203 lines
39 KiB
XML
203 lines
39 KiB
XML
<document id="345A7FE00AE62362A1D92D904B8FC9D6" ID-DOI="10.5281/zenodo.7666308" ID-ISSN="2305-2562" ID-Zenodo-Dep="7666308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677091000000" checkinUser="felipe" docAuthor="Redman, Guy T., Hamer, Michelle L. & Barraclough, David A." docDate="2003" docId="7E3D87A088128011FE4CBC0B000DF58B" docLanguage="en" docName="AfrInvertebr.44.2.203–277.pdf" docOrigin="African Invertebrates 44 (2)" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Harpagophora arida Redman & Hamer & Barraclough 2003, sp. n." docType="treatment" docVersion="3" lastPageNumber="217" masterDocId="8204FFD8881E801FFFECBF750212F119" masterDocTitle="Revision of the Harpagophoridae (Diplopoda, Spirostreptida) of southern Africa, including descriptions of five new species" masterLastPageNumber="277" masterPageNumber="203" pageNumber="215" updateTime="1699455364425" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F1726CD982A00B5BF4038D7B063405FE">Revision of the Harpagophoridae (Diplopoda, Spirostreptida) of southern Africa, including descriptions of five new species</mods:title>
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<mods:name id="E502E8A1E704B521C1034B32C9C2E9C2" type="personal">
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<mods:namePart id="6AF56C62DAA6B7FCA4B641441722BBD2">Redman, Guy T.</mods:namePart>
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<mods:namePart id="A80D115BF4404204F12345915DAD4FA0">Hamer, Michelle L.</mods:namePart>
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<mods:namePart id="DEF2B83D08963BAC6C32044127322174">Barraclough, David A.</mods:namePart>
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<mods:typeOfResource id="DEBF8902D2710277C50FF00D23831614">text</mods:typeOfResource>
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<mods:title id="FA5AC863CDC5E377EBD2D86D1B365E60">African Invertebrates</mods:title>
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<mods:part id="BC59CACA9B34FC30E8CF4F5D576C7EFE">
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<mods:date id="7C3659DCB910755851CF49350A7DF783">2003</mods:date>
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<mods:number id="D6BB15212AEEFBD618602727DE9A68C7">2003-12-31</mods:number>
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<mods:number id="5DE508FA42A1EE2353334536404FB6ED">44</mods:number>
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<treatment id="7E3D87A088128011FE4CBC0B000DF58B" ID-DOI="http://doi.org/10.5281/zenodo.7667034" ID-Zenodo-Dep="7667034" LSID="urn:lsid:plazi:treatment:7E3D87A088128011FE4CBC0B000DF58B" httpUri="http://treatment.plazi.org/id/7E3D87A088128011FE4CBC0B000DF58B" lastPageId="14" lastPageNumber="217" pageId="12" pageNumber="215">
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<subSubSection id="BE8E653D88128013FE4CBC0B012DF28E" box="[416,831,894,919]" pageId="12" pageNumber="215" type="nomenclature">
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<paragraph id="F62B36B688128013FE4CBC0B012DF28E" blockId="12.[144,1105,894,1671]" box="[416,831,894,919]" pageId="12" pageNumber="215">
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<heading id="AD6381DA88128013FE4CBC0B012DF28E" box="[416,831,894,919]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="215" reason="2">
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<taxonomicName id="31944D3588128013FE4CBC0B008AF28E" ID-CoL="3JSC6" authority="Redman & Hamer & Barraclough, 2003" authorityName="Redman & Hamer & Barraclough" authorityYear="2003" box="[416,664,894,919]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="215" phylum="Arthropoda" rank="species" species="arida" status="sp. nov.">
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<emphasis id="C4E0EAA488128013FE4CBC0B008AF28E" bold="true" box="[416,664,894,919]" pageId="12" pageNumber="215">Harpagophora arida</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="DFD357DF88128013FD73BC0B00C9F28E" box="[671,731,894,919]" pageId="12" pageNumber="215" rank="species">sp. n.</taxonomicNameLabel>
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Redman
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="BE8E653D88128011FDCFBCD3000DF58B" lastPageId="14" lastPageNumber="217" pageId="12" pageNumber="215" type="description">
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<paragraph id="F62B36B688128013FDCFBCD300AFF2A6" blockId="12.[144,1105,894,1671]" box="[547,701,934,959]" pageId="12" pageNumber="215">
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<figureCitation id="6EAF2A3388128013FDCFBCD3007AF2A6" box="[547,616,934,959]" captionStart="Fig" captionStartId="1.[182,215,845,865]" captionTargetBox="[251,1061,203,829]" captionTargetId="figure-7@1.[232,1095,193,840]" captionTargetPageId="1" captionText="Fig. 1. Distribution of Harpagophora, Metaphora and Poratophilus species in southern Africa. G = Gauteng; KZN = KwaZulu-Natal; L = Lesotho; MP = Mpumalanga; S = Swaziland; NW Prov = North West Province; W Cape = Western Cape." figureDoi="http://doi.org/10.5281/zenodo.7666310" httpUri="https://zenodo.org/record/7666310/files/figure.png" pageId="12" pageNumber="215">Figs 1</figureCitation>
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,
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<figureCitation id="6EAF2A3388128013FD9ABCD300AFF2A6" box="[630,701,934,959]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="12" pageNumber="215">20–26</figureCitation>
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</paragraph>
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<paragraph id="F62B36B688128013FF7CBCBB0019F51E" blockId="12.[144,1105,894,1671]" pageId="12" pageNumber="215">
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<materialsCitation id="46FC3CEB88128013FF7CBCBB0019F51E" collectingDate="1968-10" collectionCode="AMSA" collectorName="R. F. Lawrence" country="Namibia" location="Ais-Ais" pageId="12" pageNumber="215" specimenCount="1" typeStatus="holotype">
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Type material (examined):
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<typeStatus id="292F881488128013FE51BCBB0038F2FE" box="[445,554,974,999]" pageId="12" pageNumber="215" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="8E83762688128013FDDFBCBB00A0F2FE" box="[563,690,974,999]" name="Namibia" pageId="12" pageNumber="215">NAMIBIA</collectingCountry>
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: 1ơ,
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<location id="F34B606D88128013FD0ABCBB012FF2FE" LSID="urn:lsid:plazi:treatment:7E3D87A088128011FE4CBC0B000DF58B:F34B606D88128013FD0ABCBB012FF2FE" box="[742,829,974,999]" country="Namibia" name="Ais-Ais" pageId="12" pageNumber="215">Ais-Ais</location>
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[2717CB],
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<date id="822A107688128013FC2DBCBB0603F2FE" box="[961,1041,974,999]" pageId="12" pageNumber="215" value="1968-10">
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<collectingDate id="926EE99E88128013FC2DBCBB0603F2FE" box="[961,1041,974,999]" pageId="12" pageNumber="215" value="1968-10">x.1968</collectingDate>
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</date>
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,
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<collectorName id="5B61536088128013FBF7BCBB02ECF51E" pageId="12" pageNumber="215">R. F. Lawrence</collectorName>
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(
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<collectionCode id="9085AE7388128013FEE0BC9B034DF51E" box="[268,351,1006,1031]" country="South Africa" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="12" pageNumber="215">AMSA</collectionCode>
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uncatalogued).
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</materialsCitation>
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</paragraph>
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<paragraph id="F62B36B688128013FF7CBB6303DAF456" blockId="12.[144,1105,894,1671]" pageId="12" pageNumber="215">
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<emphasis id="C4E0EAA488128013FF7CBB630310F537" box="[144,258,1046,1070]" italics="true" pageId="12" pageNumber="215">Diagnosis</emphasis>
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: Larger femoral spine apically bifid with two divergent points of unequal length (
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<figureCitation id="6EAF2A3388128013FF04BB43032DF556" box="[232,319,1078,1103]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="12" pageNumber="215">Fig. 24</figureCitation>
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). In addition to two femoral spines, a third short tooth is present at flexure of telopodite (
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<figureCitation id="6EAF2A3388128013FE64BB2303CCF576" box="[392,478,1110,1135]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="12" pageNumber="215">Fig. 24</figureCitation>
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). Spine branch short, narrow, cylindrical and bifurcate, terminating in two spines of unequal length (
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<figureCitation id="6EAF2A3388128013FD9BBB0300D8F596" box="[631,714,1142,1167]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="12" pageNumber="215">Fig. 24</figureCitation>
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). Pectinophore an elongate, ribbonlike laminate plate curving apically. Length of pectinophore about half length of posterior limb of telopodite (
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<figureCitation id="6EAF2A3388128013FE9EBBC303DBF5D6" box="[370,457,1206,1231]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="12" pageNumber="215">Fig. 24</figureCitation>
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). Proximal margin of pectinophore dentate, while distal margin with several spiniform processes emerging closer to apex (
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<figureCitation id="6EAF2A3388128013FCB5BBA301B8F5F6" box="[857,938,1238,1263]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="12" pageNumber="215">Fig. 22</figureCitation>
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). Distal margin of telocoxite with shallow depression over centre of distal surface (
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<figureCitation id="6EAF2A3388128013FC79BB8301F9F416" box="[917,1003,1270,1295]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="12" pageNumber="215">Fig. 22</figureCitation>
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). Lateral margin of telocoxite folded back on itself (i.e. fold directed distally) in area around gonocoel opening (
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<figureCitation id="6EAF2A3388128013FE8BBA4303A9F456" box="[359,443,1334,1359]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="12" pageNumber="215">Fig. 22</figureCitation>
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).
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</paragraph>
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<paragraph id="F62B36B688128013FF7CBA2B0107F48E" blockId="12.[144,1105,894,1671]" pageId="12" pageNumber="215">
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Etymology: L.
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<taxonomicName id="31944D3588128013FED5BA2B0364F46F" authorityName="Redman & Hamer & Barraclough" authorityYear="2003" box="[313,374,1374,1398]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="215" phylum="Arthropoda" rank="species" species="arida">
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<emphasis id="C4E0EAA488128013FED5BA2B0364F46F" box="[313,374,1374,1398]" italics="true" pageId="12" pageNumber="215">arida</emphasis>
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</taxonomicName>
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= dry. Refers to the arid, desert
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<typeStatus id="292F881488128013FD35BA2B011BF46E" box="[729,777,1374,1399]" pageId="12" pageNumber="215">type</typeStatus>
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habitat of the
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<typeStatus id="292F881488128013FC44BA2B01CAF46E" box="[936,984,1374,1399]" pageId="12" pageNumber="215">type</typeStatus>
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specimen. Lawrence proposed the name (manuscript name) in 1968.
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</paragraph>
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<paragraph id="F62B36B688128013FF7CBAD30309F4A6" blockId="12.[144,1105,894,1671]" box="[144,283,1446,1471]" pageId="12" pageNumber="215">Description:</paragraph>
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<paragraph id="F62B36B688128013FF7CBABB0077F71E" blockId="12.[144,1105,894,1671]" pageId="12" pageNumber="215">
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<emphasis id="C4E0EAA488128013FF7CBABB0306F4FF" box="[144,276,1486,1510]" italics="true" pageId="12" pageNumber="215">Dimensions</emphasis>
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: Male, n = 1. Body width 8.2; collum width 8.1; body length 80.0; legs detached and missing; antenna length 5.7.
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</paragraph>
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<paragraph id="F62B36B688128013FF7CB97B03B5F73E" blockId="12.[144,1105,894,1671]" box="[144,423,1550,1575]" pageId="12" pageNumber="215">
|
||
<emphasis id="C4E0EAA488128013FF7CB97B0366F73F" box="[144,372,1550,1574]" italics="true" pageId="12" pageNumber="215">Number of segments</emphasis>
|
||
: 54.
|
||
</paragraph>
|
||
<paragraph id="F62B36B688128013FF7CB95B00C4F79E" blockId="12.[144,1105,894,1671]" pageId="12" pageNumber="215">
|
||
<emphasis id="C4E0EAA488128013FF7CB95B02CCF75F" box="[144,222,1582,1606]" italics="true" pageId="12" pageNumber="215">Colour</emphasis>
|
||
: Head and clypeus light brown. Collum brown laterally and light brown dorsally. Prozonites greyish (bleached), mesozonites light brown, metazonites brown.Anal valve, caudal spine, and legs brown, antennae light brown.
|
||
</paragraph>
|
||
<caption id="A2EB663E88138012FF5ABAEE038DF74E" ID-DOI="http://doi.org/10.5281/zenodo.7666320" ID-Zenodo-Dep="7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="13" pageNumber="216" startId="13.[182,221,1435,1455]" targetBox="[179,1169,254,1412]" targetPageId="13">
|
||
<paragraph id="F62B36B688138012FF5ABAEE038DF74E" blockId="13.[182,1143,1435,1623]" pageId="13" pageNumber="216">
|
||
Figs 20–26.
|
||
<taxonomicName id="31944D3588138012FECFBAEE03CDF4B6" authority="Redman & Hamer & Barraclough, 2003" authorityName="Redman & Hamer & Barraclough" authorityYear="2003" box="[291,479,1435,1455]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="216" phylum="Arthropoda" rank="species" species="arida" status="sp. nov.">
|
||
<emphasis id="C4E0EAA488138012FECFBAEE03CDF4B6" box="[291,479,1435,1455]" italics="true" pageId="13" pageNumber="216">Harpagophora arida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DFD357DF88138012FE0FBAEE0000F4B6" box="[483,530,1435,1455]" pageId="13" pageNumber="216" rank="species">sp. n.</taxonomicNameLabel>
|
||
20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F62B36B688108011FF7CBFBB03A1F1FE" blockId="14.[144,1104,206,1170]" box="[144,435,206,231]" pageId="14" pageNumber="217">
|
||
<emphasis id="C4E0EAA488108011FF7CBFBB0334F1FF" box="[144,294,206,230]" italics="true" pageId="14" pageNumber="217">First ozopore</emphasis>
|
||
: Segment 6.
|
||
</paragraph>
|
||
<paragraph id="F62B36B688108011FF7CBF9B008FF05E" blockId="14.[144,1104,206,1170]" pageId="14" pageNumber="217">
|
||
<emphasis id="C4E0EAA488108011FF7CBF9B02F0F01F" box="[144,226,238,262]" italics="true" pageId="14" pageNumber="217">Collum</emphasis>
|
||
: Anterior corner rounded and obtuse; posterior corner extending a little beyond the posterior border and overlapping second segment. One submarginal groove with smaller groove along vertical margin (
|
||
<figureCitation id="6EAF2A3388108011FDD0BE5B0082F05E" box="[572,656,302,327]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F62B36B688108011FF7CBE3B0605F25E" blockId="14.[144,1104,206,1170]" pageId="14" pageNumber="217">
|
||
<emphasis id="C4E0EAA488108011FF7CBE3B0310F07F" box="[144,258,334,358]" italics="true" pageId="14" pageNumber="217">Gonopods</emphasis>
|
||
: Longer femoral spine directed distally and partially concealed under laterally directed fold on aboral surface of telocoxite, apically bifid with two divergent points of unequal length (
|
||
<figureCitation id="6EAF2A3388108011FE83BEFB03D5F0BE" box="[367,455,398,423]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Fig. 24</figureCitation>
|
||
). Short, simple, partially curved, second spine wrapped around base of larger femoral spine and directed laterally. A third small spine present at flexure of telopodite (
|
||
<figureCitation id="6EAF2A3388108011FE4DBEBB03E4F0FE" box="[417,502,462,487]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Fig. 24</figureCitation>
|
||
). Pectinophore elongate, ribbon-like, curling distally. Length of pectinophore about half length of posterior limb of telopodite. Proximal margin of pectinophore dentate, while distal margin with several spiniform processes emerging closer to apex (
|
||
<figureCitation id="6EAF2A3388108011FE55BD5B0050F35E" box="[441,578,558,583]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Figs 22, 25</figureCitation>
|
||
). Spine branch short, narrow, cylindrical and apically bifid with two spines of unequal length, emerging halfway down telopodite (
|
||
<figureCitation id="6EAF2A3388108011FF74BD1B02FCF39E" box="[152,238,622,647]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Fig. 24</figureCitation>
|
||
). Telocoxal spines obscure and aborally directed (
|
||
<figureCitation id="6EAF2A3388108011FCC6BD1B0192F39E" box="[810,896,622,647]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Fig. 22</figureCitation>
|
||
). Distal margin of telocoxite simple, without medial protrusion but with shallow depression over centre of distal surface (indicated by arrow,
|
||
<figureCitation id="6EAF2A3388108011FDC0BDDB006CF3DE" box="[556,638,686,711]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Fig. 22</figureCitation>
|
||
). Lateral margin of telocoxite folded back on itself (i.e. fold directed distally) in area around gonocoel opening (indicated by broken arrow,
|
||
<figureCitation id="6EAF2A3388108011FED4BD9B039CF21E" box="[312,398,750,775]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Fig. 22</figureCitation>
|
||
). Gonocoel opening extending from opening where posterior limb of telopodite emerges, down lateral surface of gonopod. Aboral surface of telocoxite with shallow depression in which larger femoral spine rests (
|
||
<figureCitation id="6EAF2A3388108011FC59BC5B0618F25E" box="[949,1034,814,839]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F62B36B688108011FF7CBC3B03AFF29E" blockId="14.[144,1104,206,1170]" pageId="14" pageNumber="217">
|
||
<emphasis id="C4E0EAA488108011FF7CBC3B0333F27F" box="[144,289,846,870]" italics="true" pageId="14" pageNumber="217">Pre-anal ring</emphasis>
|
||
: Caudal spine short, not extending beyond margin of anal valve, moderately upturned distally (
|
||
<figureCitation id="6EAF2A3388108011FEB0BC1B03A2F29E" box="[348,432,878,903]" captionStart="Figs 20–26" captionStartId="13.[182,221,1435,1455]" captionTargetBox="[179,1169,254,1412]" captionTargetId="figure-8@13.[176,1171,251,1413]" captionTargetPageId="13" captionText="Figs 20–26. Harpagophora arida sp. n. 20. Lateral view of collum; arrow indicates anterior end of millipede. 21. Oral view of first pair of male legs. 22. Aboral view of gonopods; solid arrow indicates shallow depression in distal margin of telocoxal fold; broken arrow indicates lateral margin folded forwards. 23. Oral view of gonopods. 24.Aboral view of telopodite. 25. Detail of apical elements of telopodite. 26. Lateral view of posterior end of millipede, showing anal valve and pre-anal ring with spines. c = telocoxal spine; f1 = large femoral spine; f2 = small femoral spine; f3 = additional femoral spine; p = pectinophore; pfp = prefemoral fold; s = spine branch; te = telepodite." figureDoi="http://doi.org/10.5281/zenodo.7666320" httpUri="https://zenodo.org/record/7666320/files/figure.png" pageId="14" pageNumber="217">Fig. 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F62B36B688108011FF7CBCE101E5F2B4" blockId="14.[144,1104,206,1170]" box="[144,1015,916,941]" pageId="14" pageNumber="217">
|
||
Distribution: The species appears to be endemic to southern
|
||
<collectingCountry id="8E83762688108011FCD8BCE10186F2B4" box="[820,916,916,941]" name="Namibia" pageId="14" pageNumber="217">Namibia</collectingCountry>
|
||
(
|
||
<figureCitation id="6EAF2A3388108011FC48BCE101F7F2B4" box="[932,997,916,941]" captionStart="Fig" captionStartId="1.[182,215,845,865]" captionTargetBox="[251,1061,203,829]" captionTargetId="figure-7@1.[232,1095,193,840]" captionTargetPageId="1" captionText="Fig. 1. Distribution of Harpagophora, Metaphora and Poratophilus species in southern Africa. G = Gauteng; KZN = KwaZulu-Natal; L = Lesotho; MP = Mpumalanga; S = Swaziland; NW Prov = North West Province; W Cape = Western Cape." figureDoi="http://doi.org/10.5281/zenodo.7666310" httpUri="https://zenodo.org/record/7666310/files/figure.png" pageId="14" pageNumber="217">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F62B36B688108011FF7CBCCC000DF58B" blockId="14.[144,1104,206,1170]" pageId="14" pageNumber="217">
|
||
Remarks: This species differs from congeners in the structure of the telocoxite, pectinophore and spine branch, and the lateral margin of the telocoxite which is curled upwards (i.e. fold directed distally) in the area around the gonocoel opening. The latter character state is unique to
|
||
<taxonomicName id="31944D3588108011FE53BB6C000FF528" authorityName="Redman & Hamer & Barraclough" authorityYear="2003" box="[447,541,1049,1074]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="217" phylum="Arthropoda" rank="species" species="arida">
|
||
<emphasis id="C4E0EAA488108011FE53BB6C03C1F528" box="[447,467,1049,1073]" italics="true" pageId="14" pageNumber="217">H</emphasis>
|
||
.
|
||
<emphasis id="C4E0EAA488108011FE0CBB6C000FF528" box="[480,541,1049,1073]" italics="true" pageId="14" pageNumber="217">arida</emphasis>
|
||
</taxonomicName>
|
||
. The spine branch emerges closer to the flexure of the telopodite than in other
|
||
<taxonomicName id="31944D3588108011FE5BBB4C00A4F54B" box="[439,694,1081,1106]" pageId="14" pageNumber="217">
|
||
<emphasis id="C4E0EAA488108011FE5BBB4C004FF548" box="[439,605,1081,1105]" italics="true" pageId="14" pageNumber="217">Harpagophora</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, where these structures (pectinophore and spine branch) occur in close proximity to each other.
|
||
<taxonomicName id="31944D3588108011FCCDBB2C0190F568" authorityName="Redman & Hamer & Barraclough" authorityYear="2003" box="[801,898,1113,1137]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="217" phylum="Arthropoda" rank="species" species="arida">
|
||
<emphasis id="C4E0EAA488108011FCCDBB2C0190F568" box="[801,898,1113,1137]" italics="true" pageId="14" pageNumber="217">H. arida</emphasis>
|
||
</taxonomicName>
|
||
is bigger than the South African species of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |