165 lines
16 KiB
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165 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.929.49987" ID-GBIF-Dataset="97c79ed9-33fd-49b8-90c5-95a159839db2" ID-PMC="PMC7192954" ID-Pensoft-Pub="1313-2970-929-1" ID-Pensoft-UUID="EFB92B457B1D56A1996483C5EECFAAF0" ID-PubMed="32377146" ID-ZooBank="4FCA3A27–973C49F7–8264–6A34D279EC5C" ModsDocID="1313-2970-929-1" checkinTime="1587627147117" checkinUser="pensoft" docAuthor="Gittenberger, Edmund, Leda, Pema, Wangchuk, Jigme, Gyeltshen, Choki & Bjoern Stelbrink," docDate="2020" docId="819F33E69E015A96978B52B692112C7D" docLanguage="en" docName="ZooKeys 929: 1-17" docOrigin="ZooKeys 929" docSource="http://dx.doi.org/10.3897/zookeys.929.49987" docTitle="Erhaia pelkiae Gittenberger & Gyeltshen 2020, sp. nov." docType="treatment" docUuid="8548A11B-5AAC-4D3E-9064-6321DF57677C" docUuidSource="ZooBank" docVersion="4" id="EFB92B457B1D56A1996483C5EECFAAF0" lastPageNumber="6" masterDocId="EFB92B457B1D56A1996483C5EECFAAF0" masterDocTitle="The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya" masterLastPageNumber="17" masterPageNumber="1" pageNumber="4" updateTime="1668168753491" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gittenberger, Edmund</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, P. O. Box 9517, NL- 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:nameIdentifier type="email">egittenberger@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Leda, Pema</mods:namePart>
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<mods:affiliation>National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wangchuk, Jigme</mods:namePart>
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<mods:affiliation>Ugyen Wangchuck Institute for Conservation and Environmental Research, Bumthang, Bhutan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-3503-4369</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gyeltshen, Choki</mods:namePart>
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<mods:affiliation>National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6790-6278</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bjoern Stelbrink,</mods:namePart>
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<mods:affiliation>Department of Animal Ecology & Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, D- 35392 Giessen, Germany & Zoological Institute, Department of Environmental Sciences, University of Basel, Vesalgasse 1, 4051 Basel, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-7471-4992</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>929</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>17</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.929.49987</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.929.49987</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-929-1</mods:identifier>
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<mods:identifier type="ZooBank">4FCA3A27-973C49F7-8264-6A34D279EC5C</mods:identifier>
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<treatment ID-GBIF-Taxon="163735478" LSID="urn:lsid:zoobank.org:act:8548A11B-5AAC-4D3E-9064-6321DF57677C" httpUri="http://treatment.plazi.org/id/819F33E69E015A96978B52B692112C7D" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
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<subSubSection pageId="3" pageNumber="4" type="nomenclature">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName LSID="819F33E6-9E01-5A96-978B-52B692112C7D" authority="Gittenberger & Gyeltshen" authorityName="Gittenberger & Gyeltshen" authorityYear="2020" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia pelkiae" order="Littorinimorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pelkiae" status="sp. nov.">Erhaia pelkiae Gittenberger & Gyeltshen</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="4">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">Figs 4</figureCitation>
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">, 5</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4">, 11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="materials_examined">
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<paragraph pageId="4" pageNumber="5">
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<pageBreakToken pageId="4" pageNumber="5" start="start">Material</pageBreakToken>
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examined.
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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<emphasis bold="true" pageId="4" pageNumber="5">
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<emphasis italics="true" pageId="4" pageNumber="5">Holotype</emphasis>
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.
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</emphasis>
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(Fig.
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="4" pageNumber="5">4</figureCitation>
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) Bhutan • District Thimphu, c. 5 km E of Chhuzom, W of Geneykha; in brooklet with a prayer wheel along the road; 2750 m a.s.l.;
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<geoCoordinate degrees="27" direction="north" minutes="18" orientation="latitude" precision="15" seconds="43" value="27.311943">27°18'43"N</geoCoordinate>
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,
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<geoCoordinate degrees="89" direction="east" minutes="36" orientation="longitude" precision="15" seconds="10" value="89.602776">89°36'10"E</geoCoordinate>
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; E. Gittenberger, Choki Gyeltshen & Pema Leda leg. 25.X.2018; NBCB 1059.
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<emphasis bold="true" pageId="4" pageNumber="5">
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<emphasis italics="true" pageId="4" pageNumber="5">Paratype</emphasis>
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.
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</emphasis>
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(Fig.
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<figureCitation captionStart="Figures 2–10" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub & Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann & S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis & Rao, 1997, holotype of E. chandeshwariensis Nesemann & S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah & Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="4" pageNumber="5">5</figureCitation>
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) 1 shell; same data as for holotype; NBCB 1060.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="diagnosis">
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<paragraph pageId="4" pageNumber="5">Diagnosis.</paragraph>
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<paragraph pageId="4" pageNumber="5">Shell with a partly reflected peristome, teleoconch with spiral lirae.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="shell">
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<paragraph pageId="4" pageNumber="5">Shell.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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Shell elongated ovoid, with
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<normalizedToken originalValue="3½">31/2</normalizedToken>
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convex, shouldered whorls that are separated by a deep suture; clearly higher than broad; light yellowish brown with fine growth lines and some brown periostracal ridges. Peristome reflected at the columellar and the basal side. Parietal and columellar side of the aperture thickened by a whitish callus. An irregular umbilical chink only in the paratype might be represent a malformation resulting from repair of the shell wall. Protoconch encrustated; teleoconch with fine spiral lines.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="description">
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<paragraph pageId="4" pageNumber="5">Measurements.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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(
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<emphasis italics="true" pageId="4" pageNumber="5">N</emphasis>
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= 2): holotype and paratype H 1.9 mm, B 1.1 mm.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="ecology">
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<paragraph pageId="5" pageNumber="6">
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<pageBreakToken pageId="5" pageNumber="6" start="start">Ecology</pageBreakToken>
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</paragraph>
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<paragraph pageId="5" pageNumber="6">
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(Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="5" pageNumber="6">11</figureCitation>
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). The snails are rare on the rocks in the shaded streamlet uphill the prayer wheel. See also the data for
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<taxonomicName lsidName="E. jannei" pageId="5" pageNumber="6" rank="species" species="jannei">
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<emphasis italics="true" pageId="5" pageNumber="6">E. jannei</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="notes">
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<paragraph pageId="5" pageNumber="6">Notes.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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The differences between the sympatric
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<taxonomicName lsidName="E. pelkiae" pageId="5" pageNumber="6" rank="species" species="pelkiae">
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<emphasis italics="true" pageId="5" pageNumber="6">E. pelkiae</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="E. jannei" pageId="5" pageNumber="6" rank="species" species="jannei">
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<emphasis italics="true" pageId="5" pageNumber="6">E. jannei</emphasis>
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</taxonomicName>
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are too large to regard as sexual dimorphism.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="6" type="etymology">
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<paragraph pageId="5" pageNumber="6">Etymology.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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The epithet
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<taxonomicName lsidName="pelkiae" pageId="5" pageNumber="6" rank="species" species="pelkiae">
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<emphasis italics="true" pageId="5" pageNumber="6">pelkiae</emphasis>
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</taxonomicName>
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refers to Ms. Pelki Yangdon, the daughter of the fourth author.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |