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<document id="F5BBAF24D405B8401B54CC9C662E5A9F" ID-DOI="10.11646/phytotaxa.392.1.6" ID-ISSN="1179-3163" ID-Zenodo-Dep="13718043" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725653160491" checkinUser="felipe" docAuthor="Trujillo, Cristina, Bonifacino, José M. &amp; Pasini, Eduardo" docDate="2019" docId="324387F2FFDC5260FF68FF05D91D278C" docLanguage="en" docName="phytotaxa.392.1.6.pdf" docOrigin="Phytotaxa 392 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.392.1.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Pamphalea tenuissima Trujillo, Bonifacino &amp; Pasini 2019, sp. nov." docType="treatment" docVersion="3" lastPageNumber="65" masterDocId="CE7AFF8AFFDD5264FFE0FFF1DA5A253C" masterDocTitle="Pamphalea tenuissima (Nassauvieae, Asteraceae), a new species from southern Brazil" masterLastPageNumber="66" masterPageNumber="61" pageNumber="62" updateTime="1725663809542" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="68C202D4455C41FAF2FDACA895A15F75">Pamphalea tenuissima (Nassauvieae, Asteraceae), a new species from southern Brazil</mods:title>
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<mods:name id="E2DBD301C7611354ABD60410F2A80373" type="personal">
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<mods:namePart id="DB76DA6CA19E2D71CCD3B9947B5CF779">Trujillo, Cristina</mods:namePart>
<mods:affiliation id="3CA6070CF02343B57A8342F42326E876">Laboratorio de Botánica, Facultad de Agronomía, Universidad de la República, Montevideo, Uruguay.</mods:affiliation>
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<mods:namePart id="B48145123E2811DA92401B7CE477D999">Bonifacino, José M.</mods:namePart>
<mods:affiliation id="68BAF05C0FEA283A965D7BDF6FFAB040">Laboratorio de Botánica, Facultad de Agronomía, Universidad de la República, Montevideo, Uruguay.</mods:affiliation>
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</mods:role>
<mods:namePart id="FB029CD14F1BE19A25CC322227C76346">Pasini, Eduardo</mods:namePart>
<mods:affiliation id="2F9606FA643A134588C55B6BCC5DE702">Garden Engenharia, Caxias do Sul, Rio Grande do Sul, Brazil</mods:affiliation>
</mods:name>
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<mods:title id="A14A10A7FA6C151148FF5E0DD05622A0">Phytotaxa</mods:title>
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<mods:date id="CAE0F2F60EE4EE6F8C1D768C55942462">2019</mods:date>
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<mods:number id="67E41DB41405B9214B78BDBB82CBE54F">2019-02-12</mods:number>
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<mods:identifier id="D27F21A6095A49946B5CAEC6F5A575B1" type="ISSN">1179-3163</mods:identifier>
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<treatment id="324387F2FFDC5260FF68FF05D91D278C" ID-DOI="http://doi.org/10.5281/zenodo.13718288" ID-Zenodo-Dep="13718288" LSID="urn:lsid:plazi:treatment:324387F2FFDC5260FF68FF05D91D278C" httpUri="http://treatment.plazi.org/id/324387F2FFDC5260FF68FF05D91D278C" lastPageId="4" lastPageNumber="65" pageId="1" pageNumber="62">
<subSubSection id="F2F0656FFFDC5265FF68FF05D9982433" box="[136,962,244,271]" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="BA5536E4FFDC5265FF68FF05D9982433" blockId="1.[136,962,244,271]" box="[136,962,244,271]" pageId="1" pageNumber="62">
<heading id="E11D8188FFDC5265FF68FF05D9982433" box="[136,962,244,271]" fontSize="11" level="2" pageId="1" pageNumber="62" reason="3">
<taxonomicName id="7DEA4D67FFDC5265FF68FF05D8852433" ID-CoL="4C8R5" authority="Trujillo, Bonifacino &amp; Pasini" authorityName="Trujillo, Bonifacino &amp; Pasini" authorityYear="2019" box="[136,735,244,271]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="62" phylum="Tracheophyta" rank="species" species="tenuissima" status="sp. nov.">
<emphasis id="889EEAF6FFDC5265FF68FF05DBD52432" bold="true" box="[136,399,244,271]" italics="true" pageId="1" pageNumber="62">Pamphalea tenuissima</emphasis>
Trujillo, Bonifacino &amp; Pasini
</taxonomicName>
,
<emphasis id="889EEAF6FFDC5265FD0AFF05D91A2432" bold="true" box="[746,832,244,270]" italics="true" pageId="1" pageNumber="62">
<taxonomicNameLabel id="93AD578DFFDC5265FD0AFF05D91A2432" box="[746,832,244,270]" pageId="1" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="22D12A61FFDC5265FCB0FF07D9E32433" box="[848,953,246,271]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[136,229,1975,1996]" captionStartId-1="2.[136,229,1490,1511]" captionStartId-2="3.[136,229,1960,1981]" captionTargetBox-0="[237,1291,415,1932]" captionTargetBox-1="[386,1222,204,1428]" captionTargetBox-2="[340,1247,645,1928]" captionTargetId-0="figure-71@1.[221,1366,397,1949]" captionTargetId-1="figure-17@2.[340,1247,190,1466]" captionTargetId-2="figure-225@3.[340,1247,645,1929]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Illustration of Pamphalea tenuissima. A. Habit. B. Leaf from basal rosette, al: apical lobe, ll: lateral lobe. C. Bract. D. Phyllary. E. Floret (achene removed); note glandular trichomes at the base of corolla. F. Stamen (note anther collar and dorsal insertion of filament). G. Style, nd: nectariferous disk. H. Achene. Drawn by Trujillo, based on the holotype (Pasini et al. 1078; MVFA)." captionText-1="FIGURE 2. Compared leaf morphology of Pamphalea cardaminifolia and Pamphalea tenuissima A. Rosette leaf of P. cardaminifolia. B. Rosette leaf of P. tenuissima. C. Cauline leaf of P. cardaminifolia. D. Cauline leaf of P. tenuissima." captionText-2="FIGURE 3. Pamphalea tenuissima. A. Habit. B, C. Cauline leaves. D. Rosette leaf. E, F. Habit, the main rosette not shown in picture. G. Head, lateral view. H. Head, superior view. Photographs: Bonifacino." figureDoi-0="http://doi.org/10.5281/zenodo.13718045" figureDoi-1="http://doi.org/10.5281/zenodo.13718047" figureDoi-2="http://doi.org/10.5281/zenodo.13718049" httpUri-0="https://zenodo.org/record/13718045/files/figure.png" httpUri-1="https://zenodo.org/record/13718047/files/figure.png" httpUri-2="https://zenodo.org/record/13718049/files/figure.png" pageId="1" pageNumber="62">Figs. 1–3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F2F0656FFFDC5260FF68FEEDDEF12450" lastPageId="4" lastPageNumber="65" pageId="1" pageNumber="62" type="materials_examined">
<paragraph id="BA5536E4FFDC5265FF68FEEDDB282445" blockId="1.[136,1452,284,377]" pageId="1" pageNumber="62">
<emphasis id="889EEAF6FFDC5265FF68FEEDDAE0240D" bold="true" box="[136,186,284,305]" pageId="1" pageNumber="62">Type</emphasis>
:—
<collectingCountry id="C2FD7674FFDC5265FF37FEEDDB69240D" box="[215,307,284,305]" name="Brazil" pageId="1" pageNumber="62">BRAZIL</collectingCountry>
.
<collectingRegion id="782EF806FFDC5265FEDCFEEDDBA8240D" box="[316,498,284,305]" country="Brazil" name="Rio Grande do Sul" pageId="1" pageNumber="62">Rio Grande do Sul</collectingRegion>
: São José dos Ausentes, distrito Silveira, RS-020, calle asfaltada, camino a Silveira, lado izquierdo, borde alambrado, sitio encharcado, con
<taxonomicName id="7DEA4D67FFDC5265FDDCFEB1D89E2469" box="[572,708,320,341]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="1" pageNumber="62" phylum="Bryophyta" rank="species" species="undetermined">
<emphasis id="889EEAF6FFDC5265FDDCFEB1D8FB2469" box="[572,673,320,341]" italics="true" pageId="1" pageNumber="62">Sphagnum</emphasis>
sp.
</taxonomicName>
, Lat. 28˚42’28.9”
<collectionCode id="DCFBAE21FFDC5265FC91FEB1D9242469" box="[881,894,320,341]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="1" pageNumber="62" type="Herbarium">S</collectionCode>
, Long. 50˚0’2.6”
<collectionCode id="DCFBAE21FFDC5265FBC0FEB1DE632469" box="[1056,1081,320,341]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="62" type="Herbarium">W</collectionCode>
,
<date id="CE541024FFDC5265FBA3FEB1DEA12469" box="[1091,1275,320,341]" pageId="1" pageNumber="62" value="2015-12-10">10 December 2015</date>
, Pasini
<emphasis id="889EEAF6FFDC5265FAA6FEB1DF2C2469" box="[1350,1398,320,341]" italics="true" pageId="1" pageNumber="62">et al.</emphasis>
1078 (
<typeStatus id="65518846FFDC5265FF24FE95DB402445" box="[196,282,356,377]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
<collectionCode id="DCFBAE21FFDC5265FEFFFE95DB3D2445" box="[287,359,356,377]" country="Uruguay" httpUri="http://grbio.org/cool/x4p2-8dvd" name="Universidad de la República" pageId="1" pageNumber="62" type="Herbarium">MVFA</collectionCode>
).
</paragraph>
<caption id="EE95666CFFDC5265FF68F846DF632D28" ID-DOI="http://doi.org/10.5281/zenodo.13718045" ID-Zenodo-Dep="13718045" httpUri="https://zenodo.org/record/13718045/files/figure.png" pageId="1" pageNumber="62" startId="1.[136,229,1975,1996]" targetBox="[237,1291,415,1932]" targetPageId="1" targetType="figure">
<paragraph id="BA5536E4FFDC5265FF68F846DF632D28" blockId="1.[136,1452,1975,2068]" pageId="1" pageNumber="62">
<emphasis id="889EEAF6FFDC5265FF68F846DB5B22F0" bold="true" box="[136,257,1975,1996]" pageId="1" pageNumber="62">FIGURE 1.</emphasis>
Illustration of
<taxonomicName id="7DEA4D67FFDC5265FE78F846D82F22F0" authorityName="Trujillo, Bonifacino &amp; Pasini" authorityYear="2019" box="[408,629,1975,1996]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="62" phylum="Tracheophyta" rank="species" species="tenuissima">
<emphasis id="889EEAF6FFDC5265FE78F846D82F22F0" box="[408,629,1975,1996]" italics="true" pageId="1" pageNumber="62">Pamphalea tenuissima</emphasis>
</taxonomicName>
.
<emphasis id="889EEAF6FFDC5265FD64F846D8C122F0" bold="true" box="[644,667,1975,1996]" pageId="1" pageNumber="62">A.</emphasis>
Habit.
<emphasis id="889EEAF6FFDC5265FD08F846D8A422F0" bold="true" box="[744,766,1975,1996]" pageId="1" pageNumber="62">B.</emphasis>
Leaf from basal rosette, al: apical lobe, ll: lateral lobe.
<emphasis id="889EEAF6FFDC5265FAD1F846DF1222F0" bold="true" box="[1329,1352,1975,1996]" pageId="1" pageNumber="62">C.</emphasis>
Bract.
<emphasis id="889EEAF6FFDC5265FA74F846DFF622F0" bold="true" box="[1428,1452,1975,1996]" pageId="1" pageNumber="62">D.</emphasis>
Phyllary.
<emphasis id="889EEAF6FFDC5265FF03F82ADAA022CC" bold="true" box="[227,250,2011,2032]" pageId="1" pageNumber="62">E.</emphasis>
Floret (achene removed); note glandular trichomes at the base of corolla.
<emphasis id="889EEAF6FFDC5265FC5BF82AD99722CC" bold="true" box="[955,973,2011,2032]" pageId="1" pageNumber="62">F.</emphasis>
Stamen (note anther collar and dorsal insertion of filament).
<emphasis id="889EEAF6FFDC5265FF0CF80EDB5F2D28" bold="true" box="[236,261,2047,2068]" pageId="1" pageNumber="62">G.</emphasis>
Style, nd: nectariferous disk.
<emphasis id="889EEAF6FFDC5265FDC4F80ED8672D28" bold="true" box="[548,573,2047,2068]" pageId="1" pageNumber="62">H.</emphasis>
Achene. Drawn by Trujillo, based on the holotype (
<emphasis id="889EEAF6FFDC5265FBD0F80EDE832D28" box="[1072,1241,2047,2068]" italics="true" pageId="1" pageNumber="62">Pasini et al. 1078</emphasis>
; MVFA).
