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<mods:title id="CB12117EEB0F58372CCA5104E29180BD">Morphology and distribution of the Middle Asian centipede genus Krateraspis Lignau, 1929 (Chilopoda, Geophilomorpha, Mecistocephalidae)</mods:title>
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<mods:namePart id="C954439955BEB57FB842AC38A03ACFC3">Dyachkov, Yurii V.</mods:namePart>
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<mods:namePart id="2A9D9A6ED391201B01372C78244A8E13">Bonato, Lucio</mods:namePart>
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<subSection id="3A0B7AF1BE2677FE5C45769395A3F7DE" pageId="0" pageNumber="143" type="results">
<treatment id="71D76AB4F01B5492A7F007E7EFC19DF6" ID-GBIF-Taxon="194619840" LSID="urn:lsid:plazi:treatment:71D76AB4F01B5492A7F007E7EFC19DF6" httpUri="http://treatment.plazi.org/id/71D76AB4F01B5492A7F007E7EFC19DF6" lastPageNumber="143" pageId="0" pageNumber="143" scope_class="Chilopoda" scope_family="Mecistocephalidae" scope_order="Geophilomorpha">
<subSubSection id="67902C49BA2161FA3C5CCDFA28979FA9" pageId="0" pageNumber="143" type="nomenclature">
<paragraph id="A7EC15418833486156B5BA0C2497B3BB" pageId="0" pageNumber="143">
<taxonomicName id="6BF8A478B6EAD6BD4C1AEC3C809724AB" ID-CoL="6NMKN" LSID="71D76AB4-F01B-5492-A7F0-07E7EFC19DF6" authority="Titova, 1975" authorityName="Titova" authorityYear="1975" class="Chilopoda" family="Mecistocephalidae" genus="Krateraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krateraspis sselivanovi" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="sselivanovi">Krateraspis sselivanovi Titova, 1975</taxonomicName>
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<paragraph id="26D22498760093A11876822BA589D685" pageId="0" pageNumber="143">
<figureCitation id="37A1A34FFEC34369239C0FD87AAA54A9" captionStart="Figures 3335" captionStartId="F8" captionText="Figures 33 - 35. Krateraspis sselivanovi Titova, 1975: 33, 34 head, forcipular and LBS 1 (ventr., dors.) 35 LBS 1 and 2 (ventr.). Specimen: ♂ from Sharak (ZMMU Rc 8154). Abbreviations: ms - median longitudinal sulcus. Scale bars: 0.2 mm (33, 34); 0.1 mm (35)." figureDoi="10.3897/zookeys.1095.80806.figures33-35" httpUri="https://binary.pensoft.net/fig/672513" pageId="0" pageNumber="143">Figures 33-35</figureCitation>
<figureCitation id="6AB770D2701EBDD3AD834EA913685020" captionStart="Figures 3639" captionStartId="F9" captionText="Figures 36 - 39. Krateraspis sselivanovi Titova, 1975: 36 left forcipule (ventr.) 37 maxillary complex (ventr.) 38 clypeus and labrum (ventr.) 39 left mandible (lateral view). Specimens: 36, 39 ♂ from Fayzobod (ZMMU Rc 8173) 37, 38 ♂ from Sharak (ZMMU Rc 8165). Abbreviations: aa - anterior ala, ap - markedly areolate anterior part of clypeus, b - bulge, c - calyx of poison gland, cp - central part of clypeus with distinct but fainter areolation, f - femur, p - plagula, pa - posterior ala, ta - tarsungulum, ti - tibia, tl - transverse thickened line, tr - trochanteroprefemur. Scale bars: 0.1 mm (36 - 38); 0.05 mm (39)." figureDoi="10.3897/zookeys.1095.80806.figures36-39" httpUri="https://binary.pensoft.net/fig/672514" pageId="0" pageNumber="143">, 36-39</figureCitation>
<figureCitation id="4E39AD2689627AD0F847FDA6889CE796" captionStart="Figures 4043" captionStartId="F10" captionText="Figures 40 - 43. Krateraspis sselivanovi Titova, 1975: 40, 41 ultimate LBS and postpedal segments of ♂ (dors., ventr.) 42, 43 ultimate LBS and postpedal segments of ♀ (dors., ventr.). Specimens from Sharak: 40, 41 ♂ (ZMMU Rc 8154) 42, 43 ♀ (ZMMU Rc 8153). Abbreviations: cp - coxopleural pores, ups - presternite of ultimate LBS, upt - pretergite of ultimate LBS, us - metasternite of ultimate LBS, ut - tergite of ultimate LBS. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1095.80806.figures40-43" httpUri="https://binary.pensoft.net/fig/672515" pageId="0" pageNumber="143">, 40-43</figureCitation>
</paragraph>
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<subSubSection id="D5A5939CA29CB716E3BED865227DCCDD" pageId="0" pageNumber="143" type="reference_group">
<paragraph id="FDCF57F75633E9D146A9963C0BCA5763" pageId="0" pageNumber="143">
<taxonomicName id="845A89C292A8B2E4B7283B6465FE39BC" authorityName="Titova" authorityYear="1975" class="Chilopoda" family="Mecistocephalidae" genus="Krateraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krateraspis sselivanovi" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="sselivanovi">Krateraspis sselivanovi</taxonomicName>
:
<bibRefCitation id="C6016D3828E36C11DD49F3117F6F0DA1" author="Titova, LP" journalOrPublisher="[Zoologicheskii zhurnal] Zoologicheskii Jurnal" pageId="0" pageNumber="143" pagination="39 - 48" refId="B34" refString="Titova, LP, 1975. Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda). [Zoologicheskii zhurnal] Zoologicheskii Jurnal 54 (1): 39 - 48" title="Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda)." volume="54" year="1975">Titova 1975</bibRefCitation>
: 41 (original description), 45, 46 (in key); fig. 2: 1-5A.
<bibRefCitation id="C99BB6CD5893944C0C497FDC6F241CA8" DOI="https://doi.org/10.1046/j.1365-3113.2003.00217.x" author="Bonato, L" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="143" pagination="539 - 579" refId="B11" refString="Bonato, L, Foddai, D, Minelli, A, 2003. Evolutionary trends and patterns in centipede segment number based on a cladistic analysis of Mecistocephalidae (Chilopoda: Geophilomorpha). Systematic Entomology 28 (4): 539 - 579, DOI: https://doi.org/10.1046/j.1365-3113.2003.00217.x" title="Evolutionary trends and patterns in centipede segment number based on a cladistic analysis of Mecistocephalidae (Chilopoda: Geophilomorpha)." url="https://doi.org/10.1046/j.1365-3113.2003.00217.x" volume="28" year="2003">Bonato et al. 2003</bibRefCitation>
: 543, 545, 546, 550, 551, 552, 577.
<bibRefCitation id="E69F143B901E4574E59AA5195AC51442" DOI="https://doi.org/10.15298/arthsel.28.3.02" author="Dyachkov, YuV" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="143" pagination="368 - 373" refId="B12" refString="Dyachkov, YuV, 2019. New data on the family Mecistocephalidae Bollman, 1893 (Chilopoda: Geophilomorpha) from Middle Asia. Arthropoda Selecta 28 (3): 368 - 373, DOI: https://doi.org/10.15298/arthsel.28.3.02" title="New data on the family Mecistocephalidae Bollman, 1893 (Chilopoda: Geophilomorpha) from Middle Asia." url="https://doi.org/10.15298/arthsel.28.3.02" volume="28" year="2019">Dyachkov 2019</bibRefCitation>
: 368, 373 (in key).
<bibRefCitation id="614FDDF9B6B828679FDC076A786E7655" DOI="https://doi.org/10.37828/em.2020.36.6" author="Dyachkov, YuV" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="143" pagination="78 - 86" refId="B13" refString="Dyachkov, YuV, 2020. New data on the centipede (Chilopoda) fauna from Tajikistan. Ecologica Montenegrina 36: 78 - 86, DOI: https://doi.org/10.37828/em.2020.36.6" title="New data on the centipede (Chilopoda) fauna from Tajikistan." url="https://doi.org/10.37828/em.2020.36.6" volume="36" year="2020">Dyachkov 2020</bibRefCitation>
: 84.
</paragraph>
</subSubSection>
<subSubSection id="9F63294576217D69818930F6F81D99B2" pageId="0" pageNumber="143" type="type locality">
<paragraph id="53143340BFC810A0AC29CE62D2AB3D1A" pageId="0" pageNumber="143">Type locality.</paragraph>
<paragraph id="F71E1DB13AE2AB592DB642FD2676898E" pageId="0" pageNumber="143">
&quot;Tajikistan, Sharak&quot; (
<bibRefCitation id="8478308853C8F5ADD2848860D4B3ED8B" author="Titova, LP" journalOrPublisher="[Zoologicheskii zhurnal] Zoologicheskii Jurnal" pageId="0" pageNumber="143" pagination="39 - 48" refId="B34" refString="Titova, LP, 1975. Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda). [Zoologicheskii zhurnal] Zoologicheskii Jurnal 54 (1): 39 - 48" title="Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda)." volume="54" year="1975">Titova 1975</bibRefCitation>
) [Tajikistan, Khatlon region, Sharak village, ca.
<geoCoordinate id="E6D303E4E449A1106771D5DCA6A0482F" degrees="38" direction="north" minutes="16" orientation="latitude" precision="925" value="38.266666">38°16'N</geoCoordinate>
,
<geoCoordinate id="7360C25719C966504C2ACC893C6864CE" degrees="70" direction="east" minutes="04" orientation="longitude" precision="925" value="70.066666">70°04'E</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection id="C145D097128DB3AB141B4E82142CDCA9" pageId="0" pageNumber="143" type="examined specimens">
<paragraph id="6930C021582F0B60EAA2B4C0686DDF8F" pageId="0" pageNumber="143">Examined specimens.</paragraph>
<paragraph id="03C533F990D2F3E2CA858319EC3E6C15" pageId="0" pageNumber="143">
<emphasis id="B27A08F31AB6FBC0A90FAD55A1CECA9E" bold="true" italics="true" pageId="0" pageNumber="143">Paratypes</emphasis>
: 1 ♂, from [Tajikistan, Khatlon region,
<normalizedToken id="CC4D121497CF224CF882B8EE90B75F87" originalValue="Muminobod">Mu'minobod</normalizedToken>
district], Sharak, 10-20 [cm deep], 31.V.[19]65 (ZMMU Rc 8154); 2 ♂♂ and 2 ♀♀, from Sharak, 0-10 [cm deep], 29.V.[19]65 (ZMMU Rc 8167); 1 ♂, from Sharak, 10-20 [cm deep], 4.VI.[19]64 (ZMMU Rc 8175).
<emphasis id="3C8C83D69FD70AC64620E4EA1D440B63" bold="true" italics="true" pageId="0" pageNumber="143">Other material</emphasis>
: 3 ♀♀, from Sharak, 10-20, 20-30, 40-50 [cm deep], 15.X.[19]64 (ZMMU Rc 8153); 1 ♂ and 2 ♀♀, from Sharak, 0-10 [cm deep], 4.VI.[19]69 (ZMMU Rc 8163); 1 ♂, from Sharak, 20-40 [cm deep], 8.X.[19]65 (ZMMU Rc 8165); 1 ♂, from [unknown region] F-bad [Fayzobod village],
<taxonomicName id="353DFB636AF805C409788361BF7405BB" class="Liliopsida" family="Poaceae" genus="Hordeum" kingdom="Plantae" lsidName="Hordeum" order="Poales" pageId="0" pageNumber="143" phylum="Tracheophyta" rank="genus">
<emphasis id="5548037A0B42059C45F34F75EC40A199" italics="true" pageId="0" pageNumber="143">Hordeum</emphasis>
</taxonomicName>
, 70-80 [cm deep], 30.VII.[19]66 (ZMMU Rc 8173); 1 ♂, from [Districts of Republican Subordination], Garm [village, ca.
<geoCoordinate id="970A82C95A7E6DD2E7AD3CD2D0F07063" degrees="39" direction="north" minutes="1" orientation="latitude" precision="925" value="39.016666">39°1'N</geoCoordinate>
,
<geoCoordinate id="0BD07BDE4BBEC69EDFD7632A804EC064" degrees="70" direction="east" minutes="22" orientation="longitude" precision="925" value="70.36667">70°22'E</geoCoordinate>
], 21.VI.[19]69 (ZMMU Rc 8187).
</paragraph>
</subSubSection>
<subSubSection id="98EA34FEE5569B4FC466BE94607021EB" pageId="0" pageNumber="143" type="remarks">
<paragraph id="74D63332BE5C3FD9E3CAB00FE656A9D0" pageId="0" pageNumber="143">Remarks on nomenclatural issues.</paragraph>
<paragraph id="4FB4BAC807EE5EF7EDDE7DB345393E96" pageId="0" pageNumber="143">
The type series of
<taxonomicName id="0FA8D560F897EF838616C1A8CAC7A932" class="Chilopoda" kingdom="Animalia" lsidName="K. sselivanovi" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="sselivanovi">
<emphasis id="CB145626C2A3905535FE01A40CB269AD" italics="true" pageId="0" pageNumber="143">K. sselivanovi</emphasis>
</taxonomicName>
comprises 21 specimens: the holotype and 19 paratypes from Sharak, and another paratype from Faizobod (
<bibRefCitation id="AAC2B80866CBED3F63A61976D845A42C" author="Titova, LP" journalOrPublisher="[Zoologicheskii zhurnal] Zoologicheskii Jurnal" pageId="0" pageNumber="143" pagination="39 - 48" refId="B34" refString="Titova, LP, 1975. Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda). [Zoologicheskii zhurnal] Zoologicheskii Jurnal 54 (1): 39 - 48" title="Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda)." volume="54" year="1975">Titova 1975</bibRefCitation>
). These specimens are expected to be at the ZMMU, but we did not find the holotype, and the paratypes are not marked as such. Nevertheless, we detected six specimens that can be recognized as paratypes according to the locality and date reported on labels (ZMMU Rc 8154, ZMMU Rc 8167, ZMMU Rc 8175), while one specimen (ZMMU Rc 8187) can be recognized as not belonging to the type series, because it is from a locality not mentioned by
<bibRefCitation id="107B6A52C37CDBF8F8C3316139E5A8E9" author="Titova, LP" journalOrPublisher="[Zoologicheskii zhurnal] Zoologicheskii Jurnal" pageId="0" pageNumber="143" pagination="39 - 48" refId="B34" refString="Titova, LP, 1975. Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda). [Zoologicheskii zhurnal] Zoologicheskii Jurnal 54 (1): 39 - 48" title="Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda)." volume="54" year="1975">Titova (1975)</bibRefCitation>
. Instead, the date on the labels of the other eight specimens (ZMMU Rc 8153, ZMMU Rc 8163, ZMMU Rc 8165, ZMMU Rc 8173) do not fully correspond to the dates reported by
<bibRefCitation id="A23729C2FC3992AB9CD8890CE4328A50" author="Titova, LP" journalOrPublisher="[Zoologicheskii zhurnal] Zoologicheskii Jurnal" pageId="0" pageNumber="143" pagination="39 - 48" refId="B34" refString="Titova, LP, 1975. Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda). [Zoologicheskii zhurnal] Zoologicheskii Jurnal 54 (1): 39 - 48" title="Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda)." volume="54" year="1975">Titova (1975)</bibRefCitation>
, so it is uncertain whether they are paratypes or not.
</paragraph>
<caption id="0AABF2AA9548057B8446AC4CC0EAF78B" doi="10.3897/zookeys.1095.80806.figures33-35" httpUri="https://binary.pensoft.net/fig/672513" pageId="0" pageNumber="143" start="Figures 3335" startId="F8">
<paragraph id="EC8194A5D49AED3188EAB37A0DD9FA82" pageId="0" pageNumber="143">
<emphasis id="D74D9C0F6F1B128D402025FD2D10A58A" bold="true" pageId="0" pageNumber="143">Figures 33-35.</emphasis>
<taxonomicName id="179E772DFB9E6B70A077A1519B4AD65F" authorityName="Titova" authorityYear="1975" class="Chilopoda" family="Mecistocephalidae" genus="Krateraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krateraspis sselivanovi" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="sselivanovi">
<emphasis id="DC1F2471B7B3C4BB2FA1FBA132ED69F5" italics="true" pageId="0" pageNumber="143">Krateraspis sselivanovi</emphasis>
</taxonomicName>
Titova, 1975:
<emphasis id="E9ADB0B6723D4DFC482CFF4AF0A588DF" bold="true" pageId="0" pageNumber="143">33, 34</emphasis>
head, forcipular and LBS 1 (ventr., dors.)
<emphasis id="79DF6C4AC0F36A6DC9561CA1EB85235C" bold="true" pageId="0" pageNumber="143">35</emphasis>
LBS 1 and 2 (ventr.). Specimen: ♂ from Sharak (ZMMU Rc 8154). Abbreviations: ms - median longitudinal sulcus. Scale bars: 0.2 mm (
<emphasis id="CC50084002FC094A39D52C3A7E6B270D" bold="true" pageId="0" pageNumber="143">33, 34</emphasis>
); 0.1 mm (
<emphasis id="E38A668F83C3A32C976C0681D1451038" bold="true" pageId="0" pageNumber="143">35</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="2FCAF37D0678694903FE20083B26E589" pageId="0" pageNumber="143" type="diagnosis">
<paragraph id="724B9299AC24EEC61BF56B8F0DC502B3" pageId="0" pageNumber="143">Diagnosis.</paragraph>
<paragraph id="3CFA020881F4A326F25309972F08613C" pageId="0" pageNumber="143">
A species of
<taxonomicName id="478F7F6129B9A709E597ECBBE9FFE740" authorityName="Lignau" authorityYear="1929" class="Chilopoda" family="Mecistocephalidae" genus="Krateraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krateraspis" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="4172049D64329093DB0F6B082BBFFC64" italics="true" pageId="0" pageNumber="143">Krateraspis</emphasis>
</taxonomicName>
with: clypeus showing the transition between marked and weak areolation at ca. 0.3-0.4 of the clypeal medial length, so that some clypeal setae are surrounded by marked areolation while other setae are surrounded by weak areolation; no spine-like sensilla on the lateral parts of the clypeus; second maxillary telopodites not distinctly surpassing the tips of the telopodites of the first maxillae; first article of the second maxillary telopodites with a distinct distal bulge on the external side; all forcipular articles with a distinct denticle with the exception of the femur; invariably 53 pairs of legs. See also Table
<tableCitation id="E4B06A1F4ED54966991CB4A9A591A924" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Main differences between Krateraspis meinerti (Sseliwanoff, 1881) and K. sselivanovi Titova, 1975." httpUri="http://table.plazi.org/id/12ADF9F3295E0FB626A48BEC10026781" pageId="0" pageNumber="143" tableUuid="12ADF9F3295E0FB626A48BEC10026781">2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D9C53A7046BC6FD8E17D5852A4FAEEFC" pageId="0" pageNumber="143" type="intraspecific variation">
<paragraph id="3FC1D1A00B1AA06F738B1136D42CC38A" pageId="0" pageNumber="143">Intraspecific variation.</paragraph>
<paragraph id="6DF574A357D3EC16897984261773692B" pageId="0" pageNumber="143">
Maximum body length: 62 mm in ♀♀ (
<emphasis id="5A2C433DA04BE6AE600857D262E8783A" italics="true" pageId="0" pageNumber="143">n</emphasis>
= 7) and 67 mm in ♂♂ (
<emphasis id="53753646116CA5864ED466AC3E9D0971" italics="true" pageId="0" pageNumber="143">n</emphasis>
= 8) but the largest specimens of both sexes are slightly macerated and stretched. Color (in ethanol 70%) usually yellow, with head, forcipular segment (except forcipular tergite), and antennae light brown (Figs
<figureCitation id="234D30083F7E73BF6C9EE14D32C03C86" captionStart="Figures 3335" captionStartId="F8" captionText="Figures 33 - 35. Krateraspis sselivanovi Titova, 1975: 33, 34 head, forcipular and LBS 1 (ventr., dors.) 35 LBS 1 and 2 (ventr.). Specimen: ♂ from Sharak (ZMMU Rc 8154). Abbreviations: ms - median longitudinal sulcus. Scale bars: 0.2 mm (33, 34); 0.1 mm (35)." figureDoi="10.3897/zookeys.1095.80806.figures33-35" httpUri="https://binary.pensoft.net/fig/672513" pageId="0" pageNumber="143">33</figureCitation>
,
<figureCitation id="84A2B1887CD9F5E4EC93F15E9F4CE676" captionStart="Figures 3335" captionStartId="F8" captionText="Figures 33 - 35. Krateraspis sselivanovi Titova, 1975: 33, 34 head, forcipular and LBS 1 (ventr., dors.) 35 LBS 1 and 2 (ventr.). Specimen: ♂ from Sharak (ZMMU Rc 8154). Abbreviations: ms - median longitudinal sulcus. Scale bars: 0.2 mm (33, 34); 0.1 mm (35)." figureDoi="10.3897/zookeys.1095.80806.figures33-35" httpUri="https://binary.pensoft.net/fig/672513" pageId="0" pageNumber="143">34</figureCitation>
).
</paragraph>
<paragraph id="95697405ADB59C302B0C9D5E8E3D7D1F" pageId="0" pageNumber="143">
<emphasis id="04BA1A0FECA8F14F1140A5DC426FA800" bold="true" pageId="0" pageNumber="143">
<emphasis id="44BD67F07AA2898A10DCD4ECA092384C" italics="true" pageId="0" pageNumber="143">Head</emphasis>
.
</emphasis>
Anterior markedly areolate part of the clypeus extending medially for 30-40% of the total length of the clypeus (Fig.
<figureCitation id="F4CEF757445F14BB798CC0F312D1ACB6" captionStart="Figures 3639" captionStartId="F9" captionText="Figures 36 - 39. Krateraspis sselivanovi Titova, 1975: 36 left forcipule (ventr.) 37 maxillary complex (ventr.) 38 clypeus and labrum (ventr.) 39 left mandible (lateral view). Specimens: 36, 39 ♂ from Fayzobod (ZMMU Rc 8173) 37, 38 ♂ from Sharak (ZMMU Rc 8165). Abbreviations: aa - anterior ala, ap - markedly areolate anterior part of clypeus, b - bulge, c - calyx of poison gland, cp - central part of clypeus with distinct but fainter areolation, f - femur, p - plagula, pa - posterior ala, ta - tarsungulum, ti - tibia, tl - transverse thickened line, tr - trochanteroprefemur. Scale bars: 0.1 mm (36 - 38); 0.05 mm (39)." figureDoi="10.3897/zookeys.1095.80806.figures36-39" httpUri="https://binary.pensoft.net/fig/672514" pageId="0" pageNumber="143">38</figureCitation>
). Invariably eight clypeal setae: 2-4 setae on the markedly areolate part, 2-4 setae located on the border between the markedly areolate part and the weakly areolate part, and two setae on the weakly areolate one; spine-like sensilla on the clypeal lateral parts always absent. Each mandible (Fig.
<figureCitation id="0DA0267CBEAC5B7153D3541B3334CF80" captionStart="Figures 3639" captionStartId="F9" captionText="Figures 36 - 39. Krateraspis sselivanovi Titova, 1975: 36 left forcipule (ventr.) 37 maxillary complex (ventr.) 38 clypeus and labrum (ventr.) 39 left mandible (lateral view). Specimens: 36, 39 ♂ from Fayzobod (ZMMU Rc 8173) 37, 38 ♂ from Sharak (ZMMU Rc 8165). Abbreviations: aa - anterior ala, ap - markedly areolate anterior part of clypeus, b - bulge, c - calyx of poison gland, cp - central part of clypeus with distinct but fainter areolation, f - femur, p - plagula, pa - posterior ala, ta - tarsungulum, ti - tibia, tl - transverse thickened line, tr - trochanteroprefemur. Scale bars: 0.1 mm (36 - 38); 0.05 mm (39)." figureDoi="10.3897/zookeys.1095.80806.figures36-39" httpUri="https://binary.pensoft.net/fig/672514" pageId="0" pageNumber="143">39</figureCitation>
) usually with six lamellae, with 5-7 teeth in each lamella. Second maxillae (Fig.
<figureCitation id="1653171DF9FCBE103DCD81F25BC06FD0" captionStart="Figures 3639" captionStartId="F9" captionText="Figures 36 - 39. Krateraspis sselivanovi Titova, 1975: 36 left forcipule (ventr.) 37 maxillary complex (ventr.) 38 clypeus and labrum (ventr.) 39 left mandible (lateral view). Specimens: 36, 39 ♂ from Fayzobod (ZMMU Rc 8173) 37, 38 ♂ from Sharak (ZMMU Rc 8165). Abbreviations: aa - anterior ala, ap - markedly areolate anterior part of clypeus, b - bulge, c - calyx of poison gland, cp - central part of clypeus with distinct but fainter areolation, f - femur, p - plagula, pa - posterior ala, ta - tarsungulum, ti - tibia, tl - transverse thickened line, tr - trochanteroprefemur. Scale bars: 0.1 mm (36 - 38); 0.05 mm (39)." figureDoi="10.3897/zookeys.1095.80806.figures36-39" httpUri="https://binary.pensoft.net/fig/672514" pageId="0" pageNumber="143">37</figureCitation>
): 1st article invariably with a distinct distal bulge on the external side; distal part of 2nd article usually with two or three setae, distal part of 3rd article with numerous setae.
</paragraph>
<caption id="483E28C7A61D1EE1AF47273B8DAFEFA4" doi="10.3897/zookeys.1095.80806.figures36-39" httpUri="https://binary.pensoft.net/fig/672514" pageId="0" pageNumber="143" start="Figures 3639" startId="F9">
<paragraph id="CEB8EE24951E22F7FFB5752336E0B155" pageId="0" pageNumber="143">
<emphasis id="5C42C873C78E89094259C4E5DD13BCF5" bold="true" pageId="0" pageNumber="143">Figures 36-39.</emphasis>
<taxonomicName id="1975DFDA5F7831022B163A4146FF7FD6" authorityName="Titova" authorityYear="1975" class="Chilopoda" family="Mecistocephalidae" genus="Krateraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krateraspis sselivanovi" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="sselivanovi">
<emphasis id="CBBA4AD737E29AD4624B791DADD01619" italics="true" pageId="0" pageNumber="143">Krateraspis sselivanovi</emphasis>
</taxonomicName>
Titova, 1975:
<emphasis id="7060DC6B12905CD2CC1D227A673E6042" bold="true" pageId="0" pageNumber="143">36</emphasis>
left forcipule (ventr.)
<emphasis id="453C64A3C228F4BC5F83DADB31047C26" bold="true" pageId="0" pageNumber="143">37</emphasis>
maxillary complex (ventr.)
<emphasis id="17575D9914FC2CE7CD9E1988DC308C1D" bold="true" pageId="0" pageNumber="143">38</emphasis>
clypeus and labrum (ventr.)
<emphasis id="02724EEE78A3108EBEB832D118BBA97A" bold="true" pageId="0" pageNumber="143">39</emphasis>
left mandible (lateral view). Specimens:
<emphasis id="69EADB7E21BE6A73441C05E55271DD07" bold="true" pageId="0" pageNumber="143">36, 39</emphasis>
♂ from Fayzobod (ZMMU Rc 8173)
<emphasis id="99DBF15EAAA4E88FE1318BE6D76E7A20" bold="true" pageId="0" pageNumber="143">37, 38</emphasis>
♂ from Sharak (ZMMU Rc 8165). Abbreviations: aa - anterior ala, ap - markedly areolate anterior part of clypeus, b - bulge, c - calyx of poison gland, cp - central part of clypeus with distinct but fainter areolation, f - femur, p - plagula, pa - posterior ala, ta - tarsungulum, ti - tibia, tl - transverse thickened line, tr - trochanteroprefemur. Scale bars: 0.1 mm (
<emphasis id="72167634823F18993A22CD7CBDF8FDA5" bold="true" pageId="0" pageNumber="143">36-38</emphasis>
); 0.05 mm (
<emphasis id="38BB342AA045A9E26781D422C9A9E357" bold="true" pageId="0" pageNumber="143">39</emphasis>
).
</paragraph>
</caption>
<paragraph id="EF665FE3936FAF51BF6EC4CFE2FDD695" pageId="0" pageNumber="143">
<emphasis id="2B23EC28456B72839E1FB1C72FCB9D83" bold="true" pageId="0" pageNumber="143">
<emphasis id="7CB60447BF93C74058E9D9A77E37201A" italics="true" pageId="0" pageNumber="143">Forcipular segment</emphasis>
.
</emphasis>
Tergite usually partially covered by the tergite 1. Forcipules, when closed, usually reaching the anterior margin of the cephalic plate (Figs
<figureCitation id="EBDB723ED2C5B4DD101EFA017F1EF6A8" captionStart="Figures 3335" captionStartId="F8" captionText="Figures 33 - 35. Krateraspis sselivanovi Titova, 1975: 33, 34 head, forcipular and LBS 1 (ventr., dors.) 35 LBS 1 and 2 (ventr.). Specimen: ♂ from Sharak (ZMMU Rc 8154). Abbreviations: ms - median longitudinal sulcus. Scale bars: 0.2 mm (33, 34); 0.1 mm (35)." figureDoi="10.3897/zookeys.1095.80806.figures33-35" httpUri="https://binary.pensoft.net/fig/672513" pageId="0" pageNumber="143">33</figureCitation>
,
<figureCitation id="E5876A46C3C3B93C855E14104014FDB2" captionStart="Figures 3335" captionStartId="F8" captionText="Figures 33 - 35. Krateraspis sselivanovi Titova, 1975: 33, 34 head, forcipular and LBS 1 (ventr., dors.) 35 LBS 1 and 2 (ventr.). Specimen: ♂ from Sharak (ZMMU Rc 8154). Abbreviations: ms - median longitudinal sulcus. Scale bars: 0.2 mm (33, 34); 0.1 mm (35)." figureDoi="10.3897/zookeys.1095.80806.figures33-35" httpUri="https://binary.pensoft.net/fig/672513" pageId="0" pageNumber="143">34</figureCitation>
). Trochanteroprefemur, tibia and tarsungulum with denticles, while femur without denticle (Figs
<figureCitation id="42613151050CDF7975348264706B8755" captionStart="Figures 3335" captionStartId="F8" captionText="Figures 33 - 35. Krateraspis sselivanovi Titova, 1975: 33, 34 head, forcipular and LBS 1 (ventr., dors.) 35 LBS 1 and 2 (ventr.). Specimen: ♂ from Sharak (ZMMU Rc 8154). Abbreviations: ms - median longitudinal sulcus. Scale bars: 0.2 mm (33, 34); 0.1 mm (35)." figureDoi="10.3897/zookeys.1095.80806.figures33-35" httpUri="https://binary.pensoft.net/fig/672513" pageId="0" pageNumber="143">33</figureCitation>
,
<figureCitation id="B7E55880961E6275341486913FE2E48A" captionStart="Figures 3639" captionStartId="F9" captionText="Figures 36 - 39. Krateraspis sselivanovi Titova, 1975: 36 left forcipule (ventr.) 37 maxillary complex (ventr.) 38 clypeus and labrum (ventr.) 39 left mandible (lateral view). Specimens: 36, 39 ♂ from Fayzobod (ZMMU Rc 8173) 37, 38 ♂ from Sharak (ZMMU Rc 8165). Abbreviations: aa - anterior ala, ap - markedly areolate anterior part of clypeus, b - bulge, c - calyx of poison gland, cp - central part of clypeus with distinct but fainter areolation, f - femur, p - plagula, pa - posterior ala, ta - tarsungulum, ti - tibia, tl - transverse thickened line, tr - trochanteroprefemur. Scale bars: 0.1 mm (36 - 38); 0.05 mm (39)." figureDoi="10.3897/zookeys.1095.80806.figures36-39" httpUri="https://binary.pensoft.net/fig/672514" pageId="0" pageNumber="143">36</figureCitation>
), with the single exception of a specimen with a denticle on the right femur (however collected together with specimens with usual morphology in the sample ZMMU Rc 8163). Worth noting is that an analogous case of asymmetry has been detected in a specimen of
<taxonomicName id="5FE205E581B67E394A3706B2CBEC4DDD" class="Chilopoda" kingdom="Animalia" lsidName="K. meinerti" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="meinerti">
<emphasis id="1BF9D3BCD76F99D8781F862A10F72AEB" italics="true" pageId="0" pageNumber="143">K. meinerti</emphasis>
</taxonomicName>
, where a denticle has been recognized on one femur but not in the other femur (see above). The distal denticle of trochanteroprefemur usually larger than both denticles on the tibia and tarsungulum (Fig.
<figureCitation id="A64F9EB25E93C1CF7A025AD91AAC57D9" captionStart="Figures 3639" captionStartId="F9" captionText="Figures 36 - 39. Krateraspis sselivanovi Titova, 1975: 36 left forcipule (ventr.) 37 maxillary complex (ventr.) 38 clypeus and labrum (ventr.) 39 left mandible (lateral view). Specimens: 36, 39 ♂ from Fayzobod (ZMMU Rc 8173) 37, 38 ♂ from Sharak (ZMMU Rc 8165). Abbreviations: aa - anterior ala, ap - markedly areolate anterior part of clypeus, b - bulge, c - calyx of poison gland, cp - central part of clypeus with distinct but fainter areolation, f - femur, p - plagula, pa - posterior ala, ta - tarsungulum, ti - tibia, tl - transverse thickened line, tr - trochanteroprefemur. Scale bars: 0.1 mm (36 - 38); 0.05 mm (39)." figureDoi="10.3897/zookeys.1095.80806.figures36-39" httpUri="https://binary.pensoft.net/fig/672514" pageId="0" pageNumber="143">36</figureCitation>
). Calyx of poison gland usually reaching the trochanterophefemur in both sexes.
</paragraph>
<caption id="0F349F9BE0B7FAD2DF79765C2457A414" doi="10.3897/zookeys.1095.80806.figures40-43" httpUri="https://binary.pensoft.net/fig/672515" pageId="0" pageNumber="143" start="Figures 4043" startId="F10">
<paragraph id="47222B3E80F34B87BF2029CDA827AA6E" pageId="0" pageNumber="143">
<emphasis id="F624943A2270B56A39F76B0BB23FE406" bold="true" pageId="0" pageNumber="143">Figures 40-43.</emphasis>
<taxonomicName id="A447E3C078985D9D16F5F151B744B4BC" authorityName="Titova" authorityYear="1975" class="Chilopoda" family="Mecistocephalidae" genus="Krateraspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Krateraspis sselivanovi" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="sselivanovi">
<emphasis id="8B26FC1E7908781C76CB8754217BE148" italics="true" pageId="0" pageNumber="143">Krateraspis sselivanovi</emphasis>
</taxonomicName>
Titova, 1975:
<emphasis id="B8CC005C9CBDD53E969D258517944C0A" bold="true" pageId="0" pageNumber="143">40, 41</emphasis>
ultimate LBS and postpedal segments of ♂ (dors., ventr.)
<emphasis id="10DF2E806F05A130001FF5DB9B015B3A" bold="true" pageId="0" pageNumber="143">42, 43</emphasis>
ultimate LBS and postpedal segments of ♀ (dors., ventr.). Specimens from Sharak:
<emphasis id="9B7F7579CA0AEDBDA1EAD2C81EA6C5AA" bold="true" pageId="0" pageNumber="143">40, 41</emphasis>
♂ (ZMMU Rc 8154)
<emphasis id="FD80FAFBBB8CD04188302BE89C33CC2B" bold="true" pageId="0" pageNumber="143">42, 43</emphasis>
♀ (ZMMU Rc 8153). Abbreviations: cp - coxopleural pores, ups - presternite of ultimate LBS, upt - pretergite of ultimate LBS, us - metasternite of ultimate LBS, ut - tergite of ultimate LBS. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="DB90D49BB63E329C5E566888BC3FEBD4" pageId="0" pageNumber="143">
<emphasis id="CC3F25EBB2A11D349371838A2FCBA19D" bold="true" pageId="0" pageNumber="143">
<emphasis id="B94958015F977D44C197B528B37D5694" italics="true" pageId="0" pageNumber="143">Leg-bearing segments</emphasis>
.
</emphasis>
Invariably 53 pairs of legs in all examined specimens. Worth noting is that
<taxonomicName id="B2BC8151C742A7421A285C3D590B2A1B" class="Chilopoda" kingdom="Animalia" lsidName="K. meinerti" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="meinerti">
<emphasis id="DB9A895CFBA58CBF811F06A89C646C29" italics="true" pageId="0" pageNumber="143">K. meinerti</emphasis>
</taxonomicName>
has invariably 45 pairs of legs and the difference of eight pairs between the two species corresponds to a putative evolutionary change that have repeatedly occurred in the
<taxonomicName id="AF4DEC373E123B9AC8E0A185C25E5604" class="Chilopoda" family="Mecistocephalidae" kingdom="Animalia" lsidName="" order="Geophilomorpha" pageId="0" pageNumber="143" phylum="Arthropoda" rank="family">Mecistocephalidae</taxonomicName>
(
<bibRefCitation id="6BE7B64C52F4789D7BFA7C2D5312C6F5" DOI="https://doi.org/10.1046/j.1365-3113.2003.00217.x" author="Bonato, L" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="143" pagination="539 - 579" refId="B11" refString="Bonato, L, Foddai, D, Minelli, A, 2003. Evolutionary trends and patterns in centipede segment number based on a cladistic analysis of Mecistocephalidae (Chilopoda: Geophilomorpha). Systematic Entomology 28 (4): 539 - 579, DOI: https://doi.org/10.1046/j.1365-3113.2003.00217.x" title="Evolutionary trends and patterns in centipede segment number based on a cladistic analysis of Mecistocephalidae (Chilopoda: Geophilomorpha)." url="https://doi.org/10.1046/j.1365-3113.2003.00217.x" volume="28" year="2003">Bonato et al. 2003</bibRefCitation>
).
</paragraph>
<paragraph id="DEAC7EC9FC8737190EF0C75F9DA05DB0" pageId="0" pageNumber="143">
<emphasis id="950DCCCC84197D39261588C30C7C0739" bold="true" pageId="0" pageNumber="143">
<emphasis id="7B3807EBD2AF126FD61E157D4E896D0D" italics="true" pageId="0" pageNumber="143">Ultimate leg-bearing segment</emphasis>
.
</emphasis>
Almost similar in both sexes (Figs
<figureCitation id="D0C7EE0EE9C4E133E28C7438EC813EC8" captionStart="Figures 4043" captionStartId="F10" captionText="Figures 40 - 43. Krateraspis sselivanovi Titova, 1975: 40, 41 ultimate LBS and postpedal segments of ♂ (dors., ventr.) 42, 43 ultimate LBS and postpedal segments of ♀ (dors., ventr.). Specimens from Sharak: 40, 41 ♂ (ZMMU Rc 8154) 42, 43 ♀ (ZMMU Rc 8153). Abbreviations: cp - coxopleural pores, ups - presternite of ultimate LBS, upt - pretergite of ultimate LBS, us - metasternite of ultimate LBS, ut - tergite of ultimate LBS. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1095.80806.figures40-43" httpUri="https://binary.pensoft.net/fig/672515" pageId="0" pageNumber="143">40-43</figureCitation>
): metasternite subtriangular, its length to width ratio varying between 0.9 and 1.0, and the anterior margin 4-5
<normalizedToken id="E537DA0DF32758C43651A3B3F7397102" originalValue="×">x</normalizedToken>
wider than the posterior one; up to 20 pores on each coxopleuron in ♂♂, and up to 50 pores in ♀♀; legs slender and densely setose, without pretarsus.
</paragraph>
<paragraph id="F59EF10A22987E9D9A523A17B11DB19A" pageId="0" pageNumber="143">
<emphasis id="44C22B7F8828AA2A8F22C5E0341B47DA" bold="true" pageId="0" pageNumber="143">
<emphasis id="8CCC3B10832BCC2BB8B498C1AA248D60" italics="true" pageId="0" pageNumber="143">Postpedal segments</emphasis>
.
</emphasis>
Densely setose in both sexes (Figs
<figureCitation id="33122275EE8B079EA6AC6439AEB5A783" captionStart="Figures 4043" captionStartId="F10" captionText="Figures 40 - 43. Krateraspis sselivanovi Titova, 1975: 40, 41 ultimate LBS and postpedal segments of ♂ (dors., ventr.) 42, 43 ultimate LBS and postpedal segments of ♀ (dors., ventr.). Specimens from Sharak: 40, 41 ♂ (ZMMU Rc 8154) 42, 43 ♀ (ZMMU Rc 8153). Abbreviations: cp - coxopleural pores, ups - presternite of ultimate LBS, upt - pretergite of ultimate LBS, us - metasternite of ultimate LBS, ut - tergite of ultimate LBS. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1095.80806.figures40-43" httpUri="https://binary.pensoft.net/fig/672515" pageId="0" pageNumber="143">40-43</figureCitation>
). Male gonopods bi-articulate, narrow, and separated by a conic projection in between (Fig.
<figureCitation id="F39B721232308A795CEF04FE786FD1DB" captionStart="Figures 4043" captionStartId="F10" captionText="Figures 40 - 43. Krateraspis sselivanovi Titova, 1975: 40, 41 ultimate LBS and postpedal segments of ♂ (dors., ventr.) 42, 43 ultimate LBS and postpedal segments of ♀ (dors., ventr.). Specimens from Sharak: 40, 41 ♂ (ZMMU Rc 8154) 42, 43 ♀ (ZMMU Rc 8153). Abbreviations: cp - coxopleural pores, ups - presternite of ultimate LBS, upt - pretergite of ultimate LBS, us - metasternite of ultimate LBS, ut - tergite of ultimate LBS. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1095.80806.figures40-43" httpUri="https://binary.pensoft.net/fig/672515" pageId="0" pageNumber="143">41</figureCitation>
). Female gonopods bi-articulate, subtriangular, and touching each other at their bases (Fig.
<figureCitation id="7D7E5A0D312665A1578A0AEBCB7F2333" captionStart="Figures 4043" captionStartId="F10" captionText="Figures 40 - 43. Krateraspis sselivanovi Titova, 1975: 40, 41 ultimate LBS and postpedal segments of ♂ (dors., ventr.) 42, 43 ultimate LBS and postpedal segments of ♀ (dors., ventr.). Specimens from Sharak: 40, 41 ♂ (ZMMU Rc 8154) 42, 43 ♀ (ZMMU Rc 8153). Abbreviations: cp - coxopleural pores, ups - presternite of ultimate LBS, upt - pretergite of ultimate LBS, us - metasternite of ultimate LBS, ut - tergite of ultimate LBS. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1095.80806.figures40-43" httpUri="https://binary.pensoft.net/fig/672515" pageId="0" pageNumber="143">43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E046F50BAF625C1AFC7B9E4B09074044" pageId="0" pageNumber="143" type="distribution">
<paragraph id="DC670F0C52B5C72772FE8B15EDB0F41A" pageId="0" pageNumber="143">Distribution.</paragraph>
<paragraph id="ADFBE81C89B9E35E7135FAE7ECCD004E" pageId="0" pageNumber="143">
Recorded from three localities in the western offshoots of Pamir Mts (Fig.
<figureCitation id="D63CF6BE72BA98FC1248EED598C1D96D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Krateraspis Lignau, 1929: square, K. meinerti (Sseliwanoff, 1881); star, K. sselivanovi Titova, 1975. Green symbols indicate type localities. Some very close localities are marked by a single symbol. The question marks indicate alternative positions of the uncertain locality &quot; Fayzabad &quot; (see Materials and methods)." figureDoi="10.3897/zookeys.1095.80806.figure1" httpUri="https://binary.pensoft.net/fig/672506" pageId="0" pageNumber="143">1</figureCitation>
), all in Tajikistan (Khatlon region and Districts of Republican Subordination) (
<bibRefCitation id="BD7B7A5453EC17F1304FDA60348A6E37" author="Titova, LP" journalOrPublisher="[Zoologicheskii zhurnal] Zoologicheskii Jurnal" pageId="0" pageNumber="143" pagination="39 - 48" refId="B34" refString="Titova, LP, 1975. Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda). [Zoologicheskii zhurnal] Zoologicheskii Jurnal 54 (1): 39 - 48" title="Geophilids of the family Mecistocephalidae in the USSR fauna (Chilopoda)." volume="54" year="1975">Titova 1975</bibRefCitation>
; present records).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>