165 lines
14 KiB
XML
165 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62593" ID-GBIF-Dataset="2b58bee7-f25a-42c9-8651-3060ada2112e" ID-PMC="PMC8222207" ID-Pensoft-Pub="1313-2970-1044-563" ID-Pensoft-UUID="8787843B77CB51DEB6FCF0B8F3A09AC3" ID-PubMed="34183886" ID-ZooBank="71DCAFC2B2BB475B91662BCDC83C7F07" ModsDocID="1313-2970-1044-563" checkinTime="1623875370007" checkinUser="pensoft" docAuthor="Neri, Paolo & Toledano, Luca" docDate="2021" docId="389DE621EB165A8AAAABA928EACB2C33" docLanguage="en" docName="ZooKeys 1044: 563-587" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62593" docTitle="Bembidion (Peryphus) obscurellum Motschulsky 1845" docType="treatment" docVersion="4" id="8787843B77CB51DEB6FCF0B8F3A09AC3" lastPageNumber="563" masterDocId="8787843B77CB51DEB6FCF0B8F3A09AC3" masterDocTitle="Geographic and taxonomic notes, addenda and corrigenda on the subtribe Bembidiina Stephens, 1827 of the 2017 ' Catalogue of Palaearctic Coleoptera' (Coleoptera, Carabidae, Bembidiina)" masterLastPageNumber="587" masterPageNumber="563" pageNumber="563" updateTime="1668150485866" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Geographic and taxonomic notes, addenda and corrigenda on the subtribe Bembidiina Stephens, 1827 of the 2017 ' Catalogue of Palaearctic Coleoptera' (Coleoptera, Carabidae, Bembidiina)</mods:title>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neri, Paolo</mods:namePart>
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<mods:affiliation>Via Guido Rossa 21 " San Lorenzo ", 47121 Forli (FC), Italy</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Toledano, Luca</mods:namePart>
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<mods:affiliation>Museo Civico di Storia Naturale di Verona, Lungadige Porta Vittoria 9, 37129 Verona, Italy</mods:affiliation>
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<mods:nameIdentifier type="email">lucatole2@libero.it</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-06-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1044</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>563</mods:start>
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<mods:end>587</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1044.62593</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.62593</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1044-563</mods:identifier>
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<mods:identifier type="ZooBank">71DCAFC2B2BB475B91662BCDC83C7F07</mods:identifier>
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<mods:identifier type="Pensoft-UUID">8787843B77CB51DEB6FCF0B8F3A09AC3</mods:identifier>
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<treatment ID-GBIF-Taxon="182753950" LSID="urn:lsid:plazi:treatment:389DE621EB165A8AAAABA928EACB2C33" httpUri="http://treatment.plazi.org/id/389DE621EB165A8AAAABA928EACB2C33" lastPageNumber="563" pageId="0" pageNumber="563">
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<subSubSection pageId="0" pageNumber="563" type="nomenclature">
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<paragraph pageId="0" pageNumber="563">
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<taxonomicName LSID="389DE621-EB16-5A8A-AAAB-A928EACB2C33" authority="Motschulsky, 1845" authorityName="Motschulsky" authorityYear="1845" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) obscurellum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="obscurellum" subGenus="Peryphus">Bembidion (Peryphus) obscurellum Motschulsky, 1845</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="563" type="reference_group">
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<paragraph pageId="0" pageNumber="563">
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<taxonomicName authorityName="Csiki" authorityYear="1928" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) obscurellum subsp. turanicum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subSpecies" species="obscurellum" subGenus="Peryphus" subSpecies="turanicum">Bembidion (Peryphus) obscurellum turanicum</taxonomicName>
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Csiki, 1928
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</paragraph>
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<paragraph pageId="0" pageNumber="563">
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<taxonomicName authorityName="Fassati" authorityYear="1957" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) obscurellum subsp. fumipenne" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subSpecies" species="obscurellum" subGenus="Peryphus" subSpecies="fumipenne">Bembidion (Peryphus) obscurellum fumipenne</taxonomicName>
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Fassati, 1957
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</paragraph>
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<paragraph pageId="0" pageNumber="563">
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<taxonomicName authorityName="Fassati" authorityYear="1957" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) obscurellum subsp. thibeticum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subSpecies" species="obscurellum" subGenus="Peryphus" subSpecies="thibeticum">Bembidion (Peryphus) obscurellum thibeticum</taxonomicName>
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Fassati, 1957
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</paragraph>
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<paragraph pageId="0" pageNumber="563">
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<taxonomicName authorityName="Lutshnik" authorityYear="1938" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) obscurellum subsp. insperatum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subSpecies" species="obscurellum" subGenus="Peryphus" subSpecies="insperatum">Bembidion (Peryphus) obscurellum insperatum</taxonomicName>
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Lutshnik, 1938
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="563" type="notes">
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<paragraph pageId="0" pageNumber="563">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="563">
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<bibRefCitation author="Fassati, M" journalOrPublisher="Casopis Cesckoslovenske Spolecnosti Entomologicke" pageId="0" pageNumber="563" pagination="312 - 321" refId="B11" refString="Fassati, M, 1957. Eine neue Art und drei Rassen von Mittelasiatischen Bembidien aus der Untergattung Peryphus Steph. Casopis Cesckoslovenske Spolecnosti Entomologicke 54: 312 - 321" title="Eine neue Art und drei Rassen von Mittelasiatischen Bembidien aus der Untergattung Peryphus Steph." volume="54" year="1957">Fassati (1957)</bibRefCitation>
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describes
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<taxonomicName class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) fuscicrum subsp. thibeticum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subSpecies" species="fuscicrum" subGenus="Peryphus" subSpecies="thibeticum">Bembidion (Peryphus) fuscicrum thibeticum</taxonomicName>
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ssp. nov. and
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<taxonomicName class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) fuscicrum subsp. fumipenne" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subSpecies" species="fuscicrum" subGenus="Peryphus" subSpecies="fumipenne">Bembidion (Peryphus) fuscicrum fumipenne</taxonomicName>
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nat. nov.; they are distinguished from the nominotypical form only by the coloration of the elytra.
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="Opuscula Entomologica, Supplementum" pageId="0" pageNumber="563" pagination="201 - 408" refId="B23" refString="Lindroth, CH, 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica, Supplementum 24: 201 - 408" title="The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3." volume="24" year="1963">Lindroth (1963)</bibRefCitation>
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synonymizes the following taxa with
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<taxonomicName lsidName="B. obscurellum" pageId="0" pageNumber="563" rank="species" species="obscurellum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. obscurellum</emphasis>
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</taxonomicName>
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Motschulsky, 1845:
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<taxonomicName lsidName="B. fuscicrum" pageId="0" pageNumber="563" rank="species" species="fuscicrum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. fuscicrum</emphasis>
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</taxonomicName>
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Motschulsky, 1855,
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<taxonomicName lsidName="B. fuscicrum subsp. turanicum" pageId="0" pageNumber="563" rank="subSpecies" species="fuscicrum" subSpecies="turanicum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. fuscicrum turanicum</emphasis>
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</taxonomicName>
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Csiki, 1928,
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<taxonomicName lsidName="B. fumipenne" pageId="0" pageNumber="563" rank="species" species="fumipenne">
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<emphasis italics="true" pageId="0" pageNumber="563">B. fumipenne</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="B. thibeticum" pageId="0" pageNumber="563" rank="species" species="thibeticum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. thibeticum</emphasis>
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</taxonomicName>
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, the last two taxa because they are only color varieties. Kryzhanovskij et al. (1995) listed
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<taxonomicName lsidName="B. turanicum" pageId="0" pageNumber="563" rank="species" species="turanicum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. turanicum</emphasis>
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</taxonomicName>
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as a subspecies of
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<taxonomicName lsidName="B. obscurellum" pageId="0" pageNumber="563" rank="species" species="obscurellum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. obscurellum</emphasis>
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</taxonomicName>
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and retained
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<taxonomicName lsidName="B. fumipenne" pageId="0" pageNumber="563" rank="species" species="fumipenne">
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<emphasis italics="true" pageId="0" pageNumber="563">B. fumipenne</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="B. thibeticum" pageId="0" pageNumber="563" rank="species" species="thibeticum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. thibeticum</emphasis>
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</taxonomicName>
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as varieties only; moreover
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<normalizedToken originalValue="Belousov’s">Belousov's</normalizedToken>
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note (Kryzhanovskij et al. 1995: 85, note no. 158) reports that the subspecific situation of this polymorphic species is provisional. Recently, some authors (e.g.,
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Brill, Leiden-Boston" pageId="0" pageNumber="563" refId="B26" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (2nd edn.). Tutzing, 530 pp." title="Systematic List of Extant Ground Beetles of the World (2 nd edn.). Tutzing, 530 pp." year="2005">Lorenz 2005</bibRefCitation>
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;
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<bibRefCitation author="Marggi, W" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="563" refId="B29" refString="Marggi, W, Toledano, L, Neri, P, 2017. Carabidae: Bembidiini: Bembidiina: 294-342. In: Loebl I, Loebl D (Eds) Catalogue of Palaearctic Coleoptera (Vol. I). Archostemata - Myxophaga - Adephaga. Brill, Leiden" title="Carabidae: Bembidiini: Bembidiina: 294 - 342. In: Loebl I, Loebl D (Eds) Catalogue of Palaearctic Coleoptera (Vol. I). Archostemata - Myxophaga - Adephaga." year="2017">Marggi et al. 2017</bibRefCitation>
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) still list
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<taxonomicName lsidName="B. turanicum" pageId="0" pageNumber="563" rank="species" species="turanicum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. turanicum</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="B. fumipenne" pageId="0" pageNumber="563" rank="species" species="fumipenne">
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<emphasis italics="true" pageId="0" pageNumber="563">B. fumipenne</emphasis>
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</taxonomicName>
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, and
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<taxonomicName lsidName="B. thibeticum" pageId="0" pageNumber="563" rank="species" species="thibeticum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. thibeticum</emphasis>
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</taxonomicName>
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as subspecies of
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<taxonomicName lsidName="B. obscurellum" pageId="0" pageNumber="563" rank="species" species="obscurellum">
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<emphasis italics="true" pageId="0" pageNumber="563">B. obscurellum</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="563">
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In light of our studies we maintain that the synonymies proposed by
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="Opuscula Entomologica, Supplementum" pageId="0" pageNumber="563" pagination="201 - 408" refId="B23" refString="Lindroth, CH, 1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica, Supplementum 24: 201 - 408" title="The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3." volume="24" year="1963">Lindroth (1963)</bibRefCitation>
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were correct; moreover, our colleague Liang Hongbin (IZCAS, Beijing) confirms our opinion based on the study of hundreds of specimens from Mongolia that clearly show the variability of the species (Liang Hongbin, pers. comm.).
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</paragraph>
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<paragraph pageId="0" pageNumber="563">
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Regarding
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<taxonomicName authorityName="Lutshnik" authorityYear="1938" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) obscurellum subsp. insperatum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subSpecies" species="obscurellum" subGenus="Peryphus" subSpecies="insperatum">Bembidion (Peryphus) obscurellum insperatum</taxonomicName>
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Lutshnik, 1938, since its discriminating characters are not exclusive but shared by other subspecies, we evaluated the synonymy expressed in
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<bibRefCitation author="Marggi, W" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="563" refId="B29" refString="Marggi, W, Toledano, L, Neri, P, 2017. Carabidae: Bembidiini: Bembidiina: 294-342. In: Loebl I, Loebl D (Eds) Catalogue of Palaearctic Coleoptera (Vol. I). Archostemata - Myxophaga - Adephaga. Brill, Leiden" title="Carabidae: Bembidiini: Bembidiina: 294 - 342. In: Loebl I, Loebl D (Eds) Catalogue of Palaearctic Coleoptera (Vol. I). Archostemata - Myxophaga - Adephaga." year="2017">Marggi et al. (2017)</bibRefCitation>
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and add to the distribution of the species the Russian Caucasus (ST), the region mentioned in the description, as already reported in Kryzhanovskij et al. (1995): Ciscaucasia, Western and Central Greater Caucasus. Current distribution:
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<emphasis bold="true" pageId="0" pageNumber="563">E</emphasis>
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: DE, NT,
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<emphasis bold="true" pageId="0" pageNumber="563">ST</emphasis>
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, SV;
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<emphasis bold="true" pageId="0" pageNumber="563">A</emphasis>
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: AF, CH, ES, IN, KA, KI, KZ, MG, TM, TR, WS, XIZ;
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<emphasis bold="true" pageId="0" pageNumber="563">NAR</emphasis>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |