309 lines
55 KiB
XML
309 lines
55 KiB
XML
<document id="5C9BDCC31D0641B0BE0F283EB78F0717" ID="10.11646/zootaxa.4236.1.2SLASH10251" ID-CLB-Dataset="36479" ID-DOI="10.11646/zootaxa.4236.1.2" ID-GBIF-Dataset="0f39b5c0-0af5-476f-8448-a02339cc1dfd" ID-ISSN="1175-5326" ID-Zenodo-Dep="322014" ID-ZooBank="23412386-CB17-49CA-9C47-BD71DD9C5372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487752177037" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N. & Yushkova, Olga V." docDate="2017" docId="4F3A5135FFB0DC084FE3FD146E5BFE82" docLanguage="en" docName="zootaxa.4236.1.2SLASH10251.pdf" docOrigin="Zootaxa 4236 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhabdoblattella alexeevi Anisyutkin, sp.nov." docType="treatment" docVersion="7" lastPageNumber="52" masterDocId="B303294DFFB8DC034F74FFCC6F0CFF9F" masterDocTitle="New data on cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from India and Sri Lanka, with descriptions of new species and the genital complex of Aptera fusca (Thunberg, 1784)" masterLastPageNumber="64" masterPageNumber="41" pageNumber="49" updateTime="1698437163562" updateUser="ExternalLinkService">
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<mods:mods id="620AD953DA3CD40A241C14877FB04275" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="7509DB66493DF0E0CA165E6037598802">
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<mods:title id="EA47F9FBF0CE13C45E2DFF0FBBED4F70">New data on cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from India and Sri Lanka, with descriptions of new species and the genital complex of Aptera fusca (Thunberg, 1784)</mods:title>
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<mods:roleTerm id="573BE970CBC4D00729F43DF08B3A1C65">Author</mods:roleTerm>
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<mods:namePart id="F197EB8C4B54442C52A91283707DA6EB">Anisyutkin, Leonid N.</mods:namePart>
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<mods:name id="862CFDC4F3787FD950ABD76C5E4A4435" type="personal">
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</mods:role>
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<mods:namePart id="5E5F2A8D892B6323013EE7911281887B">Yushkova, Olga V.</mods:namePart>
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<mods:typeOfResource id="FF7E7690C59336330602462F8D390828">text</mods:typeOfResource>
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<mods:title id="75A63A1D5C3AD2DA7226BAE914E81D26">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1008FA9F80C01CE90D7A7C628DFC37F2">
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<mods:date id="C1DD82C7E53B7469EE422C4FB3ADA739">2017</mods:date>
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<mods:detail id="36AC1545D27C2E7E8F61BD2490F5464D" type="volume">
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<mods:number id="C5A30721C908C74764EB103EAAFA6D87">4236</mods:number>
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<mods:detail id="6503207163B4B59922D4434A9B6ED09B" type="issue">
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<mods:number id="416F9E6FDE8022B2D2EFA42C8A947BDB">1</mods:number>
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<mods:identifier id="C492E0A0FA347B600D93903498D0F585" type="CLB-Dataset">36479</mods:identifier>
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<mods:identifier id="CA23865AE8B8BF93852162D57761C813" type="DOI">10.11646/zootaxa.4236.1.2</mods:identifier>
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<mods:identifier id="EA59A69750345C4047A87A3C3EC07A4F" type="GBIF-Dataset">0f39b5c0-0af5-476f-8448-a02339cc1dfd</mods:identifier>
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<mods:identifier id="738E3FE774D53E556DFDC72E22827C62" type="ISSN">1175-5326</mods:identifier>
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<treatment id="4F3A5135FFB0DC084FE3FD146E5BFE82" ID-DOI="http://doi.org/10.5281/zenodo.6015703" ID-GBIF-Taxon="127697125" ID-Zenodo-Dep="6015703" LSID="urn:lsid:plazi:treatment:4F3A5135FFB0DC084FE3FD146E5BFE82" httpUri="http://treatment.plazi.org/id/4F3A5135FFB0DC084FE3FD146E5BFE82" lastPageId="11" lastPageNumber="52" pageId="8" pageNumber="49">
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<subSubSection id="8F89B3A8FFB0DC0B4FE3FD146E71FC8C" pageId="8" pageNumber="49" type="nomenclature">
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<paragraph id="C72CE023FFB0DC0B4FE3FD146DBAFD6D" blockId="8.[151,694,728,787]" box="[151,694,728,754]" pageId="8" pageNumber="49">
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<heading id="9C64574FFFB0DC0B4FE3FD146DBAFD6D" bold="true" box="[151,694,728,754]" fontSize="11" level="1" pageId="8" pageNumber="49" reason="1">
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<emphasis id="F5E73C31FFB0DC0B4FE3FD146DBAFD6D" bold="true" box="[151,694,728,754]" pageId="8" pageNumber="49">
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<taxonomicName id="00939BA0FFB0DC0B4FE3FD146D58FD6D" authority="Anisyutkin" authorityName="Anisyutkin" box="[151,596,728,754]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov.">
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<emphasis id="F5E73C31FFB0DC0B4FE3FD146ECCFD6D" bold="true" box="[151,448,728,754]" italics="true" pageId="8" pageNumber="49">Rhabdoblattella alexeevi</emphasis>
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Anisyutkin
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</taxonomicName>
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,
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<taxonomicNameLabel id="EED4814AFFB0DC0B4D14FD146DBAFD6D" box="[608,694,728,754]" pageId="8" pageNumber="49" rank="species">sp.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C72CE023FFB0DC0B4FE3FD366E71FC8C" blockId="8.[151,694,728,787]" box="[151,381,762,787]" pageId="8" pageNumber="49">
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(
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<figureCitation id="5FA8FCA6FFB0DC0B4FEBFD366E28FC8C" box="[159,292,762,787]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="8" pageNumber="49">Figs. 4A–C</figureCitation>
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,
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<figureCitation id="5FA8FCA6FFB0DC0B4E5BFD366E37FC8C" box="[303,315,762,787]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">5</figureCitation>
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A–N)
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</paragraph>
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</subSubSection>
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<subSubSection id="8F89B3A8FFB0DC0B4FE3FC8E6A8CFCE0" pageId="8" pageNumber="49" type="materials_examined">
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<paragraph id="C72CE023FFB0DC0B4FE3FC8E6A8CFCE0" blockId="8.[151,1437,834,2011]" pageId="8" pageNumber="49">
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<materialsCitation id="77FBEA7EFFB0DC0B4FE3FC8E6A8CFCE0" ID-GBIF-Occurrence="1434656635" collectingDate="1972-11-22" collectionCode="SOUTH, MHNG" collectorName="C. Besuchet & I. Lobl" country="India" county="Cardamon Hills" elevation="1700" location="Munnar" municipality="Muttapatti" pageId="8" pageNumber="49" specimenCount="1" stateProvince="Kerala">
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<emphasis id="F5E73C31FFB0DC0B4FE3FC8E6E0AFCC4" bold="true" box="[151,262,834,859]" pageId="8" pageNumber="49">Material.</emphasis>
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Holotype—male,
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<collectionCode id="A18278E6FFB0DC0B4E94FC8F6D31FCC4" box="[480,573,835,859]" pageId="8" pageNumber="49">SOUTH</collectionCode>
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<collectingCountry id="BF84A0B3FFB0DC0B4D3DFC8F6D90FCC4" box="[585,668,835,859]" name="India" pageId="8" pageNumber="49">INDIA</collectingCountry>
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, state
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<collectingRegion id="05572EC1FFB0DC0B4D9FFC8E6C34FCC4" box="[747,824,834,859]" country="India" name="Kerala" pageId="8" pageNumber="49">Kerala</collectingRegion>
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,
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<collectingCounty id="2E4D98AFFFB0DC0B4C3DFC8E6B0FFCC4" box="[841,1027,834,859]" pageId="8" pageNumber="49">Cardamon Hills</collectingCounty>
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,
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<collectingMunicipality id="27487A59FFB0DC0B4B67FC8E6B84FCC4" box="[1043,1160,834,859]" pageId="8" pageNumber="49">Muttapatti</collectingMunicipality>
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near
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<location id="C24CB6F8FFB0DC0B4BA5FC8F6A27FCC5" LSID="urn:lsid:plazi:treatment:4F3A5135FFB0DC084FE3FD146E5BFE82:C24CB6F8FFB0DC0B4BA5FC8F6A27FCC5" box="[1233,1323,835,858]" country="India" county="Cardamon Hills" municipality="Muttapatti" name="Munnar" pageId="8" pageNumber="49" stateProvince="Kerala">Munnar</location>
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,
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<elevation id="4CBE0710FFB0DC0B4A4FFC8F6A9BFCC5" box="[1339,1431,835,859]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="49" unit="m" value="1700.0">
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<quantity id="006B4DC6FFB0DC0B4A4FFC8F6A9BFCC5" box="[1339,1431,835,859]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="49" unit="m" value="1700.0">1700 m</quantity>
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</elevation>
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, sieving in forest at the foot of a group of tree ferns,
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<collectingDate id="A3693F0BFFB0DC0B4DAEFCAB6CB9FCE0" box="[730,949,870,895]" pageId="8" pageNumber="49" value="1972-11-22">
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||
<date id="B32DC6E3FFB0DC0B4DAEFCAB6CB9FCE0" box="[730,949,870,895]" pageId="8" pageNumber="49" value="1972-11-22">22 November 1972</date>
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</collectingDate>
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, coll.
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<collectorName id="6A6685F5FFB0DC0B4C8CFCAB6B8FFCE0" box="[1016,1155,870,895]" pageId="8" pageNumber="49">C. Besuchet</collectorName>
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/
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<collectorName id="6A6685F5FFB0DC0B4BF8FCAA6BD8FCE0" box="[1164,1236,870,895]" pageId="8" pageNumber="49">I. Löbl</collectorName>
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(48) (
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<collectionCode id="A18278E6FFB0DC0B4A6CFCAB6A78FCE0" box="[1304,1396,871,895]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="49">MHNG</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="8F89B3A8FFB0DC0B4FB3FC476E55FC74" pageId="8" pageNumber="49" type="etymology">
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<paragraph id="C72CE023FFB0DC0B4FB3FC476E55FC74" blockId="8.[151,1437,834,2011]" pageId="8" pageNumber="49">
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<emphasis id="F5E73C31FFB0DC0B4FB3FC476E42FC3C" bold="true" box="[199,334,907,931]" pageId="8" pageNumber="49">Etymology.</emphasis>
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This species is named in honour of the late Prof. Dr. Andrey N. Alexeev (
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<date id="B32DC6E3FFB0DC0B4BE9FC466A2DFC3C" box="[1181,1313,906,931]" pageId="8" pageNumber="49" value="1930" valueMax="2015">1930–2015</date>
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), a famous parasitologist and a true Russian intellectual, under whose supervision the authors of the present paper had a pleasure to work.
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</paragraph>
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</subSubSection>
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<subSubSection id="8F89B3A8FFB0DC084FB3FC3A6E5BFE82" lastPageId="11" lastPageNumber="52" pageId="8" pageNumber="49" type="description">
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<paragraph id="C72CE023FFB0DC0B4FB3FC3A6CA1F924" blockId="8.[151,1437,834,2011]" pageId="8" pageNumber="49">
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<emphasis id="F5E73C31FFB0DC0B4FB3FC3A6E55FB90" bold="true" box="[199,345,1014,1039]" pageId="8" pageNumber="49">Description.</emphasis>
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Male (the
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<typeStatus id="18285E81FFB0DC0B4EA0FC3A6D37FB90" box="[468,571,1014,1039]" pageId="8" pageNumber="49" type="holotype">holotype</typeStatus>
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). General colour reddish brown (
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<figureCitation id="5FA8FCA6FFB0DC0B4CDAFC3A6B26FB90" box="[942,1066,1014,1039]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="8" pageNumber="49">Fig. 4A–C</figureCitation>
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); facial part of head brown; eyes black; pronotum, tegmina and abdomen densely speckled, with small dark maculae; legs dirty yellowish. Surfaces smooth and lustrous. Head convex (
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<figureCitation id="5FA8FCA6FFB0DC0B4D53FBF26D85FBC8" box="[551,649,1086,1111]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="8" pageNumber="49">Figs. 4C</figureCitation>
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,
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<figureCitation id="5FA8FCA6FFB0DC0B4DE0FBF26DAEFBC8" box="[660,674,1086,1111]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">5</figureCitation>
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A); epicranial sutures present; ocellar spots small, but distinct; weak transverse hollow located between antennal sockets; distance between eyes 1.2 times eye length; distance between antennal sockets 2.5 times of the scape length (about
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<quantity id="006B4DC6FFB0DC0B4C74FB4B6C50FB00" box="[768,860,1159,1183]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="49" unit="mm" value="0.4">0.4 mm</quantity>
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); approximate length ratio of 3rd–5th segments of maxillary palps 1.3: 1.0: 1.8. Pronotum transverse; widely rounded along anterolateral margins, caudal margin very weakly angulate (
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<figureCitation id="5FA8FCA6FFB0DC0B4ED7FB026D3CFB79" box="[419,560,1230,1255]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="8" pageNumber="49">Figs. 4A, B</figureCitation>
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,
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<figureCitation id="5FA8FCA6FFB0DC0B4D4BFB026D42FB78" box="[575,590,1230,1255]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">5</figureCitation>
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B). Scutellum very small. Tegmina and wings shortened (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4B80FB026A5EFB78" box="[1268,1362,1230,1255]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="8" pageNumber="49">Fig. 4A</figureCitation>
|
||
), only reaching 7th abdominal tergite; rounded apically; venation distinct; all main veins present;
|
||
<emphasis id="F5E73C31FFB0DC0B4BB1FB3F6BD3FA95" box="[1221,1247,1267,1290]" italics="true" pageId="8" pageNumber="49">Sc</emphasis>
|
||
thickened (well visible on ventral side of tegmen). Wings completely hidden under tegmina, much shorter than tegmina, with a simplified venation. Anterior margin of fore femur armed as in the
|
||
<typeStatus id="18285E81FFB0DC0B4CE5FAF76CCDFACC" box="[913,961,1339,1363]" pageId="8" pageNumber="49">type</typeStatus>
|
||
B, with 6 spines, including 2 apical ones. Fore tibiae not thickened distally. Structure of hind tarsi as described for the genus (see above); arolium vestigial. Abdominal tergites 5th and 6th specialized (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4DF1FA4E6DCBFA04" box="[645,711,1410,1435]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
C). 5th tergite with median membranous area and thin transverse sclerotized strip located in the centre of this area (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4D84FA6A6C35FA20" box="[752,825,1446,1471]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
,
|
||
<emphasis id="F5E73C31FFB0DC0B4C3EFA6B6C72FA21" box="[842,894,1447,1470]" italics="true" pageId="8" pageNumber="49">s.str.</emphasis>
|
||
). 6th tergite with larger membranous area, as compared with 5th tergite, membranous area separated by transverse sclerotized strip into two parts (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4A73FA066A40FA7C" box="[1287,1356,1482,1507]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
,
|
||
<emphasis id="F5E73C31FFB0DC0B4A2CFA006A80FA7C" box="[1368,1420,1484,1507]" italics="true" pageId="8" pageNumber="49">s.str.</emphasis>
|
||
), anterior part with membranous tubercle and small transverse sclerotized plate at base of tubercle. Posterolateral angles of tergites obtuse (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4EB4F9DE6D0AF9B4" box="[448,518,1554,1579]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
C). Anal plate (tergite X) with the widely rounded caudal margin and a distinct median incision (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4E2CF9FA6E96F9D0" box="[344,410,1590,1615]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
D, E). Cerci short, flatten, with segments distinctly separated (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4B26F9FA6B99F9D0" box="[1106,1173,1590,1615]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
D, E). Paraprocts of the blaberid-type, right paraproct with a sclerotized outgrowth directed cranially (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4B4DF9966B8EF9EC" box="[1081,1154,1626,1651]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
E). Hypandrium weakly asymmetrical, with caudal margin rounded and projected caudally (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4CD7F9B26CE5F908" box="[931,1001,1662,1687]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
F); a median tooth present (Fig. F, G,
|
||
<emphasis id="F5E73C31FFB0DC0B4FE3F9686FCDF924" box="[151,193,1700,1723]" italics="true" pageId="8" pageNumber="49">m.t.</emphasis>
|
||
); right stylus fusiform, bent at its base (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4DF4F96E6DCFF924" box="[640,707,1698,1723]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
F), left stylus absent.
|
||
</paragraph>
|
||
<paragraph id="C72CE023FFB0DC0B4FB3F90A6D22F828" blockId="8.[151,1437,834,2011]" pageId="8" pageNumber="49">
|
||
Male genitalia (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4E0EF90A6EB1F940" box="[378,445,1734,1759]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
H–N). Right phallomere (R+N) with caudal part of sclerite R1T transverse (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4A50F90A6A64F940" box="[1316,1384,1734,1759]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
H, I,
|
||
<emphasis id="F5E73C31FFB0DC0B4FE3F9276FE2F89C" box="[151,238,1771,1795]" italics="true" pageId="8" pageNumber="49">c.p.R1T</emphasis>
|
||
), cranial part of R1T and R2 rounded (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4DD5F9266DEFF89C" box="[673,739,1770,1795]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
H, I,
|
||
<emphasis id="F5E73C31FFB0DC0B4C6FF9276C76F89C" box="[795,890,1771,1795]" italics="true" pageId="8" pageNumber="49">cr.p.R1T</emphasis>
|
||
); R3 with branches of unequal length and width; R5 wide, weakly sclerotized; bristles absent. Sclerite L2D (L1) divided into basal and apical parts (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4B8FF8C26A4CF8B8" box="[1275,1344,1806,1831]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
J); basal part rod-like, moderately widened cranially (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4DE0F8FE6DD4F8D4" box="[660,728,1842,1867]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
J,
|
||
<emphasis id="F5E73C31FFB0DC0B4D87F8FE6C35F8D4" box="[755,825,1842,1867]" italics="true" pageId="8" pageNumber="49">b.L2D</emphasis>
|
||
); caudally with "curved upward subsclerite" (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4A32F8FE6A86F8D4" box="[1350,1418,1842,1867]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
J, K,
|
||
<emphasis id="F5E73C31FFB0DC0B4FC8F89B6FEEF8F1" box="[188,226,1879,1902]" italics="true" pageId="8" pageNumber="49">u.s.</emphasis>
|
||
); apical part in shape of sclerotized and folded plate (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4C23F89A6C90F8F0" box="[855,924,1878,1903]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
J, K); bristles absent. Sclerite L3 (L2d) with distinct basal subsclerite (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4EC8F8B66EF3F80C" box="[444,511,1914,1939]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
L–N,
|
||
<emphasis id="F5E73C31FFB0DC0B4D36F8B66D78F80C" box="[578,628,1914,1939]" italics="true" pageId="8" pageNumber="49">b.L3</emphasis>
|
||
); "folded structure" distinct, with bristles (
|
||
<figureCitation id="5FA8FCA6FFB0DC0B4B29F8B66BACF80C" box="[1117,1184,1914,1939]" captionStart="FIGURE 5" captionStartId="10.[151,249,1831,1853]" captionTargetBox="[158,1422,205,1795]" captionTargetId="figure@10.[151,1436,193,1810]" captionTargetPageId="10" captionText="FIGURE 5. Rhabdoblattella alexeevi sp. nov. (male, holotype). A—facial part of head; B—pronotum and head from above; C—IV – VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L – Nsclerite L 3. Dotted area shows membranous parts. Abbreviations: IƑ – ƑIII — 4 th – 8 th abdominal tergites respectively, b. L 2 D, b. L 3, c. p. R 1 T, cr. p. R 1 T, f. s., m. t., R 2, R 3, R 5, u. s. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L – N." httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="8" pageNumber="49">Fig. 5</figureCitation>
|
||
L–N,
|
||
<emphasis id="F5E73C31FFB0DC0B4B97F8B76A09F80C" box="[1251,1285,1915,1939]" italics="true" pageId="8" pageNumber="49">f.s.</emphasis>
|
||
); groove hge present. Sclerite L4U (L3d) distinct.
|
||
</paragraph>
|
||
<paragraph id="C72CE023FFB0DC0B4FB3F80E6E9CF844" blockId="8.[151,1437,834,2011]" box="[199,400,1986,2011]" pageId="8" pageNumber="49">Female unknown.</paragraph>
|
||
<paragraph id="C72CE023FFB1DC0A4FB3FF546FCDFF4B" blockId="9.[151,1436,151,392]" pageId="9" pageNumber="50">
|
||
<emphasis id="F5E73C31FFB1DC0A4FB3FF546E7AFF2F" bold="true" box="[199,374,152,176]" pageId="9" pageNumber="50">Measurements</emphasis>
|
||
(mm). Head length: 1.9, width 1.9, pronotum length 2.5, width 3.7, tegmina length 4.8, width 2.3.
|
||
</paragraph>
|
||
<paragraph id="C72CE023FFB1DC0A4FB3FF136D19FE17" blockId="9.[151,1436,151,392]" pageId="9" pageNumber="50">
|
||
<emphasis id="F5E73C31FFB1DC0A4FB3FF136E6DFF67" bold="true" box="[199,353,223,248]" pageId="9" pageNumber="50">Comparison.</emphasis>
|
||
The new species readily differs from all its congeners in having the distinctly shortened tegmina and wings.
|
||
<taxonomicName id="00939BA0FFB1DC0A4E6CFEC96E81FE82" box="[280,397,260,285]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFB1DC0A4E6CFEC96E81FE82" box="[280,397,260,285]" italics="true" pageId="9" pageNumber="50">R alexeevi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EED4814AFFB1DC0A4EEFFEC86EE1FE83" box="[411,493,260,284]" pageId="9" pageNumber="50" rank="species">
|
||
<emphasis id="F5E73C31FFB1DC0A4EEFFEC86EE1FE83" bold="true" box="[411,493,260,284]" pageId="9" pageNumber="50">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="00939BA0FFB1DC0A4DF5FEC96DFAFE83" box="[641,758,261,284]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="euptera" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFB1DC0A4DF5FEC96DFAFE83" box="[641,758,261,284]" italics="true" pageId="9" pageNumber="50">R. euptera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EED4814AFFB1DC0A4C71FEC86C5BFE83" box="[773,855,260,284]" pageId="9" pageNumber="50" rank="species">
|
||
<emphasis id="F5E73C31FFB1DC0A4C71FEC86C5BFE83" bold="true" box="[773,855,260,284]" pageId="9" pageNumber="50">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in the presence of abdominal tergal glands, but can be readily distinguished from it by the location and structure of these glands (on the 5th and 6th tergites in
|
||
<taxonomicName id="00939BA0FFB1DC0A4AF0FEE56FFFFEFA" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFB1DC0A4AF0FEE56FFFFEFA" italics="true" pageId="9" pageNumber="50">R. alexeevi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EED4814AFFB1DC0A4E77FE806E59FEFB" box="[259,341,332,356]" pageId="9" pageNumber="50" rank="species">
|
||
<emphasis id="F5E73C31FFB1DC0A4E77FE806E59FEFB" bold="true" box="[259,341,332,356]" pageId="9" pageNumber="50">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, contrary to the 6th and 7th ones in
|
||
<taxonomicName id="00939BA0FFB1DC0A4D89FE816C7FFEFB" box="[765,883,333,356]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="euptera" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFB1DC0A4D89FE816C7FFEFB" box="[765,883,333,356]" italics="true" pageId="9" pageNumber="50">R. euptera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EED4814AFFB1DC0A4CF0FE806CDAFEFB" box="[900,982,332,356]" pageId="9" pageNumber="50" rank="species">
|
||
<emphasis id="F5E73C31FFB1DC0A4CF0FE806CDAFEFB" bold="true" box="[900,982,332,356]" pageId="9" pageNumber="50">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
), and the presence of the apical part of sclerite L2D in the male genitalia.
|
||
</paragraph>
|
||
<caption id="93ECB0ABFFB1DC0A4FE3F8BA6A81F855" httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="9" pageNumber="50" targetBox="[156,1430,440,1893]" targetPageId="9">
|
||
<paragraph id="C72CE023FFB1DC0A4FE3F8BA6A81F855" blockId="9.[151,1436,1910,1994]" pageId="9" pageNumber="50">
|
||
<emphasis id="F5E73C31FFB1DC0A4FE3F8BA6E18F813" bold="true" box="[151,276,1910,1932]" pageId="9" pageNumber="50">FIGURE 4.</emphasis>
|
||
A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A–C—
|
||
<taxonomicName id="00939BA0FFB1DC0A4D4BF8596C31F834" box="[575,829,1941,1963]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFB1DC0A4D4BF8596C31F834" box="[575,829,1941,1963]" italics="true" pageId="9" pageNumber="50">Rhabdoblattella alexeevi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFB1DC0A4C31F8596C83F834" box="[837,911,1941,1963]" pageId="9" pageNumber="50" rank="species">
|
||
<emphasis id="F5E73C31FFB1DC0A4C31F8596C83F834" bold="true" box="[837,911,1941,1963]" pageId="9" pageNumber="50">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(male, holotype); D, E—
|
||
<taxonomicName id="00939BA0FFB1DC0A4BE3F85A6A0EF834" box="[1175,1282,1942,1963]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="euptera" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFB1DC0A4BE3F85A6A0EF834" box="[1175,1282,1942,1963]" italics="true" pageId="9" pageNumber="50">R. euptera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFB1DC0A4A7EF8596A59F834" box="[1290,1365,1941,1963]" pageId="9" pageNumber="50" rank="species">
|
||
<emphasis id="F5E73C31FFB1DC0A4A7EF8596A59F834" bold="true" box="[1290,1365,1941,1963]" pageId="9" pageNumber="50">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(male, holotype); F, G—
|
||
<taxonomicName id="00939BA0FFB1DC0A4E3DF8786D38F855" box="[329,564,1972,1994]" class="Insecta" family="Blaberidae" genus="Placoblatta" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="beybienkoi" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFB1DC0A4E3DF8786D38F855" box="[329,564,1972,1994]" italics="true" pageId="9" pageNumber="50">Placoblatta beybienkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFB1DC0A4D4EF8786D89F855" box="[570,645,1972,1994]" pageId="9" pageNumber="50" rank="species">
|
||
<emphasis id="F5E73C31FFB1DC0A4D4EF8786D89F855" bold="true" box="[570,645,1972,1994]" pageId="9" pageNumber="50">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="93ECB0ABFFB2DC094FE3F8EB6EAFF86A" httpUri="https://zenodo.org/record/322019/files/figure.png" pageId="10" pageNumber="51" targetBox="[158,1422,205,1795]" targetPageId="10">
|
||
<paragraph id="C72CE023FFB2DC094FE3F8EB6EAFF86A" blockId="10.[151,1436,1831,2037]" pageId="10" pageNumber="51">
|
||
<emphasis id="F5E73C31FFB2DC094FE3F8EB6E18F8A2" bold="true" box="[151,276,1831,1853]" pageId="10" pageNumber="51">FIGURE 5.</emphasis>
|
||
<taxonomicName id="00939BA0FFB2DC094E68F8EB6D17F8A2" box="[284,539,1831,1853]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFB2DC094E68F8EB6D17F8A2" box="[284,539,1831,1853]" italics="true" pageId="10" pageNumber="51">Rhabdoblattella alexeevi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EED4814AFFB2DC094D57F8EB6D61F8A2" box="[547,621,1831,1853]" pageId="10" pageNumber="51" rank="species">
|
||
<emphasis id="F5E73C31FFB2DC094D57F8EB6D61F8A2" bold="true" box="[547,621,1831,1853]" pageId="10" pageNumber="51">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(male, holotype). A—facial part of head; B—pronotum and head from above; C—IV–VIII abdominal tergites from above; D—anal plate and cerci from above; E—abdominal apex from below, hypandrium and genitalia removed; F—hypandrium from below; G—caudal margin of hypandrium from caudal, outlines; H, I—right phallomere from above (H) and below (I); J—sclerite L2D from above; K—apical part of sclerite L2D from above; L–Nsclerite L3. Dotted area shows membranous parts. Abbreviations:
|
||
<emphasis id="F5E73C31FFB2DC094C3EF86F6C9BF827" box="[842,919,1955,1976]" italics="true" pageId="10" pageNumber="51">IƑ–ƑIII</emphasis>
|
||
—4th–8th abdominal tergites respectively,
|
||
<emphasis id="F5E73C31FFB2DC094A21F86E6A9AF827" box="[1365,1430,1954,1976]" italics="true" pageId="10" pageNumber="51">b.L2D</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFB2DC094FE3F80D6FC8F848" box="[151,196,1985,2007]" italics="true" pageId="10" pageNumber="51">b.L3</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFB2DC094FA5F80D6E11F848" box="[209,285,1985,2007]" italics="true" pageId="10" pageNumber="51">c.p.R1T</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFB2DC094E5EF80D6E72F848" box="[298,382,1985,2007]" italics="true" pageId="10" pageNumber="51">cr.p.R1T</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFB2DC094EFFF80D6EA5F848" box="[395,425,1985,2007]" italics="true" pageId="10" pageNumber="51">f.s.</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFB2DC094EC0F80E6ED7F848" box="[436,475,1986,2007]" italics="true" pageId="10" pageNumber="51">m.t.</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFB2DC094E93F80D6D0FF848" box="[487,515,1985,2007]" italics="true" pageId="10" pageNumber="51">R2</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFB2DC094D7BF80D6D27F848" box="[527,555,1985,2007]" italics="true" pageId="10" pageNumber="51">R3</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFB2DC094D4CF80E6D5FF848" box="[568,595,1986,2007]" italics="true" pageId="10" pageNumber="51">R5</emphasis>
|
||
,
|
||
<emphasis id="F5E73C31FFB2DC094D14F80E6D8FF848" box="[608,643,1986,2007]" italics="true" pageId="10" pageNumber="51">u.s.</emphasis>
|
||
—see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G; h = H, I; i = J; j = K; k = L–N.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C72CE023FFB3DC084FB3FF546E5BFE82" blockId="11.[151,1436,151,285]" pageId="11" pageNumber="52">
|
||
<emphasis id="F5E73C31FFB3DC084FB3FF546E0AFF2F" bold="true" box="[199,262,152,176]" pageId="11" pageNumber="52">Note.</emphasis>
|
||
The shortening of tegmina, the reduction of ocelli and arolia are probably evidence of a shift to a more reclusive lifestyle. The well visible epicranial sutures and the reduced ocelli are rather larval characters, i.e. the modus of evolution
|
||
<taxonomicName id="00939BA0FFB3DC084EFFFF2D6D00FF67" box="[395,524,223,248]" class="Insecta" family="Blaberidae" genus="Rhabdoblattella" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="52" phylum="Arthropoda" rank="species" species="alexeevi" status="sp. nov.">
|
||
<emphasis id="F5E73C31FFB3DC084EFFFF2D6D00FF67" box="[395,524,223,248]" italics="true" pageId="11" pageNumber="52">R. alexeevi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="EED4814AFFB3DC084D55FF2C6D7FFF67" box="[545,627,224,248]" pageId="11" pageNumber="52" rank="species">
|
||
<emphasis id="F5E73C31FFB3DC084D55FF2C6D7FFF67" bold="true" box="[545,627,224,248]" pageId="11" pageNumber="52">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
seems to be the retardational paedomorphosis (retrogenesis) (
|
||
<emphasis id="F5E73C31FFB3DC084A2AFF2D6A90FF67" box="[1374,1436,225,248]" italics="true" pageId="11" pageNumber="52">sensu</emphasis>
|
||
<bibRefCitation id="A3029DD2FFB3DC084FE3FEC86E47FE82" author="Iordansky" box="[151,331,260,285]" pageId="11" pageNumber="52" refString="Iordansky, N. N. (2005) Paedomorphosis, neoteny, and evolution. Zoologicheskiy Zhurnal, 84 (10), 1176 - 1187." type="journal article" year="2005">Iordansky 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |