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<document id="437504F18D71265E04463B4B1F6FE9DA" ID-DOI="10.5281/zenodo.7666308" ID-ISSN="2305-2562" ID-Zenodo-Dep="7666308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677091000000" checkinUser="felipe" docAuthor="Redman, Guy T., Hamer, Michelle L. &amp; Barraclough, David A." docDate="2003" docId="7E3D87A088078006FE1BBFBB0009F569" docLanguage="en" docName="AfrInvertebr.44.2.203–277.pdf" docOrigin="African Invertebrates 44 (2)" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Metaphora Redman &amp; Hamer &amp; Barraclough 2003, gen. n." docType="treatment" docVersion="3" lastPageNumber="228" masterDocId="8204FFD8881E801FFFECBF750212F119" masterDocTitle="Revision of the Harpagophoridae (Diplopoda, Spirostreptida) of southern Africa, including descriptions of five new species" masterLastPageNumber="277" masterPageNumber="203" pageNumber="228" updateTime="1699455364425" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D25429891A6DBE21044256CDFAFDA3C3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2E0A94AE4CB9B772B86CD155079A2CDC">Revision of the Harpagophoridae (Diplopoda, Spirostreptida) of southern Africa, including descriptions of five new species</mods:title>
</mods:titleInfo>
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<mods:role id="2407693B0192E7BA0950B457964596E0">
<mods:roleTerm id="D7E98F139A03B7E2C3FCB8A4EAFD59CE">Author</mods:roleTerm>
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<mods:namePart id="C42739E3E5E9648C6197C1FE0B9C0AFD">Redman, Guy T.</mods:namePart>
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<mods:namePart id="FDEFA9A23CB393418DB927D90E330F17">Hamer, Michelle L.</mods:namePart>
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<mods:namePart id="2E5EEAB695E64BE299275101C6B110E9">Barraclough, David A.</mods:namePart>
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<mods:titleInfo id="B9010E40662DE21DF86635DFA81A3864">
<mods:title id="C8C0097DF6C333B0BDEDACE283ED4A3E">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="F7CE0E42FBC562EF217F64200F9A90EB">
<mods:date id="A1AC4E5D9E6CE7741A4C4C17E5124CDC">2003</mods:date>
<mods:detail id="374CDAA93DC73426BCDF357EE77BC483" type="pubDate">
<mods:number id="9CEC0C06700EAF819231A7F5D678BA38">2003-12-31</mods:number>
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<mods:detail id="B51A8B54727C6D7EB460F94D938D153A" type="volume">
<mods:number id="B86DAEDE8807F9F486BA27550D2CD15A">44</mods:number>
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<mods:number id="5C8EFB2A5A21A36D5F93EC7A3BCAF59B">2</mods:number>
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<mods:classification id="E96280A201FD4F76D39288314A7AE413">journal article</mods:classification>
<mods:identifier id="9BCDDDBC445F64523003938B7350F556" type="DOI">10.5281/zenodo.7666308</mods:identifier>
<mods:identifier id="331A3B5B79B27252F61D5B41ED4C4C65" type="ISSN">2305-2562</mods:identifier>
<mods:identifier id="58CD50107B9ACF4FC09FCA6E69DD882C" type="Zenodo-Dep">7666308</mods:identifier>
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<treatment id="7E3D87A088078006FE1BBFBB0009F569" ID-DOI="http://doi.org/10.5281/zenodo.7667036" ID-Zenodo-Dep="7667036" LSID="urn:lsid:plazi:treatment:7E3D87A088078006FE1BBFBB0009F569" httpUri="http://treatment.plazi.org/id/7E3D87A088078006FE1BBFBB0009F569" lastPageNumber="228" pageId="25" pageNumber="228">
<subSubSection id="BE8E653D88078006FE1BBFBB0126F1FE" box="[503,820,206,231]" pageId="25" pageNumber="228" type="nomenclature">
<paragraph id="F62B36B688078006FE1BBFBB0126F1FE" blockId="25.[181,1142,206,639]" box="[503,820,206,231]" pageId="25" pageNumber="228">
<heading id="AD6381DA88078006FE1BBFBB0126F1FE" box="[503,820,206,231]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="228" reason="2">
<taxonomicName id="31944D3588078006FE1BBFBB006FF1FE" ID-CoL="7PNYK" authority="Redman &amp; Hamer &amp; Barraclough, 2003" authorityName="Redman &amp; Hamer &amp; Barraclough" authorityYear="2003" box="[503,637,206,231]" class="Diplopoda" family="Harpagophoridae" genus="Metaphora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C4E0EAA488078006FE1BBFBB006FF1FE" bold="true" box="[503,637,206,231]" pageId="25" pageNumber="228">Metaphora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DFD357DF88078006FD68BFBB00C2F1FE" box="[644,720,206,231]" pageId="25" pageNumber="228" rank="genus">gen. n.</taxonomicNameLabel>
Redman
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BE8E653D88078006FF59BF830009F569" pageId="25" pageNumber="228" type="materials_examined">
<paragraph id="F62B36B688078006FF59BF830662F016" blockId="25.[181,1142,206,639]" box="[181,1136,246,271]" pageId="25" pageNumber="228">
<typeStatus id="292F881488078006FF59BF8302FFF016" box="[181,237,246,271]" pageId="25" pageNumber="228">Type</typeStatus>
species here designated:
<taxonomicName id="31944D3588078006FDE6BF83067EF016" authority="(Karsch, 1881)" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[522,1132,246,271]" class="Diplopoda" family="Spirostreptidae" genus="Spirostreptus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="species" species="spirobolinus" subGenus="Nodopyge">
<emphasis id="C4E0EAA488078006FDE6BF83008FF017" box="[522,669,246,270]" italics="true" pageId="25" pageNumber="228">Spirostreptus</emphasis>
(
<emphasis id="C4E0EAA488078006FD41BF83010DF017" box="[685,799,246,270]" italics="true" pageId="25" pageNumber="228">Nodopyge</emphasis>
)
<emphasis id="C4E0EAA488078006FCC3BF8301A8F017" box="[815,954,246,270]" italics="true" pageId="25" pageNumber="228">spirobolinus</emphasis>
(
<bibRefCitation id="92054B4788078006FC25BF830671F016" author="KARSCH, F." box="[969,1123,246,271]" pageId="25" pageNumber="228" pagination="1 - 79" refId="ref35007" refString="KARSCH, F. 1881. Neue Juliden des Berliner Museums, als Prodromus einer Juliden Monographie. Zeitschrift fur die Gesamtenh Naturwissenschaften 54: 1 - 79." type="journal article" year="1881">Karsch, 1881</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="F62B36B688078006FF59BE6B0118F04E" blockId="25.[181,1142,206,639]" pageId="25" pageNumber="228">
Etymology: L.
<emphasis id="C4E0EAA488078006FEB0BE6B0380F02F" box="[348,402,286,310]" italics="true" pageId="25" pageNumber="228">meta</emphasis>
= conical or pyramidal structure. This refers to the conical shape of the lateral margin of the telocoxite. Gender feminine.
</paragraph>
<paragraph id="F62B36B688078006FF59BE130337F366" blockId="25.[181,1142,206,639]" pageId="25" pageNumber="228">
<emphasis id="C4E0EAA488078006FF59BE13033CF067" box="[181,302,358,382]" italics="true" pageId="25" pageNumber="228">Diagnosis</emphasis>
: Telopodite with two femoral spines, subequal in length (
<figureCitation id="6EAF2A3388078006FBE6BE130675F066" box="[1034,1127,358,383]" captionStart="Figures 59–67" captionStartId="26.[144,212,1432,1452]" captionTargetBox="[146,1097,232,1394]" captionTargetId="figure-11@26.[139,1110,270,1407]" captionTargetPageId="26" captionText="Figures 59–67. Metaphora spirobolina (Karsch, 1881). 59. Lateral view of anterior region of millipede. 60. Oral view of first pair of male legs. 61. Aboral view of gonopods (SAMC B7674). 62. Aboral view of gonopods (NMSA 19102). 63. Aboral view of gonopod (SAMC B7674) of different specimen, showing more obvious telocoxal spine. 64–66.Apical elements, different orientations. 67. Lateral view of anal valve and pre-anal ring. af = anterior telocoxal fold; br = additional branch on apical elements of telopodite; c = telecoxal spine; f1, f2 = femoral spines of telopodite; lp = lateral process of telocoxal fold; p = pectinophore; pf = posterior telocoxal fold; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666332" httpUri="https://zenodo.org/record/7666332/files/figure.png" pageId="25" pageNumber="228">Fig. 61</figureCitation>
). Posterior limb of telopodite comprising three branches apically: (a) elongate and ribbon-like pectinophore, (b) concave spine branch with dentate margin, (c) long, narrow, cylindrical, tapering spine with slight curvature and spiniform processes on its surface (
<figureCitation id="6EAF2A3388078006FE8ABE9303ADF0E6" box="[358,447,486,511]" captionStart="Figures 59–67" captionStartId="26.[144,212,1432,1452]" captionTargetBox="[146,1097,232,1394]" captionTargetId="figure-11@26.[139,1110,270,1407]" captionTargetPageId="26" captionText="Figures 59–67. Metaphora spirobolina (Karsch, 1881). 59. Lateral view of anterior region of millipede. 60. Oral view of first pair of male legs. 61. Aboral view of gonopods (SAMC B7674). 62. Aboral view of gonopods (NMSA 19102). 63. Aboral view of gonopod (SAMC B7674) of different specimen, showing more obvious telocoxal spine. 64–66.Apical elements, different orientations. 67. Lateral view of anal valve and pre-anal ring. af = anterior telocoxal fold; br = additional branch on apical elements of telopodite; c = telecoxal spine; f1, f2 = femoral spines of telopodite; lp = lateral process of telocoxal fold; p = pectinophore; pf = posterior telocoxal fold; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666332" httpUri="https://zenodo.org/record/7666332/files/figure.png" pageId="25" pageNumber="228">Fig. 65</figureCitation>
). Telocoxite reflexed, but fold on the distal rather than the lateral margin of telocoxite (
<figureCitation id="6EAF2A3388078006FE17BD730042F306" box="[507,592,518,543]" captionStart="Figures 59–67" captionStartId="26.[144,212,1432,1452]" captionTargetBox="[146,1097,232,1394]" captionTargetId="figure-11@26.[139,1110,270,1407]" captionTargetPageId="26" captionText="Figures 59–67. Metaphora spirobolina (Karsch, 1881). 59. Lateral view of anterior region of millipede. 60. Oral view of first pair of male legs. 61. Aboral view of gonopods (SAMC B7674). 62. Aboral view of gonopods (NMSA 19102). 63. Aboral view of gonopod (SAMC B7674) of different specimen, showing more obvious telocoxal spine. 64–66.Apical elements, different orientations. 67. Lateral view of anal valve and pre-anal ring. af = anterior telocoxal fold; br = additional branch on apical elements of telopodite; c = telecoxal spine; f1, f2 = femoral spines of telopodite; lp = lateral process of telocoxal fold; p = pectinophore; pf = posterior telocoxal fold; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666332" httpUri="https://zenodo.org/record/7666332/files/figure.png" pageId="25" pageNumber="228">Fig. 61</figureCitation>
). First pair of male legs with apex of prefemoral process subtriangular and directed orally obliquely at point of contact with coxal shelf. Syncoxosternum medially suturate between coxal elements. Coxal shelf tumid (
<figureCitation id="6EAF2A3388078006FF52BD130305F366" box="[190,279,614,639]" captionStart="Figures 59–67" captionStartId="26.[144,212,1432,1452]" captionTargetBox="[146,1097,232,1394]" captionTargetId="figure-11@26.[139,1110,270,1407]" captionTargetPageId="26" captionText="Figures 59–67. Metaphora spirobolina (Karsch, 1881). 59. Lateral view of anterior region of millipede. 60. Oral view of first pair of male legs. 61. Aboral view of gonopods (SAMC B7674). 62. Aboral view of gonopods (NMSA 19102). 63. Aboral view of gonopod (SAMC B7674) of different specimen, showing more obvious telocoxal spine. 64–66.Apical elements, different orientations. 67. Lateral view of anal valve and pre-anal ring. af = anterior telocoxal fold; br = additional branch on apical elements of telopodite; c = telecoxal spine; f1, f2 = femoral spines of telopodite; lp = lateral process of telocoxal fold; p = pectinophore; pf = posterior telocoxal fold; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666332" httpUri="https://zenodo.org/record/7666332/files/figure.png" pageId="25" pageNumber="228">Fig. 60</figureCitation>
).
</paragraph>
<paragraph id="F62B36B688078006FF59BDFA0302F3D1" blockId="25.[181,1142,655,1136]" pageId="25" pageNumber="228">
Distribution: Northern and
<collectingRegion id="3450F85488078006FE0EBDFA006EF3B1" box="[482,636,655,680]" country="South Africa" name="Western Cape" pageId="25" pageNumber="228">Western Cape</collectingRegion>
in the fynbos and the succulent Karoo biomes (
<figureCitation id="6EAF2A3388078006FF52BDDA02EDF3D1" box="[190,255,687,712]" captionStart="Fig" captionStartId="1.[182,215,845,865]" captionTargetBox="[251,1061,203,829]" captionTargetId="figure-7@1.[232,1095,193,840]" captionTargetPageId="1" captionText="Fig. 1. Distribution of Harpagophora, Metaphora and Poratophilus species in southern Africa. G = Gauteng; KZN = KwaZulu-Natal; L = Lesotho; MP = Mpumalanga; S = Swaziland; NW Prov = North West Province; W Cape = Western Cape." figureDoi="http://doi.org/10.5281/zenodo.7666310" httpUri="https://zenodo.org/record/7666310/files/figure.png" pageId="25" pageNumber="228">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="F62B36B688078006FF59BDA20009F569" blockId="25.[181,1142,655,1136]" pageId="25" pageNumber="228">
Remarks: The telocoxal folds are formed from the distal end of the telocoxite rather than from the lateral surface, as is observed in
<taxonomicName id="31944D3588078006FD07BD8D0171F209" authorityName="Chamberlain" authorityYear="1927" box="[747,867,760,784]" class="Diplopoda" family="Harpagophoridae" genus="Zinophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="C4E0EAA488078006FD07BD8D0171F209" box="[747,867,760,784]" italics="true" pageId="25" pageNumber="228">Zinophora</emphasis>
</taxonomicName>
and
<taxonomicName id="31944D3588078006FC4FBD8D062AF209" authorityName="Silvestri" authorityYear="1897" box="[931,1080,760,784]" class="Diplopoda" family="Harpagophoridae" genus="Poratophilus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="C4E0EAA488078006FC4FBD8D062AF209" box="[931,1080,760,784]" italics="true" pageId="25" pageNumber="228">Poratophilus</emphasis>
</taxonomicName>
. The pectinophore in
<taxonomicName id="31944D3588078006FE85BC6D001CF229" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[361,526,791,816]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="species" species="spirobolina">
<emphasis id="C4E0EAA488078006FE85BC6D0392F229" box="[361,384,792,816]" italics="true" pageId="25" pageNumber="228">M</emphasis>
.
<emphasis id="C4E0EAA488078006FE62BC6D001CF229" box="[398,526,792,816]" italics="true" pageId="25" pageNumber="228">spirobolina</emphasis>
</taxonomicName>
resembles that observed in
<taxonomicName id="31944D3588078006FCACBC6D018CF229" authorityName="Redman &amp; Hamer &amp; Barraclough" authorityYear="2003" box="[832,926,792,816]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="species" species="arida">
<emphasis id="C4E0EAA488078006FCACBC6D018CF229" box="[832,926,792,816]" italics="true" pageId="25" pageNumber="228">H. arida</emphasis>
</taxonomicName>
and
<taxonomicName id="31944D3588078006FC3FBC6D067DF229" authorityName="Attems" authorityYear="1909" box="[979,1135,791,816]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="species" species="diplocrada">
<emphasis id="C4E0EAA488078006FC3FBC6D01F5F229" box="[979,999,792,816]" italics="true" pageId="25" pageNumber="228">H</emphasis>
.
<emphasis id="C4E0EAA488078006FC18BC6D067DF229" box="[1012,1135,792,816]" italics="true" pageId="25" pageNumber="228">diplocrada</emphasis>
</taxonomicName>
, but the third branch (a narrow, curved and pointed spine directed towards other apical elements) clearly distinguishes
<taxonomicName id="31944D3588078006FDF1BC2D008BF269" authorityName="Redman &amp; Hamer &amp; Barraclough" authorityYear="2003" box="[541,665,856,880]" class="Diplopoda" family="Harpagophoridae" genus="Metaphora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="C4E0EAA488078006FDF1BC2D008BF269" box="[541,665,856,880]" italics="true" pageId="25" pageNumber="228">Metaphora</emphasis>
</taxonomicName>
from
<taxonomicName id="31944D3588078006FD0EBC2D0198F269" authorityName="Attems" authorityYear="1909" box="[738,906,856,880]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="C4E0EAA488078006FD0EBC2D0198F269" box="[738,906,856,880]" italics="true" pageId="25" pageNumber="228">Harpagophora</emphasis>
</taxonomicName>
. The third branch is also not the same as the third branch in
<taxonomicName id="31944D3588078006FD4ABC0D0132F289" authorityName="Chamberlain" authorityYear="1927" box="[678,800,888,912]" class="Diplopoda" family="Harpagophoridae" genus="Zinophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="C4E0EAA488078006FD4ABC0D0132F289" box="[678,800,888,912]" italics="true" pageId="25" pageNumber="228">Zinophora</emphasis>
</taxonomicName>
and
<taxonomicName id="31944D3588078006FC8DBC0D01EBF289" authorityName="Silvestri" authorityYear="1897" box="[865,1017,888,912]" class="Diplopoda" family="Harpagophoridae" genus="Poratophilus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="C4E0EAA488078006FC8DBC0D01EBF289" box="[865,1017,888,912]" italics="true" pageId="25" pageNumber="228">Poratophilus</emphasis>
</taxonomicName>
, therefore representing a character unique to
<taxonomicName id="31944D3588078006FDC0BCED00DCF2A9" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[556,718,919,944]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="species" species="spirobolina">
<emphasis id="C4E0EAA488078006FDC0BCED0051F2A9" box="[556,579,920,944]" italics="true" pageId="25" pageNumber="228">M</emphasis>
.
<emphasis id="C4E0EAA488078006FDA3BCED00DCF2A9" box="[591,718,920,944]" italics="true" pageId="25" pageNumber="228">spirobolina</emphasis>
</taxonomicName>
. While the two femoral spines initially suggest that
<taxonomicName id="31944D3588078006FEA4BCCD03E2F2C9" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[328,496,951,976]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="species" species="spirobolina">
<emphasis id="C4E0EAA488078006FEA4BCCD034DF2C9" box="[328,351,952,976]" italics="true" pageId="25" pageNumber="228">M</emphasis>
.
<emphasis id="C4E0EAA488078006FE9CBCCD03E2F2C9" box="[368,496,952,976]" italics="true" pageId="25" pageNumber="228">spirobolina</emphasis>
</taxonomicName>
is a member of
<taxonomicName id="31944D3588078006FD59BCCD014FF2C9" authorityName="Attems" authorityYear="1909" box="[693,861,952,976]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="C4E0EAA488078006FD59BCCD014FF2C9" box="[693,861,952,976]" italics="true" pageId="25" pageNumber="228">Harpagophora</emphasis>
</taxonomicName>
, the form of the abovementioned characters clearly separate it from that genus. The femoral spines in
<taxonomicName id="31944D3588078006FBBBBCAD0327F509" baseAuthorityName="Karsch" baseAuthorityYear="1881" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="species" species="spirobolina">
<emphasis id="C4E0EAA488078006FBBBBCAD067CF2E9" box="[1111,1134,984,1008]" italics="true" pageId="25" pageNumber="228">M</emphasis>
.
<emphasis id="C4E0EAA488078006FF59BC8D0327F509" box="[181,309,1016,1040]" italics="true" pageId="25" pageNumber="228">spirobolina</emphasis>
</taxonomicName>
are subequal in length and diameter, unlike in
<taxonomicName id="31944D3588078006FCD4BC8D01F2F509" authorityName="Attems" authorityYear="1909" box="[824,992,1016,1040]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="C4E0EAA488078006FCD4BC8D01F2F509" box="[824,992,1016,1040]" italics="true" pageId="25" pageNumber="228">Harpagophora</emphasis>
</taxonomicName>
where one of the femoral spines is always longer, larger and more robust than the other. It is here proposed—based on the evidence presented above—that
<taxonomicName id="31944D3588078006FCDDBB4D01C4F549" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[817,982,1079,1104]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="228" phylum="Arthropoda" rank="species" species="spirobolina">
<emphasis id="C4E0EAA488078006FCDDBB4D015AF549" box="[817,840,1080,1104]" italics="true" pageId="25" pageNumber="228">M</emphasis>
.
<emphasis id="C4E0EAA488078006FCBABB4D01C4F549" box="[854,982,1080,1104]" italics="true" pageId="25" pageNumber="228">spirobolina</emphasis>
</taxonomicName>
be recognised as representing a distinct genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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