367 lines
68 KiB
XML
367 lines
68 KiB
XML
<document id="FBD1A385F9D85CAEADA79B7986B1E33F" ID="10.11646/zootaxa.4236.1.2SLASH10251" ID-CLB-Dataset="36479" ID-DOI="10.11646/zootaxa.4236.1.2" ID-GBIF-Dataset="0f39b5c0-0af5-476f-8448-a02339cc1dfd" ID-ISSN="1175-5326" ID-Zenodo-Dep="322014" ID-ZooBank="23412386-CB17-49CA-9C47-BD71DD9C5372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487752177037" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N. & Yushkova, Olga V." docDate="2017" docId="4F3A5135FFB7DC114FE3FD3C6D13FE4F" docLanguage="en" docName="zootaxa.4236.1.2SLASH10251.pdf" docOrigin="Zootaxa 4236 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Morphna indica Anisyutkin, sp.nov." docType="treatment" docVersion="7" lastPageNumber="59" masterDocId="B303294DFFB8DC034F74FFCC6F0CFF9F" masterDocTitle="New data on cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from India and Sri Lanka, with descriptions of new species and the genital complex of Aptera fusca (Thunberg, 1784)" masterLastPageNumber="64" masterPageNumber="41" pageNumber="56" updateTime="1698437163562" updateUser="ExternalLinkService">
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<mods:mods id="03288E9A8911A62C1D088692439F002D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B2E34700D83A657370F107054DC2CC74">
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<mods:title id="1AAB3C7926AB0F6CCB0502C381DC1FFB">New data on cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from India and Sri Lanka, with descriptions of new species and the genital complex of Aptera fusca (Thunberg, 1784)</mods:title>
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</mods:titleInfo>
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<mods:name id="425FC7781B9AE5D88279662EEACA80F9" type="personal">
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<mods:roleTerm id="B7EA41A10E66452746019F75164939BE">Author</mods:roleTerm>
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<mods:namePart id="DC54F3008823A382B171157DC85AC178">Anisyutkin, Leonid N.</mods:namePart>
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</mods:name>
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<mods:name id="4F44FA46CC59ED5D2BBE56BD38A0F31C" type="personal">
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<mods:role id="41E0C26A992FF23191C4CF26B7DE30C4">
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<mods:roleTerm id="0A62D8483C7558704FD0E50088241B54">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D1069BD8D7E15720F0F8C79400FFDFE5">Yushkova, Olga V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="AC7C9F52CFB12F642900E7FD12151C1E">text</mods:typeOfResource>
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<mods:titleInfo id="E1278041753E98319FBE4ECE2F77AC53">
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<mods:title id="F97B8D81FB9F7D8A618E43B65889866F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="986FD44375750F842669088126D516A5">
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<mods:date id="C3468177D140718DEFEB7C5052EAB2BA">2017</mods:date>
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<mods:detail id="E0F617A5246E6B4AABD4B995FBE1E26A" type="volume">
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<mods:number id="6E21C1C527189D3E626E71C4AE5EC579">4236</mods:number>
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</mods:detail>
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<mods:detail id="C91B79559D31E31B81151D7DADFA8B84" type="issue">
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<mods:number id="AE1DB8F50DB5C72EEA17C52327BE7C89">1</mods:number>
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<mods:classification id="0AA68C21D9E8D2785884031241281A56">journal article</mods:classification>
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<mods:identifier id="8B35DEC0EA0D7EFF05D4662E51A13144" type="CLB-Dataset">36479</mods:identifier>
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<mods:identifier id="C4F4CAD818BE7C33C5BB4E1A5B348006" type="DOI">10.11646/zootaxa.4236.1.2</mods:identifier>
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<mods:identifier id="C0CF8E4BB36D2428C84FC7653DB7755B" type="GBIF-Dataset">0f39b5c0-0af5-476f-8448-a02339cc1dfd</mods:identifier>
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<mods:identifier id="F08893CF06A67F9884F777D0EF996F8D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="7AC35CBA0925B7E7EF80EAEF1FCB3F7D" type="Zenodo-Dep">322014</mods:identifier>
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<mods:identifier id="EB0932F111B778762D994913F6CD3AE9" type="ZooBank">23412386-CB17-49CA-9C47-BD71DD9C5372</mods:identifier>
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</mods:mods>
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<treatment id="4F3A5135FFB7DC114FE3FD3C6D13FE4F" ID-DOI="http://doi.org/10.5281/zenodo.6015715" ID-GBIF-Taxon="127697114" ID-Zenodo-Dep="6015715" LSID="urn:lsid:plazi:treatment:4F3A5135FFB7DC114FE3FD3C6D13FE4F" httpUri="http://treatment.plazi.org/id/4F3A5135FFB7DC114FE3FD3C6D13FE4F" lastPageId="18" lastPageNumber="59" pageId="15" pageNumber="56">
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<subSubSection id="8F89B3A8FFB7DC0C4FE3FD3C6E74FCB4" pageId="15" pageNumber="56" type="nomenclature">
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<paragraph id="C72CE023FFB7DC0C4FE3FD3C6D5DFC95" blockId="15.[151,593,752,811]" box="[151,593,752,778]" pageId="15" pageNumber="56">
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<heading id="9C64574FFFB7DC0C4FE3FD3C6D5DFC95" bold="true" box="[151,593,752,778]" fontSize="11" level="1" pageId="15" pageNumber="56" reason="1">
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<emphasis id="F5E73C31FFB7DC0C4FE3FD3C6D5DFC95" bold="true" box="[151,593,752,778]" pageId="15" pageNumber="56">
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<taxonomicName id="00939BA0FFB7DC0C4FE3FD3C6EE2FC95" authority="Anisyutkin" authorityName="Anisyutkin" box="[151,494,752,778]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
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<emphasis id="F5E73C31FFB7DC0C4FE3FD3C6E57FC95" bold="true" box="[151,347,752,778]" italics="true" pageId="15" pageNumber="56">Morphna indica</emphasis>
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Anisyutkin
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</taxonomicName>
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,
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<taxonomicNameLabel id="EED4814AFFB7DC0C4E8FFD3C6D5DFC95" box="[507,593,752,778]" pageId="15" pageNumber="56" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C72CE023FFB7DC0C4FE3FCDE6E74FCB4" blockId="15.[151,593,752,811]" box="[151,376,786,811]" pageId="15" pageNumber="56">
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(
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<figureCitation id="5FA8FCA6FFB7DC0C4FEBFCDE6FE6FCB4" box="[159,234,786,811]" captionStart="FIGURE 8" captionStartId="16.[151,249,1593,1615]" captionTargetBox="[153,1429,219,1567]" captionTargetId="figure@16.[149,1434,211,1572]" captionTargetPageId="16" captionText="FIGURE 8. Morphna ssp. A, B, E, F—facial part of head; C, D, G, H—anterior part of body from above. A – D— M. indica sp. nov., male, holotype (A, C) and female, paratype (B, D); E – H— M. srilankensis sp. nov., male, holotype (E, G) and female, paratype (F, H). Pronotum on fig. G damaged. Out of scale." httpUri="https://zenodo.org/record/322022/files/figure.png" pageId="15" pageNumber="56">Figs. 8</figureCitation>
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A–D, 9C–P)
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</paragraph>
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</subSubSection>
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<subSubSection id="8F89B3A8FFB7DC0C4FE3FC986E6FFC29" pageId="15" pageNumber="56" type="materials_examined">
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<paragraph id="C72CE023FFB7DC0C4FE3FC986E6FFC29" blockId="15.[151,1437,852,2030]" pageId="15" pageNumber="56">
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<materialsCitation id="77FBEA7EFFB7DC0C4FE3FC986BA3FC0D" ID-GBIF-Occurrence="1434656654" collectingDate="1972-11-09" collectionCode="SOUTH, MHNG" collectorName="C. Besuchet & I. Lobl" country="India" elevation="950" location="Cardamon Hills" pageId="15" pageNumber="56" specimenCount="1" stateProvince="Kerala">
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<emphasis id="F5E73C31FFB7DC0C4FE3FC986FF3FCF2" bold="true" box="[151,255,852,877]" pageId="15" pageNumber="56">Material</emphasis>
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. Holotype—male,
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<collectionCode id="A18278E6FFB7DC0C4EA2FC9A6D3FFCF1" box="[470,563,854,878]" pageId="15" pageNumber="56">SOUTH</collectionCode>
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<collectingCountry id="BF84A0B3FFB7DC0C4D4EFC9A6D80FCF1" box="[570,652,854,878]" name="India" pageId="15" pageNumber="56">INDIA</collectingCountry>
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, state
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<collectingRegion id="05572EC1FFB7DC0C4DA6FC996C13FCF1" box="[722,799,853,878]" country="India" name="Kerala" pageId="15" pageNumber="56">Kerala</collectingRegion>
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,
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<location id="C24CB6F8FFB7DC0C4C5FFC996CD3FCF1" LSID="urn:lsid:plazi:treatment:4F3A5135FFB7DC114FE3FD3C6D13FE4F:C24CB6F8FFB7DC0C4C5FFC996CD3FCF1" box="[811,991,853,878]" country="India" name="Cardamon Hills" pageId="15" pageNumber="56" stateProvince="Kerala">Cardamon Hills</location>
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,
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<locationDeviation id="D69EF901FFB7DC0C4C9FFC996A6BFCF1" box="[1003,1383,853,878]" pageId="15" pageNumber="56">between Pambanar and Peermade</locationDeviation>
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,
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<elevation id="4CBE0710FFB7DC0C4A06FC996FA3FC0E" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="15" pageNumber="56" unit="m" value="950.0">
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<quantity id="006B4DC6FFB7DC0C4A06FC996FA3FC0E" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="15" pageNumber="56" unit="m" value="950.0">950 m</quantity>
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</elevation>
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, sieving in the forest near river,
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<collectingDate id="A3693F0BFFB7DC0C4D6DFCB66DEBFC0D" box="[537,743,889,914]" pageId="15" pageNumber="56" value="1972-11-09">
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<date id="B32DC6E3FFB7DC0C4D6DFCB66DEBFC0D" box="[537,743,889,914]" pageId="15" pageNumber="56" value="1972-11-09">9 November 1972</date>
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</collectingDate>
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, coll.
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<collectorName id="6A6685F5FFB7DC0C4C5EFCB66CB9FC0D" box="[810,949,889,914]" pageId="15" pageNumber="56">C. Besuchet</collectorName>
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/
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<collectorName id="6A6685F5FFB7DC0C4CCAFCB56B0AFC0D" box="[958,1030,889,914]" pageId="15" pageNumber="56">I. Löbl</collectorName>
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(18) (
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<collectionCode id="A18278E6FFB7DC0C4B3FFCB66BABFC0D" box="[1099,1191,890,914]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="56">MHNG</collectionCode>
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)
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</materialsCitation>
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; paratypes (male and female) (MHNG).
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</paragraph>
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</subSubSection>
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<subSubSection id="8F89B3A8FFB7DC0C4FB3FC0D6C7FFC45" box="[199,883,961,986]" pageId="15" pageNumber="56" type="etymology">
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<paragraph id="C72CE023FFB7DC0C4FB3FC0D6C7FFC45" blockId="15.[151,1437,852,2030]" box="[199,883,961,986]" pageId="15" pageNumber="56">
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||
<emphasis id="F5E73C31FFB7DC0C4FB3FC0D6E42FC46" bold="true" box="[199,334,961,985]" pageId="15" pageNumber="56">Etymology.</emphasis>
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The species is named after the country of origin.
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</paragraph>
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</subSubSection>
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<subSubSection id="8F89B3A8FFB7DC114FB3FC286D13FE4F" lastPageId="18" lastPageNumber="59" pageId="15" pageNumber="56" type="description">
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||
<paragraph id="C72CE023FFB7DC0C4FB3FC286D18F951" blockId="15.[151,1437,852,2030]" pageId="15" pageNumber="56">
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<emphasis id="F5E73C31FFB7DC0C4FB3FC286E5DFC62" bold="true" box="[199,337,996,1021]" pageId="15" pageNumber="56">Description</emphasis>
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. Male (
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<typeStatus id="18285E81FFB7DC0C4EC5FC296D16FC61" box="[433,538,997,1022]" pageId="15" pageNumber="56" type="holotype">holotype</typeStatus>
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). General colour yellowish, partly blackish (
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<figureCitation id="5FA8FCA6FFB7DC0C4B52FC296B60FC61" box="[1062,1132,997,1022]" captionStart="FIGURE 8" captionStartId="16.[151,249,1593,1615]" captionTargetBox="[153,1429,219,1567]" captionTargetId="figure@16.[149,1434,211,1572]" captionTargetPageId="16" captionText="FIGURE 8. Morphna ssp. A, B, E, F—facial part of head; C, D, G, H—anterior part of body from above. A – D— M. indica sp. nov., male, holotype (A, C) and female, paratype (B, D); E – H— M. srilankensis sp. nov., male, holotype (E, G) and female, paratype (F, H). Pronotum on fig. G damaged. Out of scale." httpUri="https://zenodo.org/record/322022/files/figure.png" pageId="15" pageNumber="56">Fig. 8</figureCitation>
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A, C); facial part of head contrastingly coloured (
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<figureCitation id="5FA8FCA6FFB7DC0C4ED2FBC56EE7FBBD" box="[422,491,1033,1058]" captionStart="FIGURE 8" captionStartId="16.[151,249,1593,1615]" captionTargetBox="[153,1429,219,1567]" captionTargetId="figure@16.[149,1434,211,1572]" captionTargetPageId="16" captionText="FIGURE 8. Morphna ssp. A, B, E, F—facial part of head; C, D, G, H—anterior part of body from above. A – D— M. indica sp. nov., male, holotype (A, C) and female, paratype (B, D); E – H— M. srilankensis sp. nov., male, holotype (E, G) and female, paratype (F, H). Pronotum on fig. G damaged. Out of scale." httpUri="https://zenodo.org/record/322022/files/figure.png" pageId="15" pageNumber="56">Fig. 8</figureCitation>
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A); pronotum (
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<figureCitation id="5FA8FCA6FFB7DC0C4DEDFBC56DD2FBBD" box="[665,734,1033,1058]" captionStart="FIGURE 8" captionStartId="16.[151,249,1593,1615]" captionTargetBox="[153,1429,219,1567]" captionTargetId="figure@16.[149,1434,211,1572]" captionTargetPageId="16" captionText="FIGURE 8. Morphna ssp. A, B, E, F—facial part of head; C, D, G, H—anterior part of body from above. A – D— M. indica sp. nov., male, holotype (A, C) and female, paratype (B, D); E – H— M. srilankensis sp. nov., male, holotype (E, G) and female, paratype (F, H). Pronotum on fig. G damaged. Out of scale." httpUri="https://zenodo.org/record/322022/files/figure.png" pageId="15" pageNumber="56">Fig. 8</figureCitation>
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C), tegmina and abdomen yellow with numerous small black dots forming larger dark spots; legs yellow, coxae and femora with sparse brown dots, spines and apical parts of tarsi darkened, dark brown. Surfaces smooth and lustrous; antennae with lustrous proximal 17–18 segments, the rest of segments dull; head, pronotum and tegmina somewhat rugose, moderately punctuated; facial part of head with weak transverse wrinkles. Head widely rounded at vertex, longer than wide, epicranial sutures present (
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<figureCitation id="5FA8FCA6FFB7DC0C4A10FB556FA8FB49" captionStart="FIGURE 8" captionStartId="16.[151,249,1593,1615]" captionTargetBox="[153,1429,219,1567]" captionTargetId="figure@16.[149,1434,211,1572]" captionTargetPageId="16" captionText="FIGURE 8. Morphna ssp. A, B, E, F—facial part of head; C, D, G, H—anterior part of body from above. A – D— M. indica sp. nov., male, holotype (A, C) and female, paratype (B, D); E – H— M. srilankensis sp. nov., male, holotype (E, G) and female, paratype (F, H). Pronotum on fig. G damaged. Out of scale." httpUri="https://zenodo.org/record/322022/files/figure.png" pageId="15" pageNumber="56">Figs. 8</figureCitation>
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A, 9C); ocellar spots distinct; distance between eyes 1.1 times eye length; distance between antennal sockets about twice of the scape length (about
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<quantity id="006B4DC6FFB7DC0C4D24FB2D6DA0FB65" box="[592,684,1249,1274]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="56" unit="mm" value="1.5">1.5 mm</quantity>
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); approximate length ratio of 3rd–5th segments of maxillary palps 1.0: 1.0: 1.2. Pronotum transverse, widely rounded along anterolateral margins, caudal margin slightly angulate (
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<figureCitation id="5FA8FCA6FFB7DC0C4FEBFAE56FE1FADD" box="[159,237,1321,1346]" captionStart="FIGURE 8" captionStartId="16.[151,249,1593,1615]" captionTargetBox="[153,1429,219,1567]" captionTargetId="figure@16.[149,1434,211,1572]" captionTargetPageId="16" captionText="FIGURE 8. Morphna ssp. A, B, E, F—facial part of head; C, D, G, H—anterior part of body from above. A – D— M. indica sp. nov., male, holotype (A, C) and female, paratype (B, D); E – H— M. srilankensis sp. nov., male, holotype (E, G) and female, paratype (F, H). Pronotum on fig. G damaged. Out of scale." httpUri="https://zenodo.org/record/322022/files/figure.png" pageId="15" pageNumber="56">Figs. 8</figureCitation>
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C, 9D). Scutellum triangular, small. Tegmina subquadrate (
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<figureCitation id="5FA8FCA6FFB7DC0C4CFFFAE56CC3FADD" box="[907,975,1321,1346]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
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D), only reaching 2nd abdominal tergite; venation subobsolete;
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<emphasis id="F5E73C31FFB7DC0C4EE2FA826EBCFAFA" box="[406,432,1358,1381]" italics="true" pageId="15" pageNumber="56">Sc</emphasis>
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thickened (well visible on ventral side of tegmen). Wings vestigial, completely hidden under tegmina. Anterior margin of fore femur armed as in the
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<typeStatus id="18285E81FFB7DC0C4C39FABE6C71FA15" box="[845,893,1394,1418]" pageId="15" pageNumber="56">type</typeStatus>
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B, with 5 spines, including 2 apical ones. Tibiae distinctly thickened distally. Structure of hind tarsi (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4C77FA596C40FA31" box="[771,844,1429,1454]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
E): metatarsus a little shorter than other segments combined, with large euplantula along lower margin; tarsal spines absent; euplantulae of 1st–4th tarsal segments with distinct spinules (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4EE1FA116ED4FA69" box="[405,472,1501,1526]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
E,
|
||
<emphasis id="F5E73C31FFB7DC0C4E83FA126D13FA6A" box="[503,543,1502,1525]" italics="true" pageId="15" pageNumber="56">spi.</emphasis>
|
||
); claws symmetrical, simple; arolium longer than half of claw length (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4A4CFA116A70FA69" box="[1336,1404,1501,1526]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
E). Abdomen without visible glandular specializations; posterolateral angles of tergites slightly attenuate caudally; spiracle-bearing outgrowths of tergite VIII with sharp and attenuate posterolateral angles (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4BFDF9E96BC7F9A1" box="[1161,1227,1573,1598]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
F,
|
||
<emphasis id="F5E73C31FFB7DC0C4B93F9EA6A1DF9A2" box="[1255,1297,1574,1597]" italics="true" pageId="15" pageNumber="56">p.a.</emphasis>
|
||
). Anal plate (tergite X) widely rounded, with weak and wide median incision on caudal margin (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4B38F9856B9CF9FD" box="[1100,1168,1609,1634]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
F). Cerci fusiform, very weakly depressed, with distinct segments (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4DF0F9A16DC6F919" box="[644,714,1645,1670]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
F). Paraprocts of the blaberid-type, as in
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4BDCF9A16BE0F919" box="[1192,1260,1645,1670]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
G. Hypandrium nearly symmetrical, its caudal margin rounded and projected caudally, with a distinct median incision (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4A47F95D6A74F935" box="[1331,1400,1681,1706]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
H); styli short and fusiform (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4ED8F9796EFCF951" box="[428,496,1717,1742]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
H).
|
||
</paragraph>
|
||
<paragraph id="C72CE023FFB7DC0C4FB3F9156E14F871" blockId="15.[151,1437,852,2030]" pageId="15" pageNumber="56">
|
||
Male genitalia (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4E0EF9156EB2F96D" box="[378,446,1753,1778]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">
|
||
Fig.
|
||
<date id="B32DC6E3FFB7DC0C4EC4F9166EB2F96D" box="[432,446,1754,1778]" pageId="15" pageNumber="56">9</date>
|
||
</figureCitation>
|
||
I–P). Right phallomere (R+N) with caudal part of sclerite R1T subrectangular in shape (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4FEBF9316FE8F889" box="[159,228,1789,1814]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">
|
||
Fig.
|
||
<date id="B32DC6E3FFB7DC0C4FA1F9326FE8F889" box="[213,228,1790,1814]" pageId="15" pageNumber="56">9</date>
|
||
</figureCitation>
|
||
I, J,
|
||
<emphasis id="F5E73C31FFB7DC0C4E6CF9326E63F889" box="[280,367,1790,1814]" italics="true" pageId="15" pageNumber="56">c.p.R1T</emphasis>
|
||
); densely covered with bristles; R2 curved; R3 elongated, widened caudally; R4 large, platelike; R
|
||
<quantity id="006B4DC6FFB7DC0C4F91F8ED6E1CF8A5" box="[229,272,1825,1850]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="15" pageNumber="56" unit="in" value="5.0">5 in</quantity>
|
||
shape of elongated plate, fused with sclerite R3. Sclerite L2D (L1) divided into basal and apical parts (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4FEBF8896FE9F8C1" box="[159,229,1861,1886]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
K); basal part rod-like, distinctly widened cranially, with "curved upward subsclerite" (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4BA6F8896A1BF8C1" box="[1234,1303,1861,1886]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
L,
|
||
<emphasis id="F5E73C31FFB7DC0C4A4DF88A6A6CF8C2" box="[1337,1376,1862,1885]" italics="true" pageId="15" pageNumber="56">u.s.</emphasis>
|
||
) and lamellate outgrowth (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4EE4F8A56EDBF81D" box="[400,471,1897,1922]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
L,
|
||
<emphasis id="F5E73C31FFB7DC0C4E8EF8A56D11F81D" box="[506,541,1897,1922]" italics="true" pageId="15" pageNumber="56">l.o.</emphasis>
|
||
) at caudal end; apical part in shape of crescentic sclerite and small "dorsal outgrows" (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4E6FF8416E53F839" box="[283,351,1933,1958]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
K, L,
|
||
<emphasis id="F5E73C31FFB7DC0C4ED7F8416EC0F839" box="[419,460,1933,1958]" italics="true" pageId="15" pageNumber="56">d.o.</emphasis>
|
||
); bristles absent. Sclerite L3 (L2d) with basal subsclerite (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4B1CF8416BA0F839" box="[1128,1196,1933,1958]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
M,
|
||
<emphasis id="F5E73C31FFB7DC0C4BA0F8416A0AF839" box="[1236,1286,1933,1958]" italics="true" pageId="15" pageNumber="56">b.L3</emphasis>
|
||
) and "folded structure" (
|
||
<figureCitation id="5FA8FCA6FFB7DC0C4E61F87D6E54F855" box="[277,344,1969,1994]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="15" pageNumber="56">Fig. 9</figureCitation>
|
||
M,
|
||
<emphasis id="F5E73C31FFB7DC0C4EF4F87E6EAEF855" box="[384,418,1970,1994]" italics="true" pageId="15" pageNumber="56">f.s.</emphasis>
|
||
) with bristles; groove hge absent. Sclerite L4U (L3d) in shape of elongated triangle, well sclerotized.
|
||
</paragraph>
|
||
<caption id="93ECB0ABFFA8DC134FE3F9F56DFDF913" httpUri="https://zenodo.org/record/322022/files/figure.png" pageId="16" pageNumber="57" targetBox="[153,1429,219,1567]" targetPageId="16">
|
||
<paragraph id="C72CE023FFA8DC134FE3F9F56DFDF913" blockId="16.[151,1436,1593,1676]" pageId="16" pageNumber="57">
|
||
<emphasis id="F5E73C31FFA8DC134FE3F9F56E1FF9D0" bold="true" box="[151,275,1593,1615]" pageId="16" pageNumber="57">FIGURE 8.</emphasis>
|
||
<taxonomicName id="00939BA0FFA8DC134E68F9F56E76F9D0" box="[284,378,1593,1615]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F5E73C31FFA8DC134E68F9F56E76F9D0" box="[284,378,1593,1615]" italics="true" pageId="16" pageNumber="57">Morphna</emphasis>
|
||
</taxonomicName>
|
||
ssp. A, B, E, F—facial part of head; C, D, G, H—anterior part of body from above. A–D—
|
||
<taxonomicName id="00939BA0FFA8DC134A4DF9F66A90F9D0" box="[1337,1436,1593,1615]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFA8DC134A4DF9F66A42F9D0" box="[1337,1358,1594,1615]" italics="true" pageId="16" pageNumber="57">M</emphasis>
|
||
.
|
||
<emphasis id="F5E73C31FFA8DC134A28F9F56A90F9D0" box="[1372,1436,1593,1615]" italics="true" pageId="16" pageNumber="57">indica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFA8DC134FE3F9946FEEF9F1" box="[151,226,1624,1646]" pageId="16" pageNumber="57" rank="species">
|
||
<emphasis id="F5E73C31FFA8DC134FE3F9946FEEF9F1" bold="true" box="[151,226,1624,1646]" pageId="16" pageNumber="57">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male, holotype (A, C) and female, paratype (B, D); E–H—
|
||
<taxonomicName id="00939BA0FFA8DC134C33F9956CEEF9F1" box="[839,994,1624,1646]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="srilankensis" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFA8DC134C33F9956C50F9F1" box="[839,860,1625,1646]" italics="true" pageId="16" pageNumber="57">M</emphasis>
|
||
.
|
||
<emphasis id="F5E73C31FFA8DC134C1DF9946CEEF9F1" box="[873,994,1624,1646]" italics="true" pageId="16" pageNumber="57">srilankensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFA8DC134C9DF9946B38F9F1" box="[1001,1076,1624,1646]" pageId="16" pageNumber="57" rank="species">
|
||
<emphasis id="F5E73C31FFA8DC134C9DF9946B38F9F1" bold="true" box="[1001,1076,1624,1646]" pageId="16" pageNumber="57">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male, holotype (E, G) and female, paratype (F, H). Pronotum on fig. G damaged. Out of scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C72CE023FFA8DC134FB3F9756BB5F94D" blockId="16.[151,1436,1721,1998]" box="[199,1209,1721,1746]" pageId="16" pageNumber="57">
|
||
<emphasis id="F5E73C31FFA8DC134FB3F9756E4BF94D" bold="true" box="[199,327,1721,1746]" pageId="16" pageNumber="57">Variations.</emphasis>
|
||
The male
|
||
<typeStatus id="18285E81FFA8DC134EB4F9766D2DF94D" box="[448,545,1722,1746]" pageId="16" pageNumber="57" type="paratype">paratype</typeStatus>
|
||
is similar with the
|
||
<typeStatus id="18285E81FFA8DC134D8CF9756C51F94D" box="[760,861,1721,1746]" pageId="16" pageNumber="57" type="holotype">holotype</typeStatus>
|
||
, but slightly smaller in its size.
|
||
</paragraph>
|
||
<paragraph id="C72CE023FFA8DC134FB3F9116E13F851" blockId="16.[151,1436,1721,1998]" pageId="16" pageNumber="57">
|
||
Female (
|
||
<figureCitation id="5FA8FCA6FFA8DC134E58F9116E7EF969" box="[300,370,1757,1782]" captionStart="FIGURE 8" captionStartId="16.[151,249,1593,1615]" captionTargetBox="[153,1429,219,1567]" captionTargetId="figure@16.[149,1434,211,1572]" captionTargetPageId="16" captionText="FIGURE 8. Morphna ssp. A, B, E, F—facial part of head; C, D, G, H—anterior part of body from above. A – D— M. indica sp. nov., male, holotype (A, C) and female, paratype (B, D); E – H— M. srilankensis sp. nov., male, holotype (E, G) and female, paratype (F, H). Pronotum on fig. G damaged. Out of scale." httpUri="https://zenodo.org/record/322022/files/figure.png" pageId="16" pageNumber="57">Fig. 8</figureCitation>
|
||
B, D). Similar to the male; distance between antennal sockets 2.3 times of the scape length (about
|
||
<quantity id="006B4DC6FFA8DC134F92F8CE6E4CF885" box="[230,320,1794,1818]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="57" unit="mm" value="1.4">1.4 mm</quantity>
|
||
); approximate length ratio of 3rd–5th segments of maxillary palps 1.0: 1.0: 1.1; anal plate (tergite X) and genital plate similar to that of
|
||
<taxonomicName id="00939BA0FFA8DC134D48F8EA6DBFF8A1" box="[572,691,1829,1854]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="decolyi">
|
||
<emphasis id="F5E73C31FFA8DC134D48F8EA6D5FF8A2" box="[572,595,1830,1853]" italics="true" pageId="16" pageNumber="57">M</emphasis>
|
||
.
|
||
<emphasis id="F5E73C31FFA8DC134D16F8E96DBFF8A1" box="[610,691,1829,1854]" italics="true" pageId="16" pageNumber="57">decolyi</emphasis>
|
||
</taxonomicName>
|
||
, but genital plate is notched in the middle of caudal margin. Measurements (mm). Head length: male 5.6–6.0 (6.0), female 6.4; head width: male 5.2–5.5 (5.5), female 5.4; pronotum length: male 7.9–8.8 (8.8), female 8.7; pronotum width: male 13.4–14.5 (14.5), female 14.2; tegmen length: male 9.2 (9.2), female 8.6; tegmen width: male 9.0 (9.0), female 9.0. Measurements in parenthesis are those of
|
||
<typeStatus id="18285E81FFA8DC134FC1F8796E17F851" box="[181,283,1973,1998]" pageId="16" pageNumber="57" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<caption id="93ECB0ABFFA9DC124FE3F8ED6DB7F870" httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="17" pageNumber="58" targetBox="[182,1402,204,1786]" targetPageId="17">
|
||
<paragraph id="C72CE023FFA9DC124FE3F8ED6DB7F870" blockId="17.[151,1437,1825,2031]" pageId="17" pageNumber="58">
|
||
<emphasis id="F5E73C31FFA9DC124FE3F8ED6E19F8A8" bold="true" box="[151,277,1825,1847]" pageId="17" pageNumber="58">FIGURE 9.</emphasis>
|
||
<taxonomicName id="00939BA0FFA9DC124E6AF8ED6E70F8A8" box="[286,380,1825,1847]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="17" pageNumber="58" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F5E73C31FFA9DC124E6AF8ED6E70F8A8" box="[286,380,1825,1847]" italics="true" pageId="17" pageNumber="58">Morphna</emphasis>
|
||
</taxonomicName>
|
||
ssp., males:
|
||
<taxonomicName id="00939BA0FFA9DC124D72F8EE6C16F8A8" authority="(Bolivar, 1897)" authorityName="Bolivar" authorityYear="1897" baseAuthorityName="Bolivar" baseAuthorityYear="1897" box="[518,794,1825,1847]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="decolyi">
|
||
<emphasis id="F5E73C31FFA9DC124D72F8EE6D7FF8A8" box="[518,627,1825,1847]" italics="true" pageId="17" pageNumber="58">M. decolyi</emphasis>
|
||
(Bolivar, 1897)
|
||
</taxonomicName>
|
||
(A, B) and
|
||
<taxonomicName id="00939BA0FFA9DC124CE8F8EE6B0CF8A8" box="[924,1024,1825,1847]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFA9DC124CE8F8EE6CBDF8A8" box="[924,945,1826,1847]" italics="true" pageId="17" pageNumber="58">M</emphasis>
|
||
.
|
||
<emphasis id="F5E73C31FFA9DC124CB5F8ED6B0CF8A8" box="[961,1024,1825,1847]" italics="true" pageId="17" pageNumber="58">indica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFA9DC124B7DF8ED6B58F8A8" box="[1033,1108,1825,1847]" pageId="17" pageNumber="58" rank="species">
|
||
<emphasis id="F5E73C31FFA9DC124B7DF8ED6B58F8A8" bold="true" box="[1033,1108,1825,1847]" pageId="17" pageNumber="58">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(C–P). A—left hind tibia from outside; B, K—sclerite L2D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L2D from above; M—sclerite L3; N–P—apical parts of sclerite L3. Dotted area shows membranous parts (not shown in fig E). Abbreviations:
|
||
<emphasis id="F5E73C31FFA9DC124FE3F8776FC8F84E" box="[151,196,1979,2001]" italics="true" pageId="17" pageNumber="58">b.L3</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124FA5F8776E11F84E" box="[209,285,1979,2001]" italics="true" pageId="17" pageNumber="58">c.p.R1T</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124E5FF8776E5DF84E" box="[299,337,1979,2001]" italics="true" pageId="17" pageNumber="58">d.o.</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124E29F8776E71F84E" box="[349,381,1979,2001]" italics="true" pageId="17" pageNumber="58">l.o.</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124EFEF8776EA5F84E" box="[394,425,1979,2001]" italics="true" pageId="17" pageNumber="58">f.s.</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124EC0F8706ED7F84E" box="[436,475,1980,2001]" italics="true" pageId="17" pageNumber="58">p.a.</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124E93F8776D0FF84E" box="[487,515,1979,2001]" italics="true" pageId="17" pageNumber="58">R2</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124D7BF8776D27F84E" box="[527,555,1979,2001]" italics="true" pageId="17" pageNumber="58">R3</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124D4CF8776D58F84E" box="[568,596,1979,2001]" italics="true" pageId="17" pageNumber="58">R4</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124D15F8776D71F84F" box="[609,637,1979,2000]" italics="true" pageId="17" pageNumber="58">R5</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124DFEF8706DA2F84E" box="[650,686,1980,2001]" italics="true" pageId="17" pageNumber="58">spi.</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFA9DC124DCEF8706C04F84E" box="[698,776,1980,2001]" italics="true" pageId="17" pageNumber="58">t.s., u.s.</emphasis>
|
||
—see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N–P.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C72CE023FFAADC114FB3FF5B6C9BFE17" blockId="18.[151,1436,151,464]" pageId="18" pageNumber="59">
|
||
<emphasis id="F5E73C31FFAADC114FB3FF5B6E6EFF2F" bold="true" box="[199,354,151,176]" pageId="18" pageNumber="59">Comparison.</emphasis>
|
||
The new species is closely related to
|
||
<taxonomicName id="00939BA0FFAADC114C59FF556CA4FF2F" box="[813,936,151,176]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="decolyi">
|
||
<emphasis id="F5E73C31FFAADC114C59FF556C48FF2F" box="[813,836,153,176]" italics="true" pageId="18" pageNumber="59">M</emphasis>
|
||
.
|
||
<emphasis id="F5E73C31FFAADC114C23FF5B6CA4FF2F" box="[855,936,151,176]" italics="true" pageId="18" pageNumber="59">decolyi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="00939BA0FFAADC114C93FF556B9BFF2F" box="[999,1175,151,176]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="srilankensis" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFAADC114C93FF556CF2FF2F" box="[999,1022,153,176]" italics="true" pageId="18" pageNumber="59">M</emphasis>
|
||
.
|
||
<emphasis id="F5E73C31FFAADC114B65FF5B6B9BFF2F" box="[1041,1175,151,176]" italics="true" pageId="18" pageNumber="59">srilankensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F5E73C31FFAADC114BD7FF546BF9FF2F" bold="true" box="[1187,1269,152,176]" pageId="18" pageNumber="59">
|
||
<taxonomicNameLabel id="EED4814AFFAADC114BD7FF546BF9FF2F" box="[1187,1269,152,176]" pageId="18" pageNumber="59" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having the shortened subquadrate tegmina.
|
||
<taxonomicName id="00939BA0FFAADC114D66FF716D8FFF4A" box="[530,643,188,213]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFAADC114D66FF716D8FFF4A" box="[530,643,188,213]" italics="true" pageId="18" pageNumber="59">M. indica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFAADC114DE4FF706DEEFF4B" box="[656,738,188,212]" pageId="18" pageNumber="59" rank="species">
|
||
<emphasis id="F5E73C31FFAADC114DE4FF706DEEFF4B" bold="true" box="[656,738,188,212]" pageId="18" pageNumber="59">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="00939BA0FFAADC114B5AFF716BA6FF4A" box="[1070,1194,188,213]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="decolyi">
|
||
<emphasis id="F5E73C31FFAADC114B5AFF716B49FF4B" box="[1070,1093,189,212]" italics="true" pageId="18" pageNumber="59">M</emphasis>
|
||
.
|
||
<emphasis id="F5E73C31FFAADC114B2DFF706BA6FF4A" box="[1113,1194,188,213]" italics="true" pageId="18" pageNumber="59">decolyi</emphasis>
|
||
</taxonomicName>
|
||
by the lighter body coloration (compare
|
||
<figureCitation id="5FA8FCA6FFAADC114EF5FF136ECDFF67" box="[385,449,223,248]" captionStart="FIGURE 8" captionStartId="16.[151,249,1593,1615]" captionTargetBox="[153,1429,219,1567]" captionTargetId="figure@16.[149,1434,211,1572]" captionTargetPageId="16" captionText="FIGURE 8. Morphna ssp. A, B, E, F—facial part of head; C, D, G, H—anterior part of body from above. A – D— M. indica sp. nov., male, holotype (A, C) and female, paratype (B, D); E – H— M. srilankensis sp. nov., male, holotype (E, G) and female, paratype (F, H). Pronotum on fig. G damaged. Out of scale." httpUri="https://zenodo.org/record/322022/files/figure.png" pageId="18" pageNumber="59">Fig. 8</figureCitation>
|
||
A–D and
|
||
<figureCitation id="5FA8FCA6FFAADC114D44FF136D61FF67" box="[560,621,223,248]" captionStart="FIGURE 1" captionStartId="3.[151,249,1841,1863]" captionTargetBox="[158,1428,206,1803]" captionTargetId="figure@3.[151,1436,193,1819]" captionTargetPageId="3" captionText="FIGURE 1. Aptera fusca (Thunberg, 1784), female (A, B) and A. brindlei sp. nov., male, holotype (C – F). A—structures of abdominal apex and female genitalia from below, genital plate removed; B—female genitalia from below; C—facial part of head; D—anterior part of body from above; E—VI and VII abdominal tergites from above; F—hypandrium and male genitalia from above. Out of scale." httpUri="https://zenodo.org/record/322015/files/figure.png" pageId="18" pageNumber="59">Fig. 1</figureCitation>
|
||
H–J in
|
||
<bibRefCitation id="A3029DD2FFAADC114DB7FF136C8FFF67" author="Anisyutkin" box="[707,899,223,248]" pageId="18" pageNumber="59" refString="Anisyutkin, L. N. (2014) On cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from South India and Sri Lanka, with descriptions of new taxa. Zootaxa, 3847 (3), 301 - 332. http: // dx. doi. org / 10.11646 / zootaxa. 3847.3.1" type="journal article" year="2014">Anisyutkin 2014</bibRefCitation>
|
||
), the presence of attenuate posterolateral angles on the tergites VIII (compare
|
||
<figureCitation id="5FA8FCA6FFAADC114E81FEC86D35FE83" box="[501,569,260,285]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="18" pageNumber="59">Fig. 9</figureCitation>
|
||
F and
|
||
<figureCitation id="5FA8FCA6FFAADC114DF3FEC86DD9FE82" box="[647,725,260,285]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII – VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI – VIII— 6 th – 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="18" pageNumber="59">Figs. 6</figureCitation>
|
||
G, 7F in
|
||
<bibRefCitation id="A3029DD2FFAADC114C35FEC86B08FE83" author="Anisyutkin" box="[833,1028,260,285]" pageId="18" pageNumber="59" refString="Anisyutkin, L. N. (2014) On cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from South India and Sri Lanka, with descriptions of new taxa. Zootaxa, 3847 (3), 301 - 332. http: // dx. doi. org / 10.11646 / zootaxa. 3847.3.1" type="journal article" year="2014">Anisyutkin 2014</bibRefCitation>
|
||
), spinules on the tarsal euplantulae (compare
|
||
<figureCitation id="5FA8FCA6FFAADC114E7CFEEB6E45FEDF" box="[264,329,295,320]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="18" pageNumber="59">Fig. 9</figureCitation>
|
||
E and
|
||
<figureCitation id="5FA8FCA6FFAADC114EE6FEEB6ED8FEDF" box="[402,468,295,320]" captionStart="FIGURE 6" captionStartId="12.[151,249,1832,1854]" captionTargetBox="[214,1354,204,1795]" captionTargetId="figure@12.[206,1362,195,1810]" captionTargetPageId="12" captionText="FIGURE 6. Rhabdoblattella euptera sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C, D—VI (C) and VII – VIII (D) abdominal tergites from above; E—abdominal apex from above; F, G—paraprocts from caudal (F) and below (G); H—hypandrium from below; I—caudal margin of hypandrium from caudal; J—right phallomere from above; K—sclerite L 2 D from above; L—apical part of sclerite L 2 D from above; M—sclerites L 3 and L 4 U; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: VI – VIII— 6 th – 8 th abdominal tergites respectively, c. p. R 1 T, cl., cr. p. R 1 T, ch. a., f. s., l. out., L 4 U, m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E; e = F; f = G; g = H; h = I; i = J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322020/files/figure.png" pageId="18" pageNumber="59">Fig. 6</figureCitation>
|
||
E, D in
|
||
<bibRefCitation id="A3029DD2FFAADC114D58FEEB6DE6FEDF" author="Anisyutkin" box="[556,746,295,320]" pageId="18" pageNumber="59" refString="Anisyutkin, L. N. (2014) On cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from South India and Sri Lanka, with descriptions of new taxa. Zootaxa, 3847 (3), 301 - 332. http: // dx. doi. org / 10.11646 / zootaxa. 3847.3.1" type="journal article" year="2014">Anisyutkin 2014</bibRefCitation>
|
||
) and the structure of the apical part of sclerite L2D (compare
|
||
<figureCitation id="5FA8FCA6FFAADC114FE3FE806FD4FEFB" box="[151,216,332,357]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="18" pageNumber="59">Fig. 9</figureCitation>
|
||
B and
|
||
<figureCitation id="5FA8FCA6FFAADC114E54FE806E6DFEFB" box="[288,353,332,357]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="18" pageNumber="59">Fig. 9</figureCitation>
|
||
K, L). From
|
||
<taxonomicName id="00939BA0FFAADC114E9AFE816D96FEFA" box="[494,666,332,357]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="srilankensis" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFAADC114E9AFE816D09FEFB" box="[494,517,333,356]" italics="true" pageId="18" pageNumber="59">M</emphasis>
|
||
.
|
||
<emphasis id="F5E73C31FFAADC114D60FE806D96FEFA" box="[532,666,332,357]" italics="true" pageId="18" pageNumber="59">srilankensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFAADC114DD4FE806DFEFEFB" box="[672,754,332,356]" pageId="18" pageNumber="59" rank="species">
|
||
<emphasis id="F5E73C31FFAADC114DD4FE806DFEFEFB" bold="true" box="[672,754,332,356]" pageId="18" pageNumber="59">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, the new species differs in the larger size and the structure of the apical part of sclerite L2D (compare
|
||
<figureCitation id="5FA8FCA6FFAADC114D2AFEA36D93FE17" box="[606,671,367,392]" captionStart="FIGURE 9" captionStartId="17.[151,249,1825,1847]" captionTargetBox="[182,1402,204,1786]" captionTargetId="figure@17.[174,1408,193,1789]" captionTargetPageId="17" captionText="FIGURE 9. Morphna ssp., males: M. decolyi (Bolivar, 1897) (A, B) and M. indica sp. nov. (C – P). A—left hind tibia from outside; B, K—sclerite L 2 D from above; C—facial part of head; D—pronotum and left tegmen from above, outlines; E—right hind tarsus from below; F—abdominal apex from above; G—abdominal apex from caudal, genitalia and hypandrium removed; H—hypandrium from below; I, J—right phallomere from above (I) and below (J); L—apical part of sclerite L 2 D from above; M—sclerite L 3; N – P—apical parts of sclerite L 3. Dotted area shows membranous parts (not shown in fig E). Abbreviations: b. L 3, c. p. R 1 T, d. o., l. o., f. s., p. a., R 2, R 3, R 4, R 5, spi., t. s., u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H; i = I, J; j = K; k = L; l = M; m = N – P." httpUri="https://zenodo.org/record/322023/files/figure.png" pageId="18" pageNumber="59">Fig. 9</figureCitation>
|
||
K, L and
|
||
<figureCitation id="5FA8FCA6FFAADC114C7DFEA36C5BFE17" box="[777,855,367,392]" captionStart="FIGURE 10" captionStartId="19.[151,249,1772,1794]" captionTargetBox="[164,1427,313,1742]" captionTargetId="figure@19.[151,1436,303,1750]" captionTargetPageId="19" captionText="FIGURE 10. Morphna srilankensis sp. nov., male, holotype. A—facial part of head; B—abdominal apex from above; Chypandrium from below; D—right phallomere from above; E—sclerites R 3 and R 5 from below, outlines; F—sclerites L 2 D and L 4 U from above; G—apical part of sclerite L 2 D from above; H – J—apical parts of sclerite L 3. Dotted area shows membranous parts. Abbreviations: ap. cr., c. p. R 1 T, l. o., L 4 U, p. a., R 3, R 4, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G; g = H – J." httpUri="https://zenodo.org/record/322024/files/figure.png" pageId="18" pageNumber="59">Fig. 10</figureCitation>
|
||
G, F).
|
||
</paragraph>
|
||
<paragraph id="C72CE023FFAADC114FB3FE596D13FE4F" blockId="18.[151,1436,151,464]" pageId="18" pageNumber="59">
|
||
From all other congeners,
|
||
<taxonomicName id="00939BA0FFAADC114E98FE596D5AFE32" box="[492,598,404,429]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="59" phylum="Arthropoda" rank="species" species="indica" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFAADC114E98FE596D0FFE33" box="[492,515,405,428]" italics="true" pageId="18" pageNumber="59">M</emphasis>
|
||
.
|
||
<emphasis id="F5E73C31FFAADC114D64FE586D5AFE32" box="[528,598,404,429]" italics="true" pageId="18" pageNumber="59">indica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFAADC114D29FE586DA3FE33" box="[605,687,404,428]" pageId="18" pageNumber="59" rank="species">
|
||
<emphasis id="F5E73C31FFAADC114D29FE586DA3FE33" bold="true" box="[605,687,404,428]" pageId="18" pageNumber="59">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be readily separated by the presence of shortened, subquadrate tegmina with an obsolete venation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |