178 lines
13 KiB
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178 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.94.85613" ID-Pensoft-Pub="1314-2607-94-163" ID-Pensoft-UUID="F9D136F20A2555CEA336E6E300207676" ID-ZooBank="0EA310AF80844448AEDA4CD39772A98B" ModsDocID="1314-2607-94-163" checkinTime="1671601408186" checkinUser="pensoft" docAuthor="Lepeco, Anderson, Barbosa, Diego N. & Melo, Gabriel A. R." docDate="2022" docId="B39C3C5B594F52D5B4081D0BF2689BD0" docLanguage="en" docName="JourHymenoptRes 94: 163-190" docOrigin="Journal of Hymenoptera Research 94" docPubDate="2022-12-20" docSource="http://dx.doi.org/10.3897/jhr.94.85613" docTitle="Trifionyx Lepeco & Melo 2022, gen. nov." docType="treatment" docUuid="C7CFFCD0-D5DF-43EF-9514-E5A12EFADBA5" docUuidSource="ZooBank" docVersion="1" id="F9D136F20A2555CEA336E6E300207676" lastPageNumber="163" masterDocId="F9D136F20A2555CEA336E6E300207676" masterDocTitle="A remarkable new family of stinging wasps from the Cretaceous of Myanmar and China (Hymenoptera, Aculeata)" masterLastPageNumber="190" masterPageNumber="163" pageNumber="163" updateTime="1671601408186" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A remarkable new family of stinging wasps from the Cretaceous of Myanmar and China (Hymenoptera, Aculeata)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lepeco, Anderson</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7467-5244</mods:nameIdentifier>
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<mods:affiliation>Laboratorio de Biologia Comparada de Hymenoptera, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Brazil & Laboratorio de Biologia Comparada e Abelhas, Departamento de Biologia, Faculdade de Filosofia, Ciencias e Letras de Ribeirao Preto, Universidade de Sao Paulo, Ribeirao Preto, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">al.lepeco@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Barbosa, Diego N.</mods:namePart>
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<mods:affiliation>Laboratorio de Biologia Comparada de Hymenoptera, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Melo, Gabriel A. R.</mods:namePart>
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<mods:affiliation>Laboratorio de Biologia Comparada de Hymenoptera, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>94</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>163</mods:start>
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<mods:end>190</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.94.85613</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.94.85613</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-94-163</mods:identifier>
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<mods:identifier type="ZooBank">0EA310AF80844448AEDA4CD39772A98B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F9D136F20A2555CEA336E6E300207676</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:C7CFFCD0-D5DF-43EF-9514-E5A12EFADBA5" httpUri="http://treatment.plazi.org/id/B39C3C5B594F52D5B4081D0BF2689BD0" lastPageNumber="163" pageId="0" pageNumber="163">
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<subSubSection pageId="0" pageNumber="163" type="nomenclature">
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<paragraph pageId="0" pageNumber="163">
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†
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<taxonomicName LSID="https://zoobank.org/C7CFFCD0-D5DF-43EF-9514-E5A12EFADBA5" authority="Lepeco & Melo" authorityName="Lepeco & Melo" authorityYear="2022" family="Trifionychidae" genus="Trifionyx" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Trifionyx" order="Hymenoptera" pageId="0" pageNumber="163" rank="genus" status="gen. nov.">Trifionyx Lepeco & Melo</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="163">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="163" type="type species">
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<paragraph pageId="0" pageNumber="163">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="163">
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†
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<taxonomicName authorityName="Lepeco & Melo" authorityYear="2022" family="Trifionychidae" genus="Trifionyx" kingdom="Animalia" lsidName="Trifionyx pilosus" order="Hymenoptera" pageId="0" pageNumber="163" rank="species" species="pilosus">
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<emphasis italics="true" pageId="0" pageNumber="163">Trifionyx pilosus</emphasis>
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</taxonomicName>
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Lepeco & Melo, sp. nov.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="163" type="diagnosis">
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<paragraph pageId="0" pageNumber="163">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="163">
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The single species allocated in this genus is differentiated from †
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<taxonomicName authorityName="Lepeco & Barbosa & Melo" authorityYear="2022" family="Trifionychidae" genus="Mirabythus" kingdom="Animalia" lsidName="Mirabythus" order="Hymenoptera" pageId="0" pageNumber="163" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="163">Mirabythus</emphasis>
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</taxonomicName>
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and †
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<taxonomicName authorityName="Lepeco & Melo" authorityYear="2022" family="Trifionychidae" genus="Trifionyximus" kingdom="Animalia" lsidName="Trifionyximus" order="Hymenoptera" pageId="0" pageNumber="163" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="163">Trifionyximus</emphasis>
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</taxonomicName>
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gen. nov. by the absence of cells enclosed by tubular veins in the forewing (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. † Trifionyx pilosus sp. nov. Holotype female (DZUP Bur- 1906) A dorsal view of head and mesosoma B dorsal view of mesosoma C ventral view of head D ventral view of mesosoma, habitus, ventral view E wings F detail of mesotarsal claw. Scale bars: 0.5 mm." figureDoi="10.3897/jhr.94.85613.figure5" httpUri="https://binary.pensoft.net/fig/784809" pageId="0" pageNumber="163">5F</figureCitation>
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) and the enlarged head, which is far wider than mesosoma. From †
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<taxonomicName authorityName="Lepeco & Melo" authorityYear="2022" family="Trifionychidae" genus="Prionaspidion" kingdom="Animalia" lsidName="Prionaspidion" order="Hymenoptera" pageId="0" pageNumber="163" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="163">Prionaspidion</emphasis>
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</taxonomicName>
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gen. nov. it can be differentiated by the trifid tarsal claws (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. † Trifionyx pilosus sp. nov. Holotype female (DZUP Bur- 1906) A dorsal view of head and mesosoma B dorsal view of mesosoma C ventral view of head D ventral view of mesosoma, habitus, ventral view E wings F detail of mesotarsal claw. Scale bars: 0.5 mm." figureDoi="10.3897/jhr.94.85613.figure5" httpUri="https://binary.pensoft.net/fig/784809" pageId="0" pageNumber="163">5G</figureCitation>
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); tibial spur formula 1-2-2; dorsal surface of propodeum with a transverse depression between spiracles (which may correspond to the metapostnotum); ocellar triangle below upper eye level and convex clypeus disc. †
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<taxonomicName authorityName="Lepeco & Melo" authorityYear="2022" family="Trifionychidae" genus="Trifionyx" kingdom="Animalia" lsidName="Trifionyx" order="Hymenoptera" pageId="0" pageNumber="163" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="163">Trifionyx</emphasis>
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</taxonomicName>
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gen. nov. is unique within †
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<taxonomicName family="Trifionychidae" lsidName="" pageId="0" pageNumber="163" rank="family">Trifionychidae</taxonomicName>
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fam. nov. in having the body with conspicuously developed pubescence, including on the compound eyes.
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</paragraph>
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<caption doi="10.3897/jhr.94.85613.figure5" httpUri="https://binary.pensoft.net/fig/784809" pageId="0" pageNumber="163" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="163">
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<emphasis bold="true" pageId="0" pageNumber="163">Figure 5.</emphasis>
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†
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<taxonomicName authorityName="Lepeco & Melo" authorityYear="2022" family="Trifionychidae" genus="Trifionyx" kingdom="Animalia" lsidName="Trifionyx pilosus" order="Hymenoptera" pageId="0" pageNumber="163" rank="species" species="pilosus">
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<emphasis italics="true" pageId="0" pageNumber="163">Trifionyx pilosus</emphasis>
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</taxonomicName>
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sp. nov. Holotype female (DZUP Bur-1906)
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<emphasis bold="true" pageId="0" pageNumber="163">A</emphasis>
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dorsal view of head and mesosoma
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<emphasis bold="true" pageId="0" pageNumber="163">B</emphasis>
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dorsal view of mesosoma
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<emphasis bold="true" pageId="0" pageNumber="163">C</emphasis>
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ventral view of head
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<emphasis bold="true" pageId="0" pageNumber="163">D</emphasis>
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ventral view of mesosoma, habitus, ventral view
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<emphasis bold="true" pageId="0" pageNumber="163">E</emphasis>
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wings
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<emphasis bold="true" pageId="0" pageNumber="163">F</emphasis>
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detail of mesotarsal claw. Scale bars: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="163" type="description">
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<paragraph pageId="0" pageNumber="163">Description.</paragraph>
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<paragraph pageId="0" pageNumber="163">
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<emphasis bold="true" pageId="0" pageNumber="163">
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<emphasis italics="true" pageId="0" pageNumber="163">Head</emphasis>
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.
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</emphasis>
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Enlarged, far wider than mesosoma; prognathous. Entire apical margin of clypeus with small denticles; disc of clypeus relatively large, convex. Frons wider than twice eye length, flat. Frontal line not indicated. Compound eye relatively small, bulging, covered with erect setae. Vertex arcuate in frontal view; extending behind lateral ocelli for at least 1.5 times length of ocellar triangle. Mid ocellus far below upper tangent of compound eye. Lower margin of lateral ocelli below upper eye tangent.
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<emphasis bold="true" pageId="0" pageNumber="163">
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<emphasis italics="true" pageId="0" pageNumber="163">Antenna</emphasis>
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.
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</emphasis>
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F1 3 times as long as maximum width, longer than F2; F2-F8 about twice as long as maximum width.
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<emphasis bold="true" pageId="0" pageNumber="163">
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<emphasis italics="true" pageId="0" pageNumber="163">Mesosoma</emphasis>
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.
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</emphasis>
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Pronotal collar relatively short, as long as half-length of mesoscutum; anterior surface of pronotum somewhat rounded in dorsal view, with transverse sulcus at mid-height; surface near posterior edge with transverse depression. Surface of mesoscutum between notauli slightly convex, level with lateral surfaces. Mesepisternum with a shalow transverse sulcus above mesepisternal pit. Metanotum with few short carinae on sublateral surfaces. Propodeum without box-like aspect, posterior slope convex; dorsal surface depressed transversally between spiracles.
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<emphasis bold="true" pageId="0" pageNumber="163">
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<emphasis italics="true" pageId="0" pageNumber="163">Legs</emphasis>
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.
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</emphasis>
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Arolia not enlarged.
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<emphasis bold="true" pageId="0" pageNumber="163">
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<emphasis italics="true" pageId="0" pageNumber="163">Forewing</emphasis>
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.
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</emphasis>
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Veins C and Sc+R completely fused. Pterostigma present. Marginal cell absent. Submarginal cell not enclosed by tubular veins. Radial, first cubital, and medial cells defined by nebulous veins. Vein M+Cu nebulous. Vein cu-a nebulous. Distal portion of vein Cu nebulous. Vein A spectral.
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<emphasis bold="true" pageId="0" pageNumber="163">
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<emphasis italics="true" pageId="0" pageNumber="163">Hindwing</emphasis>
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.
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</emphasis>
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Vein C present, nebulous. Vein A absent.
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<emphasis bold="true" pageId="0" pageNumber="163">
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<emphasis italics="true" pageId="0" pageNumber="163">Metasoma</emphasis>
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.
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</emphasis>
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Apparently short, shorter than mesosoma; segments apparently not telescoped within one another.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="163" type="etymology">
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<paragraph pageId="0" pageNumber="163">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="163">
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The name is derived from Latin and results from the combination of
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<emphasis italics="true" pageId="0" pageNumber="163">trifidus</emphasis>
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, meaning "cleft into three", and
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<emphasis italics="true" pageId="0" pageNumber="163">onyx</emphasis>
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, meaning
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<normalizedToken originalValue="“claw”">"claw"</normalizedToken>
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. The name is masculine.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |