J retrolateral. Asterisks are artifacts in the rendering. Scale bars: 0.1 mm (only A-C, F, I to scale)." figureDoi="10.3897/asp.80.e86008.figure5" httpUri="https://binary.pensoft.net/fig/749999" pageId="0" pageNumber="541">5C</figureCitation>
|
|
). Palps (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Drymusa aculicaput (Wunderlich, 2004) comb. nov., holotype (SMNG 07 / 36287 - 422), rendered volume after image stack obtained with synchrotron radiation micro-computed tomography. Spinnerets (A - C), left palp (D - H), and right palp (I, J). A ventral. B apical. C dorsal. D subprolateral. E retrolateral. F prolateral. G bulb and cymbium, apical. H bulb and cymbium, dorsal. I prolateral. J retrolateral. Asterisks are artifacts in the rendering. Scale bars: 0.1 mm (only A-C, F, I to scale)." figureDoi="10.3897/asp.80.e86008.figure5" httpUri="https://binary.pensoft.net/fig/749999" pageId="0" pageNumber="541">5D-J</figureCitation>
|
|
): tibia shorter than femur, incrassate; cymbium short with a small apical extension; bulb with globose, rounded base and thin, straight, needle-shaped embolus. -
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="541">Measurements</emphasis>
|
|
[mm]: Total length (excluding chelicerae and spinnerets) 2.34. Carapace length 1.15, width 0.89. Abdomen length 1.27, width 0.66. Palpal femur length 0.52. Palpal tibia length 0.24, height 0.21. Leg I: femur 2.87, patella 0.27, tibia 3.05, metatarsus 0.91, tarsus 2.25. Leg II: femur 2.74, patella 0.38, tibia 2.67, metatarsus 3.65, tarsus 0.66. Leg III: femur 2.13, patella 0.27, tibia 1.88, metatarsus 1.95, tarsus 0.72. Leg IV: femur 2.58, patella 0.33, tibia 2.67, metatarsus 2.55, tarsus 0.72.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</subSection>
|
|
</document> |