</paragraph>
</caption>
<caption id="EE95666CFFDF5266FF68FA23DE652337" ID-DOI="http://doi.org/10.5281/zenodo.13718047" ID-Zenodo-Dep="13718047" httpUri="https://zenodo.org/record/13718047/files/figure.png" pageId="2" pageNumber="63" startId="2.[136,229,1490,1511]" targetBox="[386,1222,204,1428]" targetPageId="2" targetType="figure">
<paragraph id="BA5536E4FFDF5266FF68FA23DE652337" blockId="2.[136,1452,1490,1547]" pageId="2" pageNumber="63">
<emphasis id="889EEAF6FFDF5266FF68FA23DAA720DB" bold="true" box="[136,253,1490,1511]" pageId="2" pageNumber="63">FIGURE 2.</emphasis>
Compared leaf morphology of
<taxonomicName id="7DEA4D67FFDF5266FDCAFA23D97120DB" authorityName="Lessing" authorityYear="1830" box="[554,811,1490,1511]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="cardaminifolia">
<emphasis id="889EEAF6FFDF5266FDCAFA23D97120DB" box="[554,811,1490,1511]" italics="true" pageId="2" pageNumber="63">Pamphalea cardaminifolia</emphasis>
</taxonomicName>
and
<taxonomicName id="7DEA4D67FFDF5266FCB8FA23DEC620DB" authority="A. Rosette" authorityName="A. Rosette" box="[856,1180,1490,1511]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="tenuissima">
<emphasis id="889EEAF6FFDF5266FCB8FA23DE6820DB" box="[856,1074,1490,1511]" italics="true" pageId="2" pageNumber="63">Pamphalea tenuissima</emphasis>
<emphasis id="889EEAF6FFDF5266FBD7FA23DE1520DB" bold="true" box="[1079,1103,1490,1511]" pageId="2" pageNumber="63">A.</emphasis>
Rosette
</taxonomicName>
leaf of
<taxonomicName id="7DEA4D67FFDF5266FB05FA23DFD020DB" authorityName="Lessing" authorityYear="1830" box="[1253,1418,1490,1511]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="cardaminifolia">
<emphasis id="889EEAF6FFDF5266FB05FA23DFD020DB" box="[1253,1418,1490,1511]" italics="true" pageId="2" pageNumber="63">P. cardaminifolia</emphasis>
</taxonomicName>
.
<emphasis id="889EEAF6FFDF5266FA75FA23DFF620DB" bold="true" box="[1429,1452,1490,1511]" pageId="2" pageNumber="63">B.</emphasis>
Rosette leaf of
<emphasis id="889EEAF6FFDF5266FEFAFA07DBC52337" box="[282,415,1526,1547]" italics="true" pageId="2" pageNumber="63">
<taxonomicName id="7DEA4D67FFDF5266FEFAFA07DBC62337" box="[282,412,1526,1547]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="tenuissima">P. tenuissima</taxonomicName>
.
</emphasis>
<emphasis id="889EEAF6FFDF5266FE44FA07DBE62337" bold="true" box="[420,444,1526,1547]" pageId="2" pageNumber="63">C.</emphasis>
Cauline leaf of
<emphasis id="889EEAF6FFDF5266FDB6FA07D9592337" box="[598,771,1526,1547]" italics="true" pageId="2" pageNumber="63">
<taxonomicName id="7DEA4D67FFDF5266FDB6FA07D8A52337" authorityName="Lessing" authorityYear="1830" box="[598,767,1526,1547]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="cardaminifolia">P. cardaminifolia</taxonomicName>
.
</emphasis>
<emphasis id="889EEAF6FFDF5266FCE8FA07D9452337" bold="true" box="[776,799,1526,1547]" pageId="2" pageNumber="63">D.</emphasis>
Cauline leaf of
<taxonomicName id="7DEA4D67FFDF5266FC59FA07DE612337" box="[953,1083,1526,1547]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="tenuissima">P. tenuissima</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="BA5536E4FFDF5267FF68F9CDDE922419" blockId="2.[136,1452,1596,2089]" lastBlockId="3.[136,1452,160,617]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
Rosulate herbs
<quantity id="7D129B01FFDF5266FED4F9CDDBE22369" box="[308,440,1596,1621]" metricMagnitude="-1" metricUnit="m" metricValue="3.775" metricValueMax="6.0" metricValueMin="1.55" pageId="2" pageNumber="63" unit="cm" value="37.75" valueMax="60.0" valueMin="15.5">15.5–60 cm</quantity>
high; stems decumbent, forming secondary rosettes and adventitious roots on basal nodes, sometimes main stems erect and secondary decumbent, internodes
<quantity id="7D129B01FFDF5266FC6DF991DE572345" box="[909,1037,1632,1657]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="8.5" metricValueMin="0.3" pageId="2" pageNumber="63" unit="cm" value="4.4" valueMax="8.5" valueMin="0.3">0.3–8.5 cm</quantity>
long, glandular trichomes sparse to absent. Leaves alternate, arranged in a rosette, sessile, 8–20.6 ×
<quantity id="7D129B01FFDF5266FCB8F975D9F523A1" box="[856,943,1668,1693]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="63" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
, pseudopetiole
<quantity id="7D129B01FFDF5266FBBDF975DEE423A1" box="[1117,1214,1668,1693]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="14.0" metricValueMin="2.0" pageId="2" pageNumber="63" unit="cm" value="8.0" valueMax="14.0" valueMin="2.0">2–14 cm</quantity>
long, flat, sheathing, lamina narrowly elliptic to obovate, margin lyrate-pinnatisect, attenuate at the base, apex acute, apical lobe 0.25–1.5 ×
<quantity id="7D129B01FFDF5266FF7FF93DDB4523D9" box="[159,287,1740,1765]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="14.0" metricValueMin="1.0" pageId="2" pageNumber="63" unit="cm" value="0.75" valueMax="1.4" valueMin="0.1">0.1–1.4 cm</quantity>
, ovate to elliptical, 3–5-lobed, deeply incised, lateral lobes (5–)6–12 pairs, 0.2–1.2 ×
<quantity id="7D129B01FFDF5266FB00F93DDF3423D9" box="[1248,1390,1740,1765]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="4.0" metricValueMin="0.5" pageId="2" pageNumber="63" unit="cm" value="0.225" valueMax="0.4" valueMin="0.05">0.05–0.4 cm</quantity>
, very thin, herbaceous, entire to 2 lobed due to a deep incision, 1 pair sometimes 1-dentate, glabrous. Cauline leaves alternate, sessile, 0.2–13.2 ×
<quantity id="7D129B01FFDF5266FEBDF8E5DBB02211" box="[349,490,1812,1837]" metricMagnitude="-2" metricUnit="m" metricValue="1.075" metricValueMax="2.1" metricValueMin="0.05" pageId="2" pageNumber="63" unit="cm" value="1.075" valueMax="2.1" valueMin="0.05">0.05–2.1 cm</quantity>
, similar in shape to basal leaves, semi-amplexicaul, apex acute, margin dentate only at base, with very deep incisions, glabrous, thin and herbaceous. Capitulescences paniculiform with peduncles attached on one side of rachis, heads 2 to 25, peduncles
<quantity id="7D129B01FFDF5266FD48F8ADD94B2249" box="[680,785,1884,1909]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="6.4" metricValueMin="2.0" pageId="2" pageNumber="63" unit="cm" value="4.2" valueMax="6.4" valueMin="2.0">2–6.4 cm</quantity>
long, glandular trichomes very sparse; bracts 2, 0.8–1.5 ×
<quantity id="7D129B01FFDF5266FF68F871DAA522A5" box="[136,255,1920,1945]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="63" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
, ovate, scarious. Involucres 2.5–3.5 ×
<quantity id="7D129B01FFDF5266FD22F871D96322A5" box="[706,825,1920,1945]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="2" pageNumber="63" unit="mm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 mm</quantity>
, cylindrical; phyllaries 8(–10), arranged in one series, imbricate, 2.2–3.4 ×
<quantity id="7D129B01FFDF5266FE96F855D85A2281" box="[374,512,1956,1981]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="2" pageNumber="63" unit="mm" value="1.15" valueMax="1.5" valueMin="0.8">0.8–1.5 mm</quantity>
, elliptic to narrowly ovate, sometimes obovate or oblong, apex mucronate, margin scarious, glandular trichomes present towards the apex. Receptacles convex, glandular trichomes sparse. Florets 12– 20, isomorphic, bisexual, corollas bilabiate, tube
<quantity id="7D129B01FFDF5266FD52F81DD9602D39" box="[690,826,2028,2053]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="2" pageNumber="63" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.3–1.6 mm</quantity>
long, glandular trichomes present mainly in the ventral and lateral areas, abaxial lip 2–3.2 ×
<quantity id="7D129B01FFDF5266FDFCF7E1D8D42D15" box="[540,654,2064,2089]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="2" pageNumber="63" unit="mm" value="1.6" valueMax="2.0" valueMin="1.2">1.2–2 mm</quantity>
, ovate to elliptical, limb 3-toothed, white, adaxial lip 1.3–2 ×
<quantity id="7D129B01FFDF5266FAD4F7E1DFFD2D15" box="[1332,1447,2064,2089]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="2" pageNumber="63" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
, ovate, white, sometimes yellow at base. Anthers
<quantity id="7D129B01FFDE5267FD49FF51D9412585" box="[681,795,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="2.0" metricValueMin="1.1" pageId="3" pageNumber="64" unit="mm" value="1.55" valueMax="2.0" valueMin="1.1">1.1–2 mm</quantity>
long, dorsifixed, base sagittate, apical appendages 0.5–0.7 ×
<quantity id="7D129B01FFDE5267FF42FF35DB6725E1" box="[162,317,196,221]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="3" pageNumber="64" unit="mm" value="0.225" valueMax="0.25" valueMin="0.2">0.2–0.25 mm</quantity>
, narrowly elliptic to ovate, collar 0.25–0.4 ×
<quantity id="7D129B01FFDE5267FCB2FF35D98525E1" box="[850,991,196,221]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="64" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
. Style
<quantity id="7D129B01FFDE5267FBD1FF35DEE625E1" box="[1073,1212,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.6" metricValueMin="1.5" pageId="3" pageNumber="64" unit="mm" value="2.05" valueMax="2.6" valueMin="1.5">1.5–2.6 mm</quantity>
long, style branches
<quantity id="7D129B01FFDE5267FF68FF19DB4A243D" box="[136,272,232,257]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="3" pageNumber="64" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4–0.7 mm</quantity>
long, apex truncate, penicillate, sterile appendages ca.
<quantity id="7D129B01FFDE5267FC9EFF19D98F243D" box="[894,981,232,257]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="64" unit="mm" value="0.1">0.1 mm</quantity>
long, nectariferous disk ca.
<quantity id="7D129B01FFDE5267FAF5FF19DF31243D" box="[1301,1387,232,257]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="64" unit="mm" value="0.1">0.1 mm</quantity>
high. Achene
<quantity id="7D129B01FFDE5267FF05FEFDDB0D2419" box="[229,343,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="3" pageNumber="64" unit="mm" value="1.95" valueMax="2.0" valueMin="1.9">1.9–2 mm</quantity>
long, obovoid, with a short carpopodium, sparsely covered by twin trichomes.
</paragraph>
<paragraph id="BA5536E4FFDE5267FF5DFEC1D88F271D" blockId="3.[136,1452,160,617]" pageId="3" pageNumber="64">
<emphasis id="889EEAF6FFDE5267FF5DFEC1DB532475" bold="true" box="[189,265,304,329]" pageId="3" pageNumber="64">Note 1</emphasis>
:—Throughout the history, the generic name was spelled two ways: “
<taxonomicName id="7DEA4D67FFDE5267FBE6FEC1DE252475" authorityName="Lagasca" authorityYear="1811" box="[1030,1151,304,329]" class="Magnoliopsida" family="Asteraceae" genus="Panphalea" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis id="889EEAF6FFDE5267FBE6FEC1DE252475" box="[1030,1151,304,329]" italics="true" pageId="3" pageNumber="64">Panphalea</emphasis>
</taxonomicName>
” with “n” or “
<taxonomicName id="7DEA4D67FFDE5267FAFFFEC1DFC52475" authorityName="Lagasca" authorityYear="1811" box="[1311,1439,304,329]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis id="889EEAF6FFDE5267FAFFFEC1DFC52475" box="[1311,1439,304,329]" italics="true" pageId="3" pageNumber="64">Pamphalea</emphasis>
</taxonomicName>
” with “m”.
<bibRefCitation id="DE7B4B15FFDE5267FEE5FEA5DBEA2451" author="Katinas, L." box="[261,432,340,365]" pageId="3" pageNumber="64" pagination="5 - 54" refId="ref2107" refString="Katinas, L. (1995) Asteraceae, parte 1. Tribu XII. Mutisieae. Subtribu 4. Nassauviinae In: Hunziker, A. T. (Ed.) Flora fanerogamica Argentina 13. IMBIV, Cordoba, pp. 5 - 54." type="book chapter" year="1995">Katinas (1995)</bibRefCitation>
and
<bibRefCitation id="DE7B4B15FFDE5267FE0FFEA5D8BB2451" author="Katinas, L. &amp; Pruski, J. F. &amp; Sancho, G. &amp; Telleria, M. C." box="[495,737,340,365]" pageId="3" pageNumber="64" pagination="469 - 716" refId="ref2155" refString="Katinas, L., Pruski, J. F., Sancho, G. &amp; Telleria, M. C. (2008) The subfamily Mutisioideae (Asteraceae). Botanical Review 74: 469 - 716. https: // doi. org / 10.1007 / s 12229 - 008 - 9016 - 6" type="journal article" year="2008">
Katinas
<emphasis id="889EEAF6FFDE5267FDAFFEA5D8D62451" box="[591,652,340,365]" italics="true" pageId="3" pageNumber="64">et al.</emphasis>
(2008)
</bibRefCitation>
proposed to use the original form “
<taxonomicName id="7DEA4D67FFDE5267FB6BFEA5DF5E2451" authorityName="Lagasca" authorityYear="1811" box="[1163,1284,340,365]" class="Magnoliopsida" family="Asteraceae" genus="Panphalea" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis id="889EEAF6FFDE5267FB6BFEA5DF5E2451" box="[1163,1284,340,365]" italics="true" pageId="3" pageNumber="64">Panphalea</emphasis>
</taxonomicName>
” as originally published by
<bibRefCitation id="DE7B4B15FFDE5267FEFFFE89DB9424AD" author="Lagasca, M." box="[287,462,376,401]" pageId="3" pageNumber="64" pagination="26 - 44" refId="ref2213" refString="Lagasca, M. (1811) Disertacion sobre un orden nuevo de plantas de la clase de las Compuestas. In: Lagasca, M. (Ed.) Amenidades Naturales de las Espanas. Imprenta de la Muy Ilustre Junta, Orihuela, pp. 26 - 44." type="book chapter" year="1811">Lagasca (1811)</bibRefCitation>
, while
<bibRefCitation id="DE7B4B15FFDE5267FDFFFE89D8B924AD" author="Platt, F. H. &amp; Bundchen, M. &amp; Sfoggia Miotto, S. T." box="[543,739,376,401]" pageId="3" pageNumber="64" pagination="139 - 140" refId="ref2346" refString="Platt, F. H., Bundchen, M. &amp; Sfoggia Miotto, S. T. (2016) Nomenclatural notes on Pamphalea (Asteraceae: Nassauvieae). Phytotaxa 273: 139 - 140. https: // doi. org / 10.11646 / phytotaxa. 273.2.6" type="journal article" year="2016">
Platt
<emphasis id="889EEAF6FFDE5267FDB9FE89D8C824AD" box="[601,658,376,401]" italics="true" pageId="3" pageNumber="64">et al.</emphasis>
(2016)
</bibRefCitation>
proposed to maintain the spelling correction made by
<bibRefCitation id="DE7B4B15FFDE5267FAA7FE89DA8C2489" author="Candolle, A. de" pageId="3" pageNumber="64" pagination="59 - 72" refId="ref1962" refString="Candolle, A. de (1812) Observations sur les plantes Composees, ou Syngeneses, troisieme memoire: Sur les Composees a corolles labie esou Labiatiflores. Annales du Museum d'Histoire Naturelle 19: 59 - 72." type="journal article" year="1812">Candolle (1812)</bibRefCitation>
. The spelling correction is in accordance with Art. 60.1 of the International Code of Nomenclature for algae, fungi and plants (
<bibRefCitation id="DE7B4B15FFDE5267FEAEFE31D87424E5" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[334,558,448,473]" pageId="3" pageNumber="64" refId="ref2480" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashutten, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="889EEAF6FFDE5267FE50FE31DBB324E5" box="[432,489,448,473]" italics="true" pageId="3" pageNumber="64">et al.</emphasis>
2018
</bibRefCitation>
). “
<taxonomicName id="7DEA4D67FFDE5267FDAEFE31D89424E5" authorityName="Lagasca" authorityYear="1811" box="[590,718,448,473]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis id="889EEAF6FFDE5267FDAEFE31D89424E5" box="[590,718,448,473]" italics="true" pageId="3" pageNumber="64">Pamphalea</emphasis>
</taxonomicName>
” is a compound word formed by the junction of the Greek roots “pan” and “phalos”, where the dental consonant “n” in juxtaposition becomes the labial “m” (
<bibRefCitation id="DE7B4B15FFDE5267FB41FE15DF0324C1" author="Platt, F. H. &amp; Bundchen, M. &amp; Sfoggia Miotto, S. T." box="[1185,1369,484,509]" pageId="3" pageNumber="64" pagination="139 - 140" refId="ref2346" refString="Platt, F. H., Bundchen, M. &amp; Sfoggia Miotto, S. T. (2016) Nomenclatural notes on Pamphalea (Asteraceae: Nassauvieae). Phytotaxa 273: 139 - 140. https: // doi. org / 10.11646 / phytotaxa. 273.2.6" type="journal article" year="2016">
Platt
<emphasis id="889EEAF6FFDE5267FB3DFE15DF4F24C1" box="[1245,1301,484,509]" italics="true" pageId="3" pageNumber="64">et al.</emphasis>
2016
</bibRefCitation>
). In the present work, the spelling “
<taxonomicName id="7DEA4D67FFDE5267FE5DFDF9D867271D" authorityName="Lagasca" authorityYear="1811" box="[445,573,520,545]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis id="889EEAF6FFDE5267FE5DFDF9D867271D" box="[445,573,520,545]" italics="true" pageId="3" pageNumber="64">Pamphalea</emphasis>
</taxonomicName>
” is followed.
</paragraph>
<paragraph id="BA5536E4FFDE5267FF5DFDDDDB112755" blockId="3.[136,1452,160,617]" pageId="3" pageNumber="64">
<emphasis id="889EEAF6FFDE5267FF5DFDDDDB4A2779" bold="true" box="[189,272,556,581]" pageId="3" pageNumber="64">Note 2</emphasis>
:—The glandular trichomes present in
<taxonomicName id="7DEA4D67FFDE5267FD3BFDDDD9852779" box="[731,991,556,581]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="tenuissima">
<emphasis id="889EEAF6FFDE5267FD3BFDDDD9852779" box="[731,991,556,581]" italics="true" pageId="3" pageNumber="64">Pamphalea tenuissima</emphasis>
</taxonomicName>
are of biseriate vesicular beta subtype (
<bibRefCitation id="DE7B4B15FFDE5267FF70FDA1DB642755" author="Ramayya, N." box="[144,318,592,617]" pageId="3" pageNumber="64" pagination="177 - 188" refId="ref2397" refString="Ramayya, N. (1962) Studies on the trichomes of some Compositae I. General structure. Bulletin of the Botanical Survey of India 4: 177 - 188." type="journal article" year="1962">Ramayya 1962</bibRefCitation>
).
</paragraph>
<caption id="EE95666CFFDE5267FF68F859D97922DD" ID-DOI="http://doi.org/10.5281/zenodo.13718049" ID-Zenodo-Dep="13718049" httpUri="https://zenodo.org/record/13718049/files/figure.png" pageId="3" pageNumber="64" startId="3.[136,229,1960,1981]" targetBox="[340,1247,645,1928]" targetPageId="3" targetType="figure">
<paragraph id="BA5536E4FFDE5267FF68F859D97922DD" blockId="3.[136,1452,1960,2017]" pageId="3" pageNumber="64">
<emphasis id="889EEAF6FFDE5267FF68F859DAA72281" bold="true" box="[136,253,1960,1981]" pageId="3" pageNumber="64">FIGURE 3.</emphasis>
<taxonomicName id="7DEA4D67FFDE5267FEE3F859DB872281" box="[259,477,1960,1981]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="64" phylum="Tracheophyta" rank="species" species="tenuissima">
<emphasis id="889EEAF6FFDE5267FEE3F859DB872281" box="[259,477,1960,1981]" italics="true" pageId="3" pageNumber="64">Pamphalea tenuissima</emphasis>
</taxonomicName>
.
<emphasis id="889EEAF6FFDE5267FE08F859D85A2281" bold="true" box="[488,512,1960,1981]" pageId="3" pageNumber="64">A.</emphasis>
Habit.
<emphasis id="889EEAF6FFDE5267FDA6F859D8232281" bold="true" box="[582,633,1960,1981]" pageId="3" pageNumber="64">B, C.</emphasis>
Cauline leaves.
<emphasis id="889EEAF6FFDE5267FCF6F859D9742281" bold="true" box="[790,814,1960,1981]" pageId="3" pageNumber="64">D.</emphasis>
Rosette leaf.
<emphasis id="889EEAF6FFDE5267FC51F859D9842281" bold="true" box="[945,990,1960,1981]" pageId="3" pageNumber="64">E, F.</emphasis>
Habit, the main rosette not shown in picture.
<emphasis id="889EEAF6FFDE5267FA73F859DFF62281" bold="true" box="[1427,1452,1960,1981]" pageId="3" pageNumber="64">G.</emphasis>
Head, lateral view.
<emphasis id="889EEAF6FFDE5267FEA2F83DDB0122DD" bold="true" box="[322,347,1996,2017]" pageId="3" pageNumber="64">H.</emphasis>
Head, superior view. Photographs: Bonifacino.
</paragraph>
</caption>
<paragraph id="BA5536E4FFD95260FF5DFF6EDEF12450" blockId="4.[136,1452,159,688]" pageId="4" pageNumber="65">
<emphasis id="889EEAF6FFD95260FF5DFF6EDB562584" bold="true" box="[189,268,159,184]" pageId="4" pageNumber="65">Note 3</emphasis>
:—
<taxonomicName id="7DEA4D67FFD95260FED0FF6ED86A2584" box="[304,560,159,184]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="tenuissima">
<emphasis id="889EEAF6FFD95260FED0FF6ED86A2584" box="[304,560,159,184]" italics="true" pageId="4" pageNumber="65">Pamphalea tenuissima</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName id="7DEA4D67FFD95260FC28FF6EDED02584" authorityName="Lessing" authorityYear="1830" box="[968,1162,159,184]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="cardaminifolia">
<emphasis id="889EEAF6FFD95260FC28FF6EDED02584" box="[968,1162,159,184]" italics="true" pageId="4" pageNumber="65">P. cardaminifolia</emphasis>
</taxonomicName>
, from which it differs by more finely dissected leaves with a softer texture (
<figureCitation id="22D12A61FFD95260FD27FF32D95625E0" box="[711,780,195,220]" captionStart="FIGURE 2" captionStartId="2.[136,229,1490,1511]" captionTargetBox="[386,1222,204,1428]" captionTargetId="figure-17@2.[340,1247,190,1466]" captionTargetPageId="2" captionText="FIGURE 2. Compared leaf morphology of Pamphalea cardaminifolia and Pamphalea tenuissima A. Rosette leaf of P. cardaminifolia. B. Rosette leaf of P. tenuissima. C. Cauline leaf of P. cardaminifolia. D. Cauline leaf of P. tenuissima." figureDoi="http://doi.org/10.5281/zenodo.13718047" httpUri="https://zenodo.org/record/13718047/files/figure.png" pageId="4" pageNumber="65">Fig. 2</figureCitation>
) and all or most of its stems being decumbent.
<taxonomicName id="7DEA4D67FFD95260FACCFF32DB77243C" authorityName="Lessing" authorityYear="1830" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="cardaminifolia">
<emphasis id="889EEAF6FFD95260FACCFF32DB77243C" italics="true" pageId="4" pageNumber="65">Pamphalea cardaminifolia</emphasis>
</taxonomicName>
has an erect stem and coarsely dissected, chartaceous leaves. Plants of
<taxonomicName id="7DEA4D67FFD95260FBAFFF16DF14243C" box="[1103,1358,231,256]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="tenuissima">
<emphasis id="889EEAF6FFD95260FBAFFF16DF14243C" box="[1103,1358,231,256]" italics="true" pageId="4" pageNumber="65">Pamphalea tenuissima</emphasis>
</taxonomicName>
seem to support themselves in surrounding vegetation, developing adventitious roots on nodes; they are generally associated with
<taxonomicName id="7DEA4D67FFD95260FF21FEDEDB042474" box="[193,350,303,328]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="4" pageNumber="65" phylum="Bryophyta" rank="species" species="undetermined">
<emphasis id="889EEAF6FFD95260FF21FEDEDB6D2474" box="[193,311,303,328]" italics="true" pageId="4" pageNumber="65">Sphagnum</emphasis>
sp.
</taxonomicName>
on flooded soils. Both
<taxonomicName id="7DEA4D67FFD95260FD82FEDED93A2474" box="[610,864,303,328]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="tenuissima">
<emphasis id="889EEAF6FFD95260FD82FEDED93A2474" box="[610,864,303,328]" italics="true" pageId="4" pageNumber="65">Pamphalea tenuissima</emphasis>
</taxonomicName>
and
<taxonomicName id="7DEA4D67FFD95260FC76FEDEDE0C2474" authorityName="Lessing" authorityYear="1830" box="[918,1110,303,328]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="cardaminifolia">
<emphasis id="889EEAF6FFD95260FC76FEDEDE0C2474" box="[918,1110,303,328]" italics="true" pageId="4" pageNumber="65">P. cardaminifolia</emphasis>
</taxonomicName>
occur in
<collectingRegion id="782EF806FFD95260FB5EFEDEDFD42474" box="[1214,1422,303,328]" country="Brazil" name="Rio Grande do Sul" pageId="4" pageNumber="65">Rio Grande do Sul</collectingRegion>
in waterlogged habitats, but
<taxonomicName id="7DEA4D67FFD95260FE49FEA2D8302450" authorityName="Lessing" authorityYear="1830" box="[425,618,339,364]" class="Magnoliopsida" family="Asteraceae" genus="Pamphalea" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="cardaminifolia">
<emphasis id="889EEAF6FFD95260FE49FEA2D8302450" box="[425,618,339,364]" italics="true" pageId="4" pageNumber="65">P. cardaminifolia</emphasis>
</taxonomicName>
has not been observed growing with
<taxonomicName id="7DEA4D67FFD95260FBEDFEA2DEF12450" box="[1037,1195,339,364]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="4" pageNumber="65" phylum="Bryophyta" rank="species" species="undetermined">
<emphasis id="889EEAF6FFD95260FBEDFEA2DED92450" box="[1037,1155,339,364]" italics="true" pageId="4" pageNumber="65">Sphagnum</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F2F0656FFFD95260FF5DFE86D91D278C" pageId="4" pageNumber="65" type="distribution">
<paragraph id="BA5536E4FFD95260FF5DFE86D81524E4" blockId="4.[136,1452,159,688]" pageId="4" pageNumber="65">
<emphasis id="889EEAF6FFD95260FF5DFE86DB8624AC" bold="true" box="[189,476,375,400]" pageId="4" pageNumber="65">Distribution and habitat</emphasis>
:—Southern
<collectingCountry id="C2FD7674FFD95260FD89FE86D8EB24AC" box="[617,689,375,400]" name="Brazil" pageId="4" pageNumber="65">Brazil</collectingCountry>
; so far considered endemic to the South Brazilian highland grasslands in the state of
<collectingRegion id="782EF806FFD95260FEC6FE6ADBA02488" box="[294,506,411,436]" country="Brazil" name="Rio Grande do Sul" pageId="4" pageNumber="65">Rio Grande do Sul</collectingRegion>
. It occurs on waterlogged soils associated with
<taxonomicName id="7DEA4D67FFD95260FBECFE6ADED82488" authorityName="C.Linnaeus" authorityYear="1753" box="[1036,1154,411,436]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" order="Sphagnales" pageId="4" pageNumber="65" phylum="Bryophyta" rank="genus">
<emphasis id="889EEAF6FFD95260FBECFE6ADED82488" box="[1036,1154,411,436]" italics="true" pageId="4" pageNumber="65">Sphagnum</emphasis>
</taxonomicName>
, or sometimes on marshes dominated by
<taxonomicName id="7DEA4D67FFD95260FEC8FE4EDB9F24E4" box="[296,453,447,472]" class="Liliopsida" family="Eriocaulaceae" kingdom="Plantae" order="Poales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="family">Eriocaulaceae</taxonomicName>
(
<figureCitation id="22D12A61FFD95260FE35FE4ED81824E4" box="[469,578,447,472]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="3.[136,229,1960,1981]" captionStartId-1="4.[136,229,1912,1933]" captionTargetBox-0="[340,1247,645,1928]" captionTargetId-0="figure-225@3.[340,1247,645,1929]" captionTargetPageId-0="3" captionText-0="FIGURE 3. Pamphalea tenuissima. A. Habit. B, C. Cauline leaves. D. Rosette leaf. E, F. Habit, the main rosette not shown in picture. G. Head, lateral view. H. Head, superior view. Photographs: Bonifacino." captionText-1="FIGURE 4. Distribution of Pamphalea tenuissima." figureDoi-0="http://doi.org/10.5281/zenodo.13718049" httpUri-0="https://zenodo.org/record/13718049/files/figure.png" pageId="4" pageNumber="65">Figs. 3–4</figureCitation>
).
</paragraph>
<paragraph id="BA5536E4FFD95260FF5DFE12DE1B24C0" blockId="4.[136,1452,159,688]" box="[189,1089,483,508]" pageId="4" pageNumber="65">
<emphasis id="889EEAF6FFD95260FF5DFE12DB6324C0" bold="true" box="[189,313,483,508]" pageId="4" pageNumber="65">Phenology</emphasis>
:—Flowering from November to December, fruiting from December.
</paragraph>
<paragraph id="BA5536E4FFD95260FF5DFDF6D91D278C" blockId="4.[136,1452,159,688]" pageId="4" pageNumber="65">
<materialsCitation id="0A823CB9FFD95260FF5DFDF6D9BD2778" collectingDate="2015-12-09" collectionCode="S, W, MVFA" collectorName="Sao Jose Dos Ausentes" country="Brazil" location="Rio Grande do Sul" pageId="4" pageNumber="65" specimenCount="2" stateProvince="Rio Grande do Sul">
<emphasis id="889EEAF6FFD95260FF5DFDF6D8BE271C" bold="true" box="[189,740,519,544]" pageId="4" pageNumber="65">
<typeStatus id="65518846FFD95260FF5DFDF6D810271C" box="[189,586,519,544]" isOtherTypeStatus="true" pageId="4" pageNumber="65">Additional specimens examined</typeStatus>
(
<typeStatus id="65518846FFD95260FD82FDF6D886271C" box="[610,732,519,544]" pageId="4" pageNumber="65" type="paratype">paratypes</typeStatus>
)
</emphasis>
:—
<collectingCountry id="C2FD7674FFD95260FCE5FDF6D934271C" box="[773,878,519,544]" name="Brazil" pageId="4" pageNumber="65">BRASIL</collectingCountry>
.
<collectingRegion id="782EF806FFD95260FC62FDF6DE29271C" box="[898,1139,519,544]" country="Brazil" name="Rio Grande do Sul" pageId="4" pageNumber="65">Rio Grande do Sul</collectingRegion>
:
<collectorName id="171F5332FFD95260FB67FDF6DFF2271C" box="[1159,1448,519,544]" pageId="4" pageNumber="65">São José Dos Ausentes</collectorName>
, [28˚42’32.7”
<collectionCode id="DCFBAE21FFD95260FEFDFDDADB762778" box="[285,300,555,580]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="4" pageNumber="65" type="Herbarium">S</collectionCode>
, 50˚11’9.7”
<collectionCode id="DCFBAE21FFD95260FE4FFDDADB952778" box="[431,463,555,580]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="4" pageNumber="65" type="Herbarium">W</collectionCode>
],
<date id="CE541024FFD95260FE01FDDAD8F12778" box="[481,683,555,580]" pageId="4" pageNumber="65" value="2015-12-09">
<collectingDate id="DE10E9CCFFD95260FE01FDDAD8F12778" box="[481,683,555,580]" pageId="4" pageNumber="65" value="2015-12-09">9 December 2015</collectingDate>
</date>
,
<emphasis id="889EEAF6FFD95260FD56FDDAD9212778" box="[694,891,555,580]" italics="true" pageId="4" pageNumber="65">Pasini et al. 1076</emphasis>
(
<collectionCode id="DCFBAE21FFD95260FC6AFDDAD9BA2778" box="[906,992,555,580]" country="Uruguay" httpUri="http://grbio.org/cool/x4p2-8dvd" name="Universidad de la República" pageId="4" pageNumber="65" type="Herbarium">MVFA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0A823CB9FFD95260FC12FDDADB3627B0" collectingDate="2015-12-10" collectionCode="S, W, MVFA" collectorName="Sao Jose dos Ausentes" country="Brazil" elevation="500" pageId="4" pageNumber="65" specimenCount="1">
from
<collectorName id="171F5332FFD95260FBCFFDDADF702778" box="[1071,1322,555,580]" pageId="4" pageNumber="65">São José dos Ausentes</collectorName>
to Silveira, left side, in humid low to
<quantity id="7D129B01FFD95260FE43FDBEDBB32754" box="[419,489,591,616]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="65" unit="m" value="500.0">
<elevation id="31C7D1D7FFD95260FE43FDBEDBB32754" box="[419,489,591,616]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="65" unit="m" value="500.0">500 m</elevation>
</quantity>
of registered coordinates, [28˚39’23.17”
<collectionCode id="DCFBAE21FFD95260FC51FDBED99A2754" box="[945,960,591,616]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="4" pageNumber="65" type="Herbarium">S</collectionCode>
, 49˚57’9.71”
<collectionCode id="DCFBAE21FFD95260FBB3FDBEDE282754" box="[1107,1138,591,616]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="4" pageNumber="65" type="Herbarium">W</collectionCode>
],
<date id="CE541024FFD95260FB64FDBEDF002754" box="[1156,1370,591,616]" pageId="4" pageNumber="65" value="2015-12-10">
<collectingDate id="DE10E9CCFFD95260FB64FDBEDF002754" box="[1156,1370,591,616]" pageId="4" pageNumber="65" value="2015-12-10">10 December 2015</collectingDate>
</date>
,
<emphasis id="889EEAF6FFD95260FA84FDBEDAA527B0" italics="true" pageId="4" pageNumber="65">Pasini et al. 1079</emphasis>
(
<collectionCode id="DCFBAE21FFD95260FEEEFD82DB3E27B0" box="[270,356,627,652]" country="Uruguay" httpUri="http://grbio.org/cool/x4p2-8dvd" name="Universidad de la República" pageId="4" pageNumber="65" type="Herbarium">MVFA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0A823CB9FFD95260FE97FD82DF4B27B0" box="[375,1297,626,652]" collectingDate="2015-12-10" collectionCode="S, W, MVFA" country="Brazil" pageId="4" pageNumber="65" specimenCount="1">
[28˚35’17.53”
<collectionCode id="DCFBAE21FFD95260FDF9FD82D87227B0" box="[537,552,627,652]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="4" pageNumber="65" type="Herbarium">S</collectionCode>
, 49˚50’38.65”
<collectionCode id="DCFBAE21FFD95260FD2BFD82D8B327B0" box="[715,745,627,652]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="4" pageNumber="65" type="Herbarium">W</collectionCode>
],
<date id="CE541024FFD95260FD1CFD82D98E27B0" box="[764,980,627,652]" pageId="4" pageNumber="65" value="2015-12-10">
<collectingDate id="DE10E9CCFFD95260FD1CFD82D98E27B0" box="[764,980,627,652]" pageId="4" pageNumber="65" value="2015-12-10">10 December 2015</collectingDate>
</date>
,
<emphasis id="889EEAF6FFD95260FC3FFD82DEFF27B0" box="[991,1189,627,652]" italics="true" pageId="4" pageNumber="65">Pasini et al. 1085</emphasis>
(
<collectionCode id="DCFBAE21FFD95260FB53FD82DF5327B0" box="[1203,1289,627,652]" country="Uruguay" httpUri="http://grbio.org/cool/x4p2-8dvd" name="Universidad de la República" pageId="4" pageNumber="65" type="Herbarium">MVFA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0A823CB9FFD95260FAFCFD82D919278C" collectingDate="2007-11-21" collectionCode="ICN" collectorName="Sao Jose dos Ausentes" country="Brazil" pageId="4" pageNumber="65" specimenCode="ICN 152230" specimenCount="1">
<collectorName id="171F5332FFD95260FAFCFD82DAA8278C" pageId="4" pageNumber="65">São José dos Ausentes</collectorName>
,
<date id="CE541024FFD95260FF1DFD66DB82278C" box="[253,472,663,688]" pageId="4" pageNumber="65" value="2007-11-21">
<collectingDate id="DE10E9CCFFD95260FF1DFD66DB82278C" box="[253,472,663,688]" pageId="4" pageNumber="65" value="2007-11-21">21 November 2007</collectingDate>
</date>
,
<emphasis id="889EEAF6FFD95260FE03FD66D8C8278C" box="[483,658,663,688]" italics="true" pageId="4" pageNumber="65">Schneider 1547</emphasis>
(
<specimenCode id="EA4C9E9FFFD95260FD40FD66D96D278C" box="[672,823,663,688]" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="4" pageNumber="65">ICN 152230</specimenCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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