397 lines
46 KiB
XML
397 lines
46 KiB
XML
<document id="222818341075889C35BD12349498B670" ID="10.11646/zootaxa.4138.2.3" ID-DOI="10.11646/zootaxa.4138.2.3" ID-GBIF-Dataset="ff393c27-4a85-4e2e-99b0-87286a6f24fb" ID-ISSN="1175-5326" ID-Zenodo-Dep="255350" ID-ZooBank="4F1E4A87-5370-420C-9C16-2BC373645110" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468565702401" checkinUser="plazi" docAuthor="Doan, Tiffany M., Mason, Andrew J., Castoe, Todd A., Sasa, Mahmood & Parkinson, Christopher L." docDate="2016" docId="E07487CFFFDB8D3AE9D3D8E3CA380619" docLanguage="en" docName="zootaxa.4138.2.3.pdf" docOrigin="Zootaxa 4138 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bothriechis nubestris Doan, Mason, Castoe, Sasa & Parkinson, 2016, new species" docType="treatment" docUuid="9A385383-FA5C-46C3-B3D0-F83F2D577DA5" docUuidSource="ZooBank" docVersion="8" lastPageNumber="281" masterDocId="1C4DFFB7FFDD8D30E944DA4DCB200704" masterDocTitle="A cryptic palm-pitviper species (Squamata: Viperidae: Bothriechis) from the Costa Rican highlands, with notes on the variation within B. nigroviridis" masterLastPageNumber="290" masterPageNumber="271" pageNumber="277" updateTime="1698365156826" updateUser="plazi">
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<mods:mods id="5A731DF59FFDF45A7A7AECC7B0147ECB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A263933456E2311B83C73DDB47E15F6B">
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<mods:title id="E65F30443426A9E14019ABB5221C6661">A cryptic palm-pitviper species (Squamata: Viperidae: Bothriechis) from the Costa Rican highlands, with notes on the variation within B. nigroviridis</mods:title>
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</mods:titleInfo>
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<mods:name id="45399D4109CF4DE8E422767885879AA4" type="personal">
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<mods:role id="DA3B7EA746AE125281CC646D5BF39792">
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<mods:roleTerm id="190A4C4CC68AB2BD19D5F9732C6F19F4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="254CAE3BE43F0696F7BBBA1217612E75">Doan, Tiffany M.</mods:namePart>
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</mods:name>
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<mods:name id="155D57233088771CC0B1FB1C59EE0BAA" type="personal">
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<mods:role id="FC7DD83FFF94E05E1FD746D161680405">
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<mods:roleTerm id="EE3EB6FC76A46DB662FDB0FB229C6663">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EA2620BBE235D07BAC8D8B6F3C37960B">Mason, Andrew J.</mods:namePart>
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</mods:name>
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<mods:name id="C3B7F576E49585FA987DCCE566A21D47" type="personal">
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<mods:role id="A86D86287BDCBFE15359241FD708FFF2">
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<mods:roleTerm id="DA555030F592E8C22BC821CC866D1DAA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="933E3326A59DAC4498211B99C0F60575">Castoe, Todd A.</mods:namePart>
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</mods:name>
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<mods:name id="C9CB1AD19D2AE12A82ED6B60FCE13710" type="personal">
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<mods:role id="BECCDB2AADB4B15E726C7577BBC47038">
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<mods:roleTerm id="CEA208A6BB58E285C879716D24004E9A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FFB1EB37C1142FE0E96C48D8139F4145">Sasa, Mahmood</mods:namePart>
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</mods:name>
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<mods:name id="2BFE46B5F0BD3C7C91C8FFF8A8F87ECC" type="personal">
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<mods:role id="719C979C079DC2F603DC0442A0CA7767">
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<mods:roleTerm id="CA788F5F48204E4BFFAA6D2D74B804C6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="49990FB0880EF582A8ACB3CBD84A6B71">Parkinson, Christopher L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="810BA0806A124A28965DA6040DEB1BD4">text</mods:typeOfResource>
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<mods:titleInfo id="B519BD134FCBD9C949DFFAB1B9682868">
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<mods:title id="254ED00F6198579D9037F02F5E933D73">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3E7F2360AA4435107B9023D9DE22D377">
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<mods:date id="5C56B82D75DCB92317BBF42CB23FD403">2016</mods:date>
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<mods:detail id="BCBBA0A372747BDFE62739857179F73B" type="volume">
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<mods:number id="659E2D3C482955EA5A5C86B813FDFB6C">4138</mods:number>
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</mods:detail>
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<mods:detail id="CA32AB4457D16405B4FE94E071137E5A" type="issue">
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<mods:number id="2A26F68B20529EA9257A09672B907ABA">2</mods:number>
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<mods:end id="D0C462CE5AA40604431184A475ECD6DC">290</mods:end>
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<mods:classification id="B2ABC44018AAE230E1F289F2BDC7D331">journal article</mods:classification>
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<mods:identifier id="1B55854A5499E4C0406A0B49C85B3D5D" type="DOI">10.11646/zootaxa.4138.2.3</mods:identifier>
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<mods:identifier id="13E293FA40FDDD5D85956FC1F157C2D2" type="GBIF-Dataset">ff393c27-4a85-4e2e-99b0-87286a6f24fb</mods:identifier>
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<mods:identifier id="61131247E51CE77ECC1EFAE643392077" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3126955F6F422A94C2C1B16CA11374F0" type="Zenodo-Dep">255350</mods:identifier>
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<mods:identifier id="CE28F0ED245889FBBC95DB00E57279B7" type="ZooBank">4F1E4A87-5370-420C-9C16-2BC373645110</mods:identifier>
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</mods:mods>
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<treatment id="E07487CFFFDB8D3AE9D3D8E3CA380619" ID-DOI="http://doi.org/10.5281/zenodo.5658645" ID-GBIF-Taxon="123195675" ID-Zenodo-Dep="5658645" LSID="urn:lsid:zoobank.org:act:9A385383-FA5C-46C3-B3D0-F83F2D577DA5" httpUri="http://treatment.plazi.org/id/E07487CFFFDB8D3AE9D3D8E3CA380619" lastPageId="10" lastPageNumber="281" pageId="6" pageNumber="277">
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<subSubSection id="20C76552FFDB8D36E9D3D8E3CA03040A" pageId="6" pageNumber="277" type="nomenclature">
|
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<paragraph id="686236D9FFDB8D36E9D3D8E3C91005CC" blockId="6.[151,560,686,782]" box="[151,560,686,712]" pageId="6" pageNumber="277">
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<heading id="332A81B5FFDB8D36E9D3D8E3C91005CC" bold="true" box="[151,560,686,712]" fontSize="11" level="1" pageId="6" pageNumber="277" reason="1">
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||
<emphasis id="5AA9EACBFFDB8D36E9D3D8E3C91005CC" bold="true" box="[151,560,686,712]" pageId="6" pageNumber="277">
|
||
<taxonomicName id="AFDD4D5AFFDB8D36E9D3D8E3CAB905CC" box="[151,409,686,712]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nubestris" status="sp. nov.">
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||
<emphasis id="5AA9EACBFFDB8D36E9D3D8E3CAB905CC" bold="true" box="[151,409,686,712]" italics="true" pageId="6" pageNumber="277">Bothriechis nubestris</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="419A57B0FFDB8D36E8E4D8E3C91005CC" box="[416,560,686,712]" pageId="6" pageNumber="277" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
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||
<paragraph id="686236D9FFDB8D36E9D3D89CCA03040A" blockId="6.[151,560,686,782]" pageId="6" pageNumber="277">
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||
Talamancan Palm-Pitviper
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<figureCitation id="F0E62A5CFFDB8D36E9D3D8B8CA3F040A" box="[151,287,757,782]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="8.[151,250,1953,1975]" captionStartId-1="9.[151,250,1658,1680]" captionTargetBox-0="[369,1215,193,1931]" captionTargetBox-1="[168,1432,225,1636]" captionTargetId-0="figure@8.[369,1217,193,1932]" captionTargetId-1="figure@9.[151,1436,213,1645]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 4. Photographs of Bothriechis nubestris sp. n. in life. Top photographer: John Tashjian; middle and bottom photographer Wayne van Devender." captionText-1="FIGURE 5. Dorsal and lateral views of the head of male paratype UCR 11151 from San Gerardo, Costa Rica. Scale bar = 20 mm. The diagnostic kidney-shaped supraoculars and partial scale rows between the supralabials and suboculars are highlighted." httpUri-0="https://zenodo.org/record/255354/files/figure.png" httpUri-1="https://zenodo.org/record/255355/files/figure.png" pageId="6" pageNumber="277">Figures 4–5</figureCitation>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="20C76552FFDB8D36E9D3D970C9B90457" box="[151,665,829,851]" pageId="6" pageNumber="277" type="reference_group">
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||
<paragraph id="686236D9FFDB8D36E9D3D970C9B90457" blockId="6.[151,665,829,851]" box="[151,665,829,851]" pageId="6" pageNumber="277">
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||
<taxonomicName id="AFDD4D5AFFDB8D36E9D3D970C9150457" authority="Peters 1859:278" authorityName="Peters" authorityPageNumber="278" authorityYear="1859" box="[151,565,829,851]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nigroviridis">
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||
<emphasis id="5AA9EACBFFDB8D36E9D3D970CAA90457" box="[151,393,829,851]" italics="true" pageId="6" pageNumber="277">Bothriechis nigroviridis</emphasis>
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||
Peters 1859:278
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</taxonomicName>
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||
. [In part.]
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="20C76552FFDB8D36E9D3D932CAB00221" pageId="6" pageNumber="277" type="materials_examined">
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||
<paragraph id="686236D9FFDB8D36E9D3D932CE6804B9" blockId="6.[151,1437,895,2037]" pageId="6" pageNumber="277">
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||
<emphasis id="5AA9EACBFFDB8D36E9D3D932CA2B049C" bold="true" box="[151,267,895,920]" pageId="6" pageNumber="277">
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||
<typeStatus id="B766887BFFDB8D36E9D3D932CA26049C" box="[151,262,895,920]" pageId="6" pageNumber="277" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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||
<collectionCode id="0ECCAE1CFFDB8D36E850D9CDCA6B049C" box="[276,331,896,920]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="6" pageNumber="277">UTA</collectionCode>
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||
<collectionCode id="0ECCAE1CFFDB8D36E810D932CA50049C" box="[340,368,895,920]" pageId="6" pageNumber="277">R-</collectionCode>
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||
9637 (
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||
<bibRefCitation id="0C4C4B28FFDB8D36E8FCD932C843049C" author="Campbell" box="[440,867,895,920]" pageId="6" pageNumber="277" refString="Campbell, J. A. & Lamar, W. W. (2004) Venomous Reptiles of the Western Hemisphere, Vol 2. Cornell University Press. Ithaca, USA, 976 pp." type="book" year="2004">Campbell and Lamar, 2004: fig. 97B</bibRefCitation>
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||
), an adult female, from San Isidro de El General, Province of San José,
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||
<collectingCountry id="10CA7649FFDB8D36E8CBD9E9C91504B8" box="[399,565,932,956]" name="Costa Rica" pageId="6" pageNumber="277">COSTA RICA</collectingCountry>
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||
; approximately
|
||
<quantity id="AF259B3CFFDB8D36EBAED9E9C86204B8" box="[746,834,932,956]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="277" unit="m" value="3000.0">3000 m</quantity>
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||
; collected in
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<date id="1C631019FFDB8D36EA92D9E9CF5004B8" box="[982,1136,932,957]" pageId="6" pageNumber="277" value="1973-10">October 1973</date>
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||
by Peter Seigfried.
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||
</paragraph>
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||
<paragraph id="686236D9FFDB8D36E983D98AC8E7032C" blockId="6.[151,1437,895,2037]" pageId="6" pageNumber="277">
|
||
<emphasis id="5AA9EACBFFDB8D36E983D98ACA6604E4" bold="true" box="[199,326,967,992]" pageId="6" pageNumber="277">
|
||
<typeStatus id="B766887BFFDB8D36E983D98ACA6104E4" box="[199,321,967,992]" pageId="6" pageNumber="277" type="paratype">Paratypes</typeStatus>
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||
.
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||
</emphasis>
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||
All from province of San José,
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||
<collectingCountry id="10CA7649FFDB8D36EBE8D985C87404E4" box="[684,852,968,992]" name="Costa Rica" pageId="6" pageNumber="277">COSTA RICA</collectingCountry>
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:
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<collectionCode id="0ECCAE1CFFDB8D36EA1AD985C8B504E4" box="[862,917,968,992]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="6" pageNumber="277">UTA</collectionCode>
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||
<collectionCode id="0ECCAE1CFFDB8D36EAD9D985C89A04E4" box="[925,954,968,992]" pageId="6" pageNumber="277">R-</collectionCode>
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||
2801, 2808, 2850, 7327, 7463, 9635, 9636, 10433, all from San Isidro de El General District;
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<collectionCode id="0ECCAE1CFFDB8D36EB8ED9A1C8240300" box="[714,772,1004,1028]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="6" pageNumber="277">UCR</collectionCode>
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||
5727, 5728, 5758, 11151, 15420, 15423, 15424, 15432, all from San Gerardo de Dota;
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<collectionCode id="0ECCAE1CFFDB8D36E88ADE5DC928032C" box="[462,520,1040,1064]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="6" pageNumber="277">UCR</collectionCode>
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12356, from División, Páramo District.
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||
</paragraph>
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||
<paragraph id="686236D9FFDB8D36E983DE79CAB00221" blockId="6.[151,1437,895,2037]" pageId="6" pageNumber="277">
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||
<emphasis id="5AA9EACBFFDB8D36E983DE79CA980349" bold="true" box="[199,440,1076,1101]" pageId="6" pageNumber="277">Referred specimens.</emphasis>
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From province of San José,
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<collectingCountry id="10CA7649FFDB8D36EBB3DE79C8BF0348" box="[759,927,1076,1100]" name="Costa Rica" pageId="6" pageNumber="277">COSTA RICA</collectingCountry>
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||
:
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||
<collectionCode id="0ECCAE1CFFDB8D36EAEDDE79C8C30348" box="[937,995,1076,1100]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="6" pageNumber="277">UCR</collectionCode>
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5757, from San Gerardo de Dota;
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<collectionCode id="0ECCAE1CFFDB8D36EC21DE79CEBC0348" box="[1381,1436,1076,1100]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="6" pageNumber="277">UTA</collectionCode>
|
||
<collectionCode id="0ECCAE1CFFDB8D36E9D3DE1ACB930374" box="[151,179,1111,1136]" pageId="6" pageNumber="277">R-</collectionCode>
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||
6799, 9364, 9365, 10432, all from San Isidro de El General District;
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||
<accessionNumber id="778EAB3AFFDB8D36EAA0DE15CF510374" box="[996,1137,1111,1136]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/CM145874" pageId="6" pageNumber="277" type="EnaNcbi">CM 145874</accessionNumber>
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||
, 148063, 148064,
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<collectionCode id="0ECCAE1CFFDB8D36EC0FDE15CEBC0374" box="[1355,1436,1112,1136]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="6" pageNumber="277">LACM</collectionCode>
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||
154553, Cerro de la Muerte near San Gerardo;
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||
<collectionCode id="0ECCAE1CFFDB8D36EBEFDE31C9C50390" box="[683,741,1148,1172]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="6" pageNumber="277">UCR</collectionCode>
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||
16726, Providencia;
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||
<collectionCode id="0ECCAE1CFFDB8D36EA92DE31CF300390" box="[982,1040,1148,1172]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="6" pageNumber="277">UCR</collectionCode>
|
||
15422, 15429, Alto Palma, Parque Nacional Braulio Carrillo, Bajo la Hondura (locality possibly in error);
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||
<collectionCode id="0ECCAE1CFFDB8D36EA86DEEDC8DC03BC" box="[962,1020,1184,1208]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="6" pageNumber="277">UCR</collectionCode>
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||
15428, Las Nubes (locality possibly in error);
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<collectionCode id="0ECCAE1CFFDB8D36E841DE89CA1E03D8" box="[261,318,1220,1244]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="6" pageNumber="277">UCR</collectionCode>
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||
3951, Naranjo, Río Naranjo;
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||
<collectionCode id="0ECCAE1CFFDB8D36EBD0DE88C9CB03D8" box="[660,747,1221,1244]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="6" pageNumber="277">UMMZ</collectionCode>
|
||
117734, no precise locality data. From province of Cartago:
|
||
<collectionCode id="0ECCAE1CFFDB8D36E9D3DEA5CBF503FB" box="[151,213,1256,1279]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="6" pageNumber="277">MVZ</collectionCode>
|
||
24228. From unknown Costa Rican locality:
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||
<collectionCode id="0ECCAE1CFFDB8D36EB96DEA5C82C0204" box="[722,780,1256,1280]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="6" pageNumber="277">UCR</collectionCode>
|
||
15439. From province of Limón:
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||
<collectionCode id="0ECCAE1CFFDB8D36EDCDDEA5CFE30204" box="[1161,1219,1256,1280]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="6" pageNumber="277">UCR</collectionCode>
|
||
22428, Talamanca, Telire, Cerro Uthyum.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="20C76552FFDB8D36E983DF62CF4501D0" pageId="6" pageNumber="277" type="diagnosis">
|
||
<paragraph id="686236D9FFDB8D36E983DF62CF4902DC" blockId="6.[151,1437,895,2037]" pageId="6" pageNumber="277">
|
||
<emphasis id="5AA9EACBFFDB8D36E983DF62CA61024C" bold="true" box="[199,321,1327,1352]" pageId="6" pageNumber="277">Diagnosis.</emphasis>
|
||
(1) a medium-sized slender arboreal pitviper; (2) dorsum green with heavy black mottling; (3) iris blackish; (4) superciliary scales absent; (5) interrictals 22–29; (6) supraoculars thin, usually kidney-shaped; (7) intersupraoculars 6–10; (8) partial rows or two rows of irregular scales between suboculars and supralabials; (9) infralabials 9–12; (10) first dorsals usually 21 (75.9%); (11) second dorsals usually 21 (58.6%); (12) third dorsals usually 17 (85.7%); (13) ventrals 150–160; (14) subcaudals 52–64; (15) tail prehensile.
|
||
</paragraph>
|
||
<paragraph id="686236D9FFDB8D36E983DFA9CF4501D0" blockId="6.[151,1437,895,2037]" pageId="6" pageNumber="277">
|
||
Specimens of
|
||
<taxonomicName id="AFDD4D5AFFDB8D36E821DFA9C97402F8" box="[357,596,1508,1532]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nubestris">
|
||
<emphasis id="5AA9EACBFFDB8D36E821DFA9C97402F8" box="[357,596,1508,1532]" italics="true" pageId="6" pageNumber="277">Bothriechis nubestris</emphasis>
|
||
</taxonomicName>
|
||
differ from
|
||
<taxonomicName id="AFDD4D5AFFDB8D36EB98DFA8C84702F8" box="[732,871,1508,1532]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="schlegelii">
|
||
<emphasis id="5AA9EACBFFDB8D36EB98DFA8C84702F8" box="[732,871,1508,1532]" italics="true" pageId="6" pageNumber="277">B. schlegelii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AFDD4D5AFFDB8D36EAD9DFA8CF6B02F8" box="[925,1099,1508,1532]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="supraciliaris">
|
||
<emphasis id="5AA9EACBFFDB8D36EAD9DFA8CF6B02F8" box="[925,1099,1508,1532]" italics="true" pageId="6" pageNumber="277">B. supraciliaris</emphasis>
|
||
</taxonomicName>
|
||
by lacking superciliary scales (present in
|
||
<taxonomicName id="AFDD4D5AFFDB8D36E85ADC44CA8D0124" box="[286,429,1544,1568]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="schlegelii">
|
||
<emphasis id="5AA9EACBFFDB8D36E85ADC44CA8D0124" box="[286,429,1544,1568]" italics="true" pageId="6" pageNumber="277">B. schlegelii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AFDD4D5AFFDB8D36E8A9DC44C9BF0124" box="[493,671,1544,1568]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="supraciliaris">
|
||
<emphasis id="5AA9EACBFFDB8D36E8A9DC44C9BF0124" box="[493,671,1544,1568]" italics="true" pageId="6" pageNumber="277">B. supraciliaris</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="AFDD4D5AFFDB8D36EBF8DC45C8900124" box="[700,944,1544,1568]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nubestris">
|
||
<emphasis id="5AA9EACBFFDB8D36EBF8DC45C8900124" box="[700,944,1544,1568]" italics="true" pageId="6" pageNumber="277">Bothriechis nubestris</emphasis>
|
||
</taxonomicName>
|
||
differs from all other
|
||
<taxonomicName id="AFDD4D5AFFDB8D36EDFADC45CE1F0124" box="[1214,1343,1544,1568]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="genus">
|
||
<emphasis id="5AA9EACBFFDB8D36EDFADC45CE1F0124" box="[1214,1343,1544,1568]" italics="true" pageId="6" pageNumber="277">Bothriechis</emphasis>
|
||
</taxonomicName>
|
||
species except
|
||
<taxonomicName id="AFDD4D5AFFDB8D36E9A3DC60CAA90140" box="[231,393,1580,1604]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nigroviridis">
|
||
<emphasis id="5AA9EACBFFDB8D36E9A3DC60CAA90140" box="[231,393,1580,1604]" italics="true" pageId="6" pageNumber="277">B. nigroviridis</emphasis>
|
||
</taxonomicName>
|
||
by having green dorsal coloration with heavy black mottling and a blackish iris.
|
||
<taxonomicName id="AFDD4D5AFFDB8D36EC5FDC61CBDE016C" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nubestris">
|
||
<emphasis id="5AA9EACBFFDB8D36EC5FDC61CBDE016C" italics="true" pageId="6" pageNumber="277">Bothriechis nubestris</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="AFDD4D5AFFDB8D36E8DEDC1CC91F016C" box="[410,575,1616,1640]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nigroviridis">
|
||
<emphasis id="5AA9EACBFFDB8D36E8DEDC1CC91F016C" box="[410,575,1616,1640]" italics="true" pageId="6" pageNumber="277">B. nigroviridis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<tableCitation id="255F0362FFDB8D36EB3BDC02C9F5016C" box="[639,725,1615,1640]" captionStart="TABLE 1" captionStartId="7.[151,239,508,531]" captionTargetBox="[159,1254,590,943]" captionTargetPageId="7" captionText="TABLE 1. Comparison of morphological characters of Bothriechis nigroviridis (n = 41) and B. nubestris (n = 27) (some specimens could not have all characters scored because of damage). The format is Range; Mode; Mean." httpUri="http://table.plazi.org/id/3CA26651FFDA8D37E9D3DBB1CFCA0537" pageId="6" pageNumber="277" tableUuid="3CA26651FFDA8D37E9D3DBB1CFCA0537">Table 1</tableCitation>
|
||
for summary) by the combination of having 150–160 ventral scales (
|
||
<quantity id="AF259B3CFFDB8D36E9AEDC39CA4B0189" box="[234,363,1652,1677]" metricMagnitude="0" metricUnit="m" metricValue="3.6195" metricValueMax="3.7845999999999997" metricValueMin="3.4543999999999997" pageId="6" pageNumber="277" unit="in" value="142.5" valueMax="149.0" valueMin="136.0">136–149 in</quantity>
|
||
<taxonomicName id="AFDD4D5AFFDB8D36E836DC38C9350188" box="[370,533,1652,1676]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nigroviridis">
|
||
<emphasis id="5AA9EACBFFDB8D36E836DC38C9350188" box="[370,533,1652,1676]" italics="true" pageId="6" pageNumber="277">B. nigroviridis</emphasis>
|
||
</taxonomicName>
|
||
), thin, often kidney-shaped supraoculars with a wide intersupraocular space (
|
||
<taxonomicName id="AFDD4D5AFFDB8D36ECC0DC38CA3A01B4" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nigroviridis">
|
||
<emphasis id="5AA9EACBFFDB8D36ECC0DC38CA3A01B4" italics="true" pageId="6" pageNumber="277">B. nigroviridis</emphasis>
|
||
</taxonomicName>
|
||
usually have wider supraoculars, never kidney-shaped, with narrow intersupraocular space), and higher average counts of interrictals, dorsals, and subcaudal scales than
|
||
<taxonomicName id="AFDD4D5AFFDB8D36EAF8DCF0CF7F01D0" box="[956,1119,1724,1748]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="277" phylum="Chordata" rank="species" species="nigroviridis">
|
||
<emphasis id="5AA9EACBFFDB8D36EAF8DCF0CF7F01D0" box="[956,1119,1724,1748]" italics="true" pageId="6" pageNumber="277">B. nigroviridis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="20C76552FFDB8D37E983DC92CBFE00F1" lastPageId="7" lastPageNumber="278" pageId="6" pageNumber="277" type="description">
|
||
<paragraph id="686236D9FFDB8D37E983DC92CFDD06D4" blockId="6.[151,1437,895,2037]" lastBlockId="7.[151,1436,151,464]" lastPageId="7" lastPageNumber="278" pageId="6" pageNumber="277">
|
||
<emphasis id="5AA9EACBFFDB8D36E983DC92CAC701FC" bold="true" box="[199,487,1759,1784]" pageId="6" pageNumber="277">
|
||
Description of
|
||
<typeStatus id="B766887BFFDB8D36E83DDC92CAC201FC" box="[377,482,1759,1784]" pageId="6" pageNumber="277" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
An adult female; SVL
|
||
<quantity id="AF259B3CFFDB8D36EBB7DCADC87501FC" box="[755,853,1760,1784]" metricMagnitude="-1" metricUnit="m" metricValue="7.79" pageId="6" pageNumber="277" unit="mm" value="779.0">779 mm</quantity>
|
||
; tail length
|
||
<quantity id="AF259B3CFFDB8D36EA98DCADCF7101FC" box="[988,1105,1760,1784]" metricMagnitude="-1" metricUnit="m" metricValue="1.3219999999999998" pageId="6" pageNumber="277" unit="mm" value="132.2">132.2 mm</quantity>
|
||
, comprising 14.51% of total; head length
|
||
<quantity id="AF259B3CFFDB8D36E86CDD49CAB30018" box="[296,403,1796,1820]" metricMagnitude="-2" metricUnit="m" metricValue="3.8899999999999997" pageId="6" pageNumber="277" unit="mm" value="38.9">38.9 mm</quantity>
|
||
; maximum head width
|
||
<quantity id="AF259B3CFFDB8D36EBEEDD49C8360018" box="[682,790,1796,1820]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="6" pageNumber="277" unit="mm" value="23.8">23.8 mm</quantity>
|
||
; rostral scute broader than high (4.2 X
|
||
<quantity id="AF259B3CFFDB8D36EDA9DD49CE6B0018" box="[1261,1355,1796,1820]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="277" unit="mm" value="3.2">3.2 mm</quantity>
|
||
); nasal divided above and below nostril, nasal fused with first supralabial on right side; loreal 1/1, contacting canthal, upper preocular, supralacunal, prelacunal, and two prefoveals; prefoveals 6/8; subfoveals absent; postfoveals 3/4; prelacunal large, contacting third supralabial on right side, second and third supralabials on left side; sublacunals 2/ 2; postlacunal absent because supralacunal and second sublacunal in contact; preoculars 3/3, upper large, middle about half size of upper, lower small and rounded; suboculars 2/2, anterior scale long; postoculars 3/3; loreal pit large, directed anteriorly, center of pit located slightly below line drawn from center of eye to naris and approximately halfway between center of eye and naris; supralabials 11/11 (including fusion of nasal with first supralabial on right side); infralabials 10/11, first pair meet posteriorly. Mental broader than long (3.6 X
|
||
<quantity id="AF259B3CFFDA8D37EC71DAF1CEAF07D0" box="[1333,1423,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="278" unit="mm" value="2.9">2.9 mm</quantity>
|
||
); three pairs of chin shields flanking mental groove, first two pairs contacting infralabials; five gulars between chin shields and preventrals; three preventrals. Internasals 3 anteriorly; canthals 2/2, anterior above nasal, posterior contacting loreal and upper preocular; 1/2 small scales between posterior canthal and supraocular; three scales between anterior canthal pair; four scales between posterior canthal pair; superciliaries absent; supraoculars narrow, approximately three times longer than broad; intersupraoculars nine; scales in parietal region small and keeled; interrictals 23. Dorsal scale rows 21-21-17; ventrals 156; cloacal entire; subcaudals 53, first divided; tail spine short, blunt, 1 ½ times as long as preceding subcaudal scale, covered by scales; tail prehensile.
|
||
</paragraph>
|
||
<caption id="3CA26651FFDA8D37E9D3DBB1CFCA0537" ID-Table-UUID="3CA26651FFDA8D37E9D3DBB1CFCA0537" httpUri="http://table.plazi.org/id/3CA26651FFDA8D37E9D3DBB1CFCA0537" pageId="7" pageNumber="278" targetBox="[159,1254,590,943]" targetIsTable="true" targetPageId="7">
|
||
<paragraph id="686236D9FFDA8D37E9D3DBB1CFCA0537" blockId="7.[151,1436,507,563]" pageId="7" pageNumber="278">
|
||
<emphasis id="5AA9EACBFFDA8D37E9D3DBB1CA2A0517" bold="true" box="[151,266,508,531]" pageId="7" pageNumber="278">TABLE 1.</emphasis>
|
||
Comparison of morphological characters of
|
||
<taxonomicName id="AFDD4D5AFFDA8D37EBACDBB6C8C70517" box="[744,999,507,531]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="278" phylum="Chordata" rank="species" species="nigroviridis">
|
||
<emphasis id="5AA9EACBFFDA8D37EBACDBB6C8C70517" box="[744,999,507,531]" italics="true" pageId="7" pageNumber="278">Bothriechis nigroviridis</emphasis>
|
||
</taxonomicName>
|
||
(n = 41) and
|
||
<taxonomicName id="AFDD4D5AFFDA8D37ED3CDBB1CFD70517" box="[1144,1271,507,531]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="278" phylum="Chordata" rank="species" species="nubestris">
|
||
<emphasis id="5AA9EACBFFDA8D37ED3CDBB1CFD70517" box="[1144,1271,507,531]" italics="true" pageId="7" pageNumber="278">B. nubestris</emphasis>
|
||
</taxonomicName>
|
||
(n = 27) (some specimens could not have all characters scored because of damage). The format is Range; Mode; Mean.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="686236D9FFDA8D37E9DBD803CF9304AB" pageId="7" pageNumber="278">
|
||
<table id="1ADDC479FFDA72CFE9DBD803CFC604AB" box="[159,1254,590,943]" gridcols="3" gridrows="9" pageId="7" pageNumber="278">
|
||
<tr id="D6ED349BFFDA72CFE9DBD803CFC60561" box="[159,1254,590,613]" gridrow="0" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFE9DBD803CA830561" box="[159,419,590,613]" gridcol="0" gridrow="0" pageId="7" pageNumber="278">Character</th>
|
||
<emphasis id="5AA9EACBFFDA8D37EB39D81DCFA00560" box="[637,1152,590,613]" italics="true" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFEB39D803C87D0561" box="[637,861,590,613]" gridcol="1" gridrow="0" pageId="7" pageNumber="278">
|
||
<taxonomicName id="AFDD4D5AFFDA8D37EB39D81DC8310560" box="[637,785,590,613]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="278" phylum="Chordata" rank="species" species="nigroviridis">
|
||
<emphasis id="5AA9EACBFFDA8D37EB39D81DC8310560" box="[637,785,590,613]" italics="true" pageId="7" pageNumber="278">B. nigroviridis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="953C5DE7FFDA72CFED43D803CFC60561" box="[1031,1254,590,613]" gridcol="2" gridrow="0" pageId="7" pageNumber="278">
|
||
<taxonomicName id="AFDD4D5AFFDA8D37ED43D81DCFA00560" box="[1031,1152,590,613]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="278" phylum="Chordata" rank="species" species="nubestris">
|
||
<emphasis id="5AA9EACBFFDA8D37ED43D81DCFA00560" box="[1031,1152,590,613]" italics="true" pageId="7" pageNumber="278">B. nubestris</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</emphasis>
|
||
</tr>
|
||
<tr id="D6ED349BFFDA72CFE9DBD835CFC6058A" box="[159,1254,632,654]" gridrow="1" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFE9DBD835CA83058A" box="[159,419,632,654]" gridcol="0" gridrow="1" pageId="7" pageNumber="278">Dorsal scales--First Third</th>
|
||
<td id="953C5DE7FFDA72CFEB39D835C87D058A" box="[637,861,632,654]" gridcol="1" gridrow="1" pageId="7" pageNumber="278">17–21; 19; 18.90</td>
|
||
<td id="953C5DE7FFDA72CFED43D835CFC6058A" box="[1031,1254,632,654]" gridcol="2" gridrow="1" pageId="7" pageNumber="278">19–23; 21; 20.93</td>
|
||
</tr>
|
||
<tr id="D6ED349BFFDA72CFE9DBD8ECCFC605B3" box="[159,1254,673,695]" gridrow="2" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFE9DBD8ECCA8305B3" box="[159,419,673,695]" gridcol="0" gridrow="2" pageId="7" pageNumber="278">Dorsal scales--Midbody</th>
|
||
<td id="953C5DE7FFDA72CFEB39D8ECC87D05B3" box="[637,861,673,695]" gridcol="1" gridrow="2" pageId="7" pageNumber="278">17–21; 19; 18.90</td>
|
||
<td id="953C5DE7FFDA72CFED43D8ECCFC605B3" box="[1031,1254,673,695]" gridcol="2" gridrow="2" pageId="7" pageNumber="278">17–23; 21; 20.21</td>
|
||
</tr>
|
||
<tr id="D6ED349BFFDA72CFE9DBD887CFC605E4" box="[159,1254,714,736]" gridrow="3" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFE9DBD887CA8305E4" box="[159,419,714,736]" gridcol="0" gridrow="3" pageId="7" pageNumber="278">Dorsal scales--Last Third</th>
|
||
<td id="953C5DE7FFDA72CFEB39D887C87D05E4" box="[637,861,714,736]" gridcol="1" gridrow="3" pageId="7" pageNumber="278">15–19; 15; 15.82</td>
|
||
<td id="953C5DE7FFDA72CFED43D887CFC605E4" box="[1031,1254,714,736]" gridcol="2" gridrow="3" pageId="7" pageNumber="278">15–17; 17; 16.71</td>
|
||
</tr>
|
||
<tr id="D6ED349BFFDA72CFE9DBD8B9CFC6040E" box="[159,1254,756,778]" gridrow="4" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFE9DBD8B9CA83040E" box="[159,419,756,778]" gridcol="0" gridrow="4" pageId="7" pageNumber="278">Ventral scales</th>
|
||
<td id="953C5DE7FFDA72CFEB39D8B9C87D040E" box="[637,861,756,778]" gridcol="1" gridrow="4" pageId="7" pageNumber="278">136–149; 141; 142.73</td>
|
||
<td id="953C5DE7FFDA72CFED43D8B9CFC6040E" box="[1031,1254,756,778]" gridcol="2" gridrow="4" pageId="7" pageNumber="278">150–160; 155; 154.64</td>
|
||
</tr>
|
||
<tr id="D6ED349BFFDA72CFE9DBD950CFC60437" box="[159,1254,797,819]" gridrow="5" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFE9DBD950CA830437" box="[159,419,797,819]" gridcol="0" gridrow="5" pageId="7" pageNumber="278">Subcaudal scales</th>
|
||
<td id="953C5DE7FFDA72CFEB39D950C87D0437" box="[637,861,797,819]" gridcol="1" gridrow="5" pageId="7" pageNumber="278">48–64; 50; 52.79</td>
|
||
<td id="953C5DE7FFDA72CFED43D950CFC60437" box="[1031,1254,797,819]" gridcol="2" gridrow="5" pageId="7" pageNumber="278">52–64; 59; 57.41</td>
|
||
</tr>
|
||
<tr id="D6ED349BFFDA72CFE9DBD90BCFC60458" box="[159,1254,838,860]" gridrow="6" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFE9DBD90BCA830458" box="[159,419,838,860]" gridcol="0" gridrow="6" pageId="7" pageNumber="278">Supralabials</th>
|
||
<td id="953C5DE7FFDA72CFEB39D90BC87D0458" box="[637,861,838,860]" gridcol="1" gridrow="6" pageId="7" pageNumber="278">9–11; 10; 9.76</td>
|
||
<td id="953C5DE7FFDA72CFED43D90BCFC60458" box="[1031,1254,838,860]" gridcol="2" gridrow="6" pageId="7" pageNumber="278">9–11; 10; 9.62</td>
|
||
</tr>
|
||
<tr id="D6ED349BFFDA72CFE9DBD93DCFC60482" box="[159,1254,880,902]" gridrow="7" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFE9DBD93DCA830482" box="[159,419,880,902]" gridcol="0" gridrow="7" pageId="7" pageNumber="278">Intersupraoculars</th>
|
||
<td id="953C5DE7FFDA72CFEB39D93DC87D0482" box="[637,861,880,902]" gridcol="1" gridrow="7" pageId="7" pageNumber="278">3–10; 6; 6.34</td>
|
||
<td id="953C5DE7FFDA72CFED43D93DCFC60482" box="[1031,1254,880,902]" gridcol="2" gridrow="7" pageId="7" pageNumber="278">6–10; 8; 7.83</td>
|
||
</tr>
|
||
<tr id="D6ED349BFFDA72CFE9DBD9D4CFC604AB" box="[159,1254,921,943]" gridrow="8" pageId="7" pageNumber="278">
|
||
<th id="953C5DE7FFDA72CFE9DBD9D4CA8304AB" box="[159,419,921,943]" gridcol="0" gridrow="8" pageId="7" pageNumber="278">Interrictals</th>
|
||
<td id="953C5DE7FFDA72CFEB39D9D4C87D04AB" box="[637,861,921,943]" gridcol="1" gridrow="8" pageId="7" pageNumber="278">19–26; 22; 21.90</td>
|
||
<td id="953C5DE7FFDA72CFED43D9D4CFC604AB" box="[1031,1254,921,943]" gridcol="2" gridrow="8" pageId="7" pageNumber="278">22–29; 24; 25.07</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="686236D9FFDA8D37E983D9A1CA5003BC" blockId="7.[151,1437,1004,2037]" pageId="7" pageNumber="278">
|
||
<emphasis id="5AA9EACBFFDA8D37E983D9A1C9270301" bold="true" box="[199,519,1004,1029]" pageId="7" pageNumber="278">Coloration in preservative.</emphasis>
|
||
Dorsum of head and body black and pale green mottled; dorsal surface of tail less mottled, grey-green speckled with black, last third yellowish brown lightly speckled with black; postocular stripe extending to rictus, above stripe faint black longitudinal striping, below stripe yellowish with some black blotches on upper lip; ventral surface of head and body yellow, becoming more yellowish green posteriorly. Subcaudal region grey-green with black speckling fading to brown with faint black speckling near tip. Coloration in life of
|
||
<typeStatus id="B766887BFFDA8D37E9D3DED2CBD903BC" box="[151,249,1183,1208]" pageId="7" pageNumber="278" type="holotype">holotype</typeStatus>
|
||
unknown.
|
||
</paragraph>
|
||
<paragraph id="686236D9FFDA8D37E983DE89C9980221" blockId="7.[151,1437,1004,2037]" pageId="7" pageNumber="278">
|
||
<emphasis id="5AA9EACBFFDA8D37E983DE89CA1B03D9" bold="true" box="[199,315,1220,1245]" pageId="7" pageNumber="278">Variation.</emphasis>
|
||
Dorsals were most commonly arranged in 21–21–17 (42.9% of specimens), whereas each of the following combinations were also observed: 23–23–17, 23–21–17, 21–21–15, 21–20–17, 21–19–17, 21–19–15, 19–21–17, 19–19–17, 19–17–17, and 19–17–15.
|
||
</paragraph>
|
||
<paragraph id="686236D9FFDA8D37E983DF62CE680294" blockId="7.[151,1437,1004,2037]" pageId="7" pageNumber="278">
|
||
Ventral scales range 151–157 (mean = 153.7) in males and
|
||
<quantity id="AF259B3CFFDA8D37EA3DDF62C8DB024C" box="[889,1019,1327,1352]" metricMagnitude="0" metricUnit="m" metricValue="3.9624" metricValueMax="4.064" metricValueMin="3.8608" pageId="7" pageNumber="278" unit="in" value="156.0" valueMax="160.0" valueMin="152.0">152–160 in</quantity>
|
||
females (mean = 155.4). Subcaudal scales range 53–62 (mean = 58.6) in males and 52–64 (mean = 56.2) in females. Tail length comprises 14–17% (mean = 15.2%) of total length in males; tail comprises 14–16% (mean = 15.0%) of total length in females.
|
||
</paragraph>
|
||
<paragraph id="686236D9FFDA8D37E983DFD1C87502F8" blockId="7.[151,1437,1004,2037]" pageId="7" pageNumber="278">
|
||
Head scale variation is as follows: prefoveals 4–7; subfoveals 0–1 (present in one specimen only, all others lack subfoveals); postfoveals 1–4; suboculars 1–4; postoculars 2–3; supralabials 9–11; infralabials 9–12; internasals 2–4; intercanthals 4–7; interrictals 22–29 (
|
||
<tableCitation id="255F0362FFDA8D37EBB4DFA9C86402F8" box="[752,836,1508,1533]" captionStart="TABLE 1" captionStartId="7.[151,239,508,531]" captionTargetBox="[159,1254,590,943]" captionTargetPageId="7" captionText="TABLE 1. Comparison of morphological characters of Bothriechis nigroviridis (n = 41) and B. nubestris (n = 27) (some specimens could not have all characters scored because of damage). The format is Range; Mode; Mean." httpUri="http://table.plazi.org/id/3CA26651FFDA8D37E9D3DBB1CFCA0537" pageId="7" pageNumber="278" tableUuid="3CA26651FFDA8D37E9D3DBB1CFCA0537">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="686236D9FFDA8D37E983DC4ACF460019" blockId="7.[151,1437,1004,2037]" pageId="7" pageNumber="278">
|
||
Coloration in preservative varied among specimens and consisted of two rather distinct morphs, with few specimens showing intermediate coloration. Twenty-six percent of the specimens had nearly identical coloration to the
|
||
<typeStatus id="B766887BFFDA8D37E985DC02CA07016C" box="[193,295,1615,1640]" pageId="7" pageNumber="278" type="holotype">holotype</typeStatus>
|
||
. Sixty percent of the specimens displayed a darker color morph with dorsal green being replaced by dark grey and the yellowish venter being replaced by beige, fading to grey-green on the tail. The same ventral pattern is present is both morphs but the yellow is replaced by beige in the darker morph. In two specimens, the dorsum is the “darker morph” but the venter is like the holotype’s with yellow. Recently-born juveniles had brown and black dorsums with beige-brown venters and the tail tips were completely brownish yellow. Coloration in life of three known specimens is nearly identical to the
|
||
<typeStatus id="B766887BFFDA8D37EB93DD49C8190019" box="[727,825,1796,1821]" pageId="7" pageNumber="278" type="holotype">holotype</typeStatus>
|
||
coloration in preservative.
|
||
</paragraph>
|
||
<paragraph id="686236D9FFDA8D37E983DD6ACBFE00F1" blockId="7.[151,1437,1004,2037]" pageId="7" pageNumber="278">
|
||
<emphasis id="5AA9EACBFFDA8D37E983DD6ACA720044" bold="true" box="[199,338,1831,1856]" pageId="7" pageNumber="278">Hemipenes.</emphasis>
|
||
<typeStatus id="B766887BFFDA8D37E81FDD64CA9D0045" box="[347,445,1833,1857]" pageId="7" pageNumber="278" type="paratype">Paratype</typeStatus>
|
||
UCR 11151 had fully everted hemipenes. The total length of the right hemipenis was equal to the length of the first five subcaudal scales. The two lobes of the organ bifurcate at an approximate length of two subcaudals and the sulcus spermaticus divides at a distance of approximately ½ the length of a subcaudal. The base of the organ is spinous, whereas the distal portion is covered in calyces. Spines are dense and evenly dispersed; some spines are quite large. Calyces begin approximately 1 ½ subcaudal scale lengths from the tip of the organ.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="3CA26651FFD58D38E9D3DDECC92300D2" httpUri="https://zenodo.org/record/255354/files/figure.png" pageId="8" pageNumber="279" targetBox="[369,1215,193,1931]" targetPageId="8">
|
||
<paragraph id="686236D9FFD58D38E9D3DDECC92300D2" blockId="8.[151,1436,1953,2006]" pageId="8" pageNumber="279">
|
||
<emphasis id="5AA9EACBFFD58D38E9D3DDECCA3900B2" bold="true" box="[151,281,1953,1975]" pageId="8" pageNumber="279">FIGURE 4.</emphasis>
|
||
Photographs of
|
||
<taxonomicName id="AFDD4D5AFFD58D38E896DDECC99000B3" box="[466,688,1953,1975]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="279" phylum="Chordata" rank="species" species="nubestris" status="sp. nov.">
|
||
<emphasis id="5AA9EACBFFD58D38E896DDECC99000B3" box="[466,688,1953,1975]" italics="true" pageId="8" pageNumber="279">Bothriechis nubestris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="419A57B0FFD58D38EBF8DDEFC9DC00B3" box="[700,764,1954,1975]" pageId="8" pageNumber="279" rank="species">
|
||
<emphasis id="5AA9EACBFFD58D38EBF8DDEFC9DC00B3" bold="true" box="[700,764,1954,1975]" pageId="8" pageNumber="279">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in life. Top photographer: John Tashjian; middle and bottom photographer Wayne van Devender.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3CA26651FFD48D39E9D3DC37CEBD01AB" httpUri="https://zenodo.org/record/255355/files/figure.png" pageId="9" pageNumber="280" targetBox="[168,1432,225,1636]" targetPageId="9">
|
||
<paragraph id="686236D9FFD48D39E9D3DC37CEBD01AB" blockId="9.[151,1437,1658,1711]" pageId="9" pageNumber="280">
|
||
<emphasis id="5AA9EACBFFD48D39E9D3DC37CA33018B" bold="true" box="[151,275,1658,1680]" pageId="9" pageNumber="280">FIGURE 5.</emphasis>
|
||
Dorsal and lateral views of the head of male paratype UCR 11151 from San Gerardo, Costa Rica. Scale bar = 20 mm. The diagnostic kidney-shaped supraoculars and partial scale rows between the supralabials and suboculars are highlighted.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="20C76552FFD48D39E983DC9BC81E00EF" pageId="9" pageNumber="280" type="distribution">
|
||
<paragraph id="686236D9FFD48D39E983DC9BC81E00EF" blockId="9.[151,1436,1750,2027]" pageId="9" pageNumber="280">
|
||
<emphasis id="5AA9EACBFFD48D39E983DC9BC97301EB" bold="true" box="[199,595,1750,1775]" pageId="9" pageNumber="280">Distribution and natural history.</emphasis>
|
||
<taxonomicName id="AFDD4D5AFFD48D39EB18DC9BC86C01EA" box="[604,844,1750,1774]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="280" phylum="Chordata" rank="species" species="nubestris">
|
||
<emphasis id="5AA9EACBFFD48D39EB18DC9BC86C01EA" box="[604,844,1750,1774]" italics="true" pageId="9" pageNumber="280">Bothriechis nubestris</emphasis>
|
||
</taxonomicName>
|
||
is known from the northern and central portions of the Cordillera de Talamanca of
|
||
<collectingCountry id="10CA7649FFD48D39EB46DCB6C95E0017" box="[514,638,1786,1811]" name="Costa Rica" pageId="9" pageNumber="280">Costa Rica</collectingCountry>
|
||
in the provinces of San José, Cartago, and Limón (see
|
||
<figureCitation id="F0E62A5CFFD48D39EDBADCB7CE630017" box="[1278,1347,1786,1811]" captionStart="FIGURE 6" captionStartId="10.[151,250,1349,1370]" captionTargetBox="[151,1436,339,1327]" captionTargetId="figure@10.[151,1436,339,1327]" captionTargetPageId="10" captionText="FIGURE 6. Topographical map indicating localities of Bothriechis nigroviridis (black symbols) and B. nubestris sp. n. (blue symbols) used in this study: 72 used in morphological analyses (circles), 21 used in genetic analyses (squares), and 3 used in both (triangles)." httpUri="https://zenodo.org/record/255356/files/figure.png" pageId="9" pageNumber="280">Fig. 6</figureCitation>
|
||
). Three specimens (UCR 15422, 15428, 15429) have locality data that indicate they are from the Cordillera Central, but they were snakes donated by locals to the Instituto Clodomiro Picado and it is possible that the locality data were recorded in error. We exclude these localities from consideration here, but acknowledge that if these localities are correct, the species also occurs in the Cordillera Central. The species has been recorded from
|
||
<quantity id="AF259B3CFFD48D39EDFDDDC6CE2F00A6" box="[1209,1295,1931,1955]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="280" unit="m" value="2400.0">2400 m</quantity>
|
||
on Cerro de la Muerte to over
|
||
<quantity id="AF259B3CFFD48D39E81BDDE2CA9300C2" box="[351,435,1967,1991]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="280" unit="m" value="3000.0">3000 m</quantity>
|
||
in San Gerardo de Dota.
|
||
<taxonomicName id="AFDD4D5AFFD48D39EB95DDE3C8E000C2" box="[721,960,1966,1990]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="280" phylum="Chordata" rank="species" species="nubestris">
|
||
<emphasis id="5AA9EACBFFD48D39EB95DDE3C8E000C2" box="[721,960,1966,1990]" italics="true" pageId="9" pageNumber="280">Bothriechis nubestris</emphasis>
|
||
</taxonomicName>
|
||
is an arboreal species usually found in the transition zone between cloud forest and montane rainforest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="20C76552FFD78D3AE983DADACA380619" pageId="10" pageNumber="281" type="etymology">
|
||
<paragraph id="686236D9FFD78D3AE983DADACA380619" blockId="10.[151,1436,151,285]" pageId="10" pageNumber="281">
|
||
<emphasis id="5AA9EACBFFD78D3AE983DADACA6E07B4" bold="true" box="[199,334,151,176]" pageId="10" pageNumber="281">Etymology.</emphasis>
|
||
The specific epithet means ‘belonging to the clouds’. It is derived from the Latin noun
|
||
<emphasis id="5AA9EACBFFD78D3AEC61DAD5CEB507AB" box="[1317,1429,152,176]" italics="true" pageId="10" pageNumber="281">nubes, –is</emphasis>
|
||
, meaning cloud, and the Latin suffix
|
||
<emphasis id="5AA9EACBFFD78D3AEB71DAF0C9A007D0" box="[565,640,189,212]" italics="true" pageId="10" pageNumber="281">–estris</emphasis>
|
||
, meaning belonging to. This name alludes to the fact that this species inhabits cloud forests. The common name Talamancan Palm-Pitviper refers to its range in the Cordillera de Talamanca.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |