165 lines
11 KiB
XML
165 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.773.24878" ID-GBIF-Dataset="3b67ab3e-bd75-4194-a90c-c12b3dc8332b" ID-Pensoft-Pub="1313-2970-773-1" ID-ZBK="900F9307844A4B9AB428DA97FF3B4B5C" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-773-1" ModsDocOrigin="ZooKeys 773" ModsDocTitle="A molecular and conchological dissection of the “scaly” Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)" checkinTime="1531209184653" checkinUser="pensoft" docAuthor="Khalik, Mohd Zacaery, Hendriks, Kasper, Vermeulen, Jaap J. & Schilthuizen, Menno" docDate="2018" docId="EAF0B755CA06762D7FCE3A73F96B1A67" docLanguage="en" docName="ZooKeys 773: 1-55" docOrigin="ZooKeys 773" docSource="http://dx.doi.org/10.3897/zookeys.773.24878" docTitle="Georissa bauensis Khalik, Hendriks, Vermeulen & Schilthuizen, 2018, sp. n." docType="treatment" docUuid="6E333906-75DD-4055-BD32-578E0D651E1F" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FFACFFA96149FF8D3477FFCCD227FF8E" masterDocTitle="A molecular and conchological dissection of the " scaly " Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)" masterLastPageNumber="55" masterPageNumber="1" pageNumber="1" updateTime="1643514872515" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A molecular and conchological dissection of the " scaly " Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Khalik, Mohd Zacaery</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hendriks, Kasper</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>773</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>55</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.773.24878</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.773.24878</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-773-1</mods:identifier>
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<mods:identifier type="ZBK">900F9307844A4B9AB428DA97FF3B4B5C</mods:identifier>
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<mods:identifier type="ZooBank">900F9307844A4B9AB428DA97FF3B4B5C</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="146053681" LSID="urn:lsid:zoobank.org:act:6E333906-75DD-4055-BD32-578E0D651E1F" httpUri="http://treatment.plazi.org/id/EAF0B755CA06762D7FCE3A73F96B1A67" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/6E333906-75DD-4055-BD32-578E0D651E1F" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa bauensis" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bauensis">Georissa bauensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Wind Cave Passage 3, Wind Cave Nature Reserve, Bau, Sarawak, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="1.4135">01°24.81'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="110.13617">110°08.17'E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Holotype. Wind Cave Passage 3, Wind Cave Nature Reserve, Bau, Sarawak, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="1.4135">01°24.81'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="110.13617">110°08.17'E</geoCoordinate>
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): MZU/MOL 17.89 (leg. MZ Khalik). Paratypes. Wind Cave Passage 3, Wind Cave Nature Reserve, Bau, Sarawak, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="1.4135">01°24.81'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="110.13617">110°08.17'E</geoCoordinate>
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): MZU/MOL 16.01 (25), MZU/MOL 16.02 (>50). Gunung Podam, near Sungai Ayup, Kampung Bogag, Bau, Sarawak, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="1.3525">01°21.15'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="110.0595">110°03.57'E</geoCoordinate>
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): MZU/MOL 16.03 (5).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">Named after the district of Bau, Sarawak, Malaysia, where the type locality Wind Cave Nature Reserve is located.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Protoconch. Color: red. Sculpture pattern: meshed - rounded or irregular mesh shape. Mesh width: 12-22
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<normalizedToken originalValue="µm">µm</normalizedToken>
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. Teleoconch. Color: orange to red. First whorl: shouldered, flat both above and below the shoulder. Subsequent whorls: convex shoulder and more rounded at the periphery. SH: 1.16-1.62 mm, SW: 1.06-1.30 mm, SI: 1.02-1.25. Total number of whorls: 2-2
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<normalizedToken originalValue="½">1/2</normalizedToken>
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. Shell sculpture. Radial sculpture: absent, only weak growth lines are visible. Spiral sculpture: present, weakly to strongly sculpted, continuous to discontinuous ribs, more prominent at the periphery. Scales: two to three major spiral series of low and small diagonal scales, regularly spaced, the upper series always stronger than the lower ones, scale series irregularly spaced. Aperture. Shape: rounded and tilted below. Basal side: rounded, angular at the columellar region. Parietal side: straight. AH: 0.57-0.78 mm, AW: 0.69-0.86 mm, AI: 0.74-0.96. Holotype dimension. SH: 1.16 mm, SW: 1.06 mm, AH: 0.58 mm, AW: 0.70 mm.
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</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 14.
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa bauensis" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bauensis">Georissa bauensis</taxonomicName>
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sp. n.
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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Holotype: MZU/MOL 17.89
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<normalizedToken originalValue="D–K">D-K</normalizedToken>
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Paratypes: MZU/MOL 16.03. A, D Shell apertural view B Shell side view C Shell rear view
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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Shell cross-section from 3D model
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<normalizedToken originalValue="G–H">G-H</normalizedToken>
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Operculum frontal and ventral view I Shell top view J Protoconch side view K Close up of protoconch from top at 1000
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification. Scale bars: 500
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<normalizedToken originalValue="A–I">A-I</normalizedToken>
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); 200
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(J); 10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(K).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="cross diagnosis">
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<paragraph pageId="0" pageNumber="1">Cross diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa bauensis" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="bauensis">Georissa bauensis</taxonomicName>
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is very similar to
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<taxonomicName genus="G." lsidName="G. kobelti" pageId="0" pageNumber="1" rank="species" species="kobelti">G. kobelti</taxonomicName>
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(although not closely related phylogenetically), in terms of general shell shape and spiral scale characters. However,
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<taxonomicName genus="G." lsidName="G. bauensis" pageId="0" pageNumber="1" rank="species" species="bauensis">G. bauensis</taxonomicName>
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is sufficiently variable to include specimens that are more similar to
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<taxonomicName genus="G." lsidName="G. hosei" pageId="0" pageNumber="1" rank="species" species="hosei">G. hosei</taxonomicName>
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and
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<taxonomicName genus="G." lsidName="G. scalinella" pageId="0" pageNumber="1" rank="species" species="scalinella">G. scalinella</taxonomicName>
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. Furthermore,
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<taxonomicName genus="G." lsidName="G. bauensis" pageId="0" pageNumber="1" rank="species" species="bauensis">G. bauensis</taxonomicName>
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has more strongly sculpted scales than
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<taxonomicName genus="G." lsidName="G. hosei" pageId="0" pageNumber="1" rank="species" species="hosei">G. hosei</taxonomicName>
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, and a more rounded and convex shell than
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<taxonomicName genus="G." lsidName="G. scalinella" pageId="0" pageNumber="1" rank="species" species="scalinella">G. scalinella</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Known from Gunung Podam and Wind Cave Nature Reserve, Bau, Sarawak.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="molecular analysis">
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<paragraph pageId="0" pageNumber="1">Molecular analysis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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ML and Bayesian analyses resolve all individuals of
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<taxonomicName genus="G." lsidName="G. bauensis" pageId="0" pageNumber="1" rank="species" species="bauensis">G. bauensis</taxonomicName>
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(16S: n = 13; CO1: n = 8) as a monophyletic group with 99% BS and 100% PP, the sister group of
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<taxonomicName genus="G." lsidName="G. silaburensis" pageId="0" pageNumber="1" rank="species" species="silaburensis">G. silaburensis</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |