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<document id="6BDF7617FA2A7F1DDAAF5900AD2128C5" ID-DOI="10.5281/zenodo.5353792" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353792" ID-ZooBank="88D45C5E-0E49-4F02-87A4-C4FA55A83B2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212659467" checkinUser="felipe" docAuthor="Freese, Edwin L., Veal, Doug A. &amp; Lago, Paul K." docDate="2020" docId="28608792FFAC3651FF76FB70BAD1F9F7" docLanguage="en" docName="0787.pdf" docOrigin="Insecta Mundi 2020 (787)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Phyllophaga vilifrons" docType="treatment" docVersion="3" lastPageNumber="41" masterDocId="D459FFEAFF86367BFFB6FFEDBB37FF84" masterDocTitle="The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist" masterLastPageNumber="83" masterPageNumber="1" pageNumber="41" updateTime="1699188000474" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="9FDB113014DE68F46409DB2A18CF4A8A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="56771E9C29A31967C1F20251C73A139E">
<mods:title id="47712AF1B8AA345698D57B3B2630BFD9">The Scarabaeoidea (Coleoptera) of Iowa: An annotated checklist</mods:title>
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<mods:name id="A31423D54560748C76EA80FB26D58B1E" type="personal">
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<mods:roleTerm id="EDEF4FA0046C626104F0CD33ACA0AE26">Author</mods:roleTerm>
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<mods:namePart id="E10FF4C2AC44E8111AD6363CF0B516D9">Freese, Edwin L.</mods:namePart>
<mods:affiliation id="D8671FDA86472EE7A174EF82DD0BA256">33493 “ S ” Avenue Adel, Iowa 50003</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="1AD46C82076E67869F80463CF88725CB">Veal, Doug A.</mods:namePart>
<mods:affiliation id="BC5D487255F700136E79C1BCD1AEAB2F">168 Thompson Drive SE Cedar Rapids, Iowa 52403</mods:affiliation>
</mods:name>
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<mods:namePart id="0AA9F0B023D40A7435AA5D62E9ECBF8E">Lago, Paul K.</mods:namePart>
<mods:affiliation id="C0C5AA5223A3CC0DDD7E9B593C86FA73">Biology Department, University of Mississippi University, Mississippi 38677</mods:affiliation>
</mods:name>
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<mods:title id="A8AC4B1F7C00D199981DDD3329EB5A98">Insecta Mundi</mods:title>
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<mods:part id="E534D4BBA490F8D3534B49121373F8AB">
<mods:date id="C541760D58D0CBAFEF51F661D2A2F6EC">2020</mods:date>
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<mods:number id="AB0751A918DFAE79F0143D619D99B917">2020-08-28</mods:number>
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<mods:number id="349482CD545E71B9554F39B13716774B">2020</mods:number>
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<mods:number id="FE8BA5D881A281AF235869F01A599D47">787</mods:number>
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<mods:identifier id="36BD8EF066071437FF94471FF2143EFE" type="DOI">10.5281/zenodo.5353792</mods:identifier>
<mods:identifier id="8C97CF2C1DDCB72D8953E634739BB752" type="ISSN">1942-1354</mods:identifier>
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<treatment id="28608792FFAC3651FF76FB70BAD1F9F7" LSID="urn:lsid:plazi:treatment:28608792FFAC3651FF76FB70BAD1F9F7" httpUri="http://treatment.plazi.org/id/28608792FFAC3651FF76FB70BAD1F9F7" lastPageNumber="41" pageId="42" pageNumber="41">
<subSubSection id="E8D3650FFFAC3651FF76FB70BA25FA84" pageId="42" pageNumber="41" type="nomenclature">
<paragraph id="A0763684FFAC3651FF76FB70B9F1FB32" blockId="42.[192,710,1181,1206]" box="[192,710,1181,1206]" pageId="42" pageNumber="41">
<heading id="FB3E81E8FFAC3651FF76FB70B9F1FB32" bold="true" box="[192,710,1181,1206]" fontSize="10" level="2" pageId="42" pageNumber="41" reason="3">
<taxonomicName id="67C94D07FFAC3651FF76FB70B9F1FB32" authority="(LeConte, 1856)" baseAuthorityName="LeConte" baseAuthorityYear="1856" box="[192,710,1181,1206]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="41" phylum="Arthropoda" rank="species" species="vilifrons">
<emphasis id="92BDEA96FFAC3651FF76FB70B9F1FB32" bold="true" box="[192,710,1181,1206]" pageId="42" pageNumber="41">
<emphasis id="92BDEA96FFAC3651FF76FB70BADDFB31" bold="true" box="[192,490,1181,1205]" italics="true" pageId="42" pageNumber="41">Phyllophaga vilifrons</emphasis>
(LeConte, 1856)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A0763684FFAC3651FF76FB26BA25FA84" blockId="42.[192,1440,1227,1280]" pageId="42" pageNumber="41">
<bibRefCitation id="C4584B75FFAC3651FF76FB26BA71FB67" author="Horn, G. W." box="[192,326,1227,1251]" pageId="42" pageNumber="41" pagination="209 - 296" refId="ref32091" refString="Horn, G. W. 1887. Revision of the species of Lachnosterna of America north of Mexico. Transactions of the American Entomological Society 14: 209 - 296." type="journal article" year="1887">Horn 1887</bibRefCitation>
;
<bibRefCitation id="C4584B75FFAC3651FEE4FB26B93AFB67" author="Wickham, H. F." box="[338,525,1227,1251]" pageId="42" pageNumber="41" pagination="1 - 40" refId="ref34735" refString="Wickham, H. F. 1911. A list of the Coleoptera of Iowa. Bulletin of the State University of Iowa Laboratory of Natural History 6 (2): 1 - 40." type="journal article" year="1911">Wickham 1911</bibRefCitation>
;
<bibRefCitation id="C4584B75FFAC3651FDACFB26B998FB67" author="Travis, B. V." box="[538,687,1227,1251]" pageId="42" pageNumber="41" pagination="313 - 365" refId="ref34468" refString="Travis, B. V. 1934. Phyllophaga of Iowa. Iowa State Journal of Science 8 (2): 313 - 365." type="journal article" year="1934">Travis 1934</bibRefCitation>
;
<bibRefCitation id="C4584B75FFAC3651FD0DFB26BF3AFB67" author="Luginbill, P. &amp; H. R. Painter" box="[699,1037,1227,1251]" pageId="42" pageNumber="41" pagination="1 - 102" refId="ref32964" refString="Luginbill, P., and H. R. Painter. 1953. May beetles of the United States and Canada. United States Department of Agriculture Technical Bulletin 1061: 1 - 102." type="journal article" year="1953">Luginbill and Painter 1953</bibRefCitation>
;
<bibRefCitation id="C4584B75FFAC3651FBACFB26BFC2FB67" author="Harpootlian, P. J." box="[1050,1269,1227,1251]" pageId="42" pageNumber="41" refId="ref31897" refString="Harpootlian, P. J. 2001. Scarab beetles (Coleoptera: Scarabaeidae) of South Carolina. Biota of South Carolina. Volume 2. Clemson University; Clemson, SC. 157 p." type="book" year="2001">Harpootlian 2001</bibRefCitation>
; ISIC; IWUC; ELFC.
</paragraph>
</subSubSection>
<subSubSection id="E8D3650FFFAC3651FF76FAE2BAD1F9F7" pageId="42" pageNumber="41" type="description">
<paragraph id="A0763684FFAC3651FF76FAE2B928FAA3" blockId="42.[192,543,1295,1319]" box="[192,543,1295,1319]" pageId="42" pageNumber="41">Collection dates: AprilJuly.</paragraph>
<paragraph id="A0763684FFAC3651FF76FADBB877FACA" blockId="42.[192,832,1334,1358]" box="[192,832,1334,1358]" pageId="42" pageNumber="41">Collected at light; also collected under rock in April.</paragraph>
<paragraph id="A0763684FFAC3651FF76FAB3BE37FAF1" blockId="42.[192,1280,1374,1398]" box="[192,1280,1374,1398]" pageId="42" pageNumber="41">
<bibRefCitation id="C4584B75FFAC3651FF76FAB3BA53FAF2" author="Travis, B. V." box="[192,356,1374,1398]" pageId="42" pageNumber="41" pagination="313 - 365" refId="ref34468" refString="Travis, B. V. 1934. Phyllophaga of Iowa. Iowa State Journal of Science 8 (2): 313 - 365." type="journal article" year="1934">Travis (1934)</bibRefCitation>
reported only
<specimenCount id="B6CFFD0DFFAC3651FDA1FAB3B9E7FAF2" box="[535,720,1374,1398]" pageId="42" pageNumber="41" type="generic">four specimens</specimenCount>
from
<collectingRegion id="620DF866FFAC3651FCAFFAB3B862FAF1" box="[793,853,1374,1397]" country="United States of America" name="Iowa" pageId="42" pageNumber="41">Iowa</collectingRegion>
indicating it was rare in the state.
</paragraph>
<paragraph id="A0763684FFAC3651FF76FA68BA9DFA70" blockId="42.[191,1440,1413,1524]" pageId="42" pageNumber="41">
<materialsCitation id="10A13CD9FFAC3651FF76FA68BE34FA19" box="[192,1283,1413,1437]" collectingDate="1929-06-01" collectionCode="ISIC" collectorName="Specimen &amp; Ames" county="Story Co." location="Davis" municipality="Ralfs" pageId="42" pageNumber="41" specimenCount="1" stateProvince="Iowa">
<collectorName id="0D3C5352FFAC3651FF76FA68BA00FA19" box="[192,311,1413,1437]" pageId="42" pageNumber="41">Specimen</collectorName>
label data:
<collectorName id="0D3C5352FFAC3651FE70FA68B939FA19" box="[454,526,1413,1437]" pageId="42" pageNumber="41">Ames</collectorName>
,
<collectingCounty id="49174E08FFAC3651FDAFFA68B9BAFA19" box="[537,653,1413,1437]" pageId="42" pageNumber="41">Story Co.</collectingCounty>
,
<collectingRegion id="620DF866FFAC3651FD2FFA6BB9EFFA19" box="[665,728,1414,1437]" country="United States of America" name="Iowa" pageId="42" pageNumber="41">Iowa</collectingRegion>
,
<date id="D4771044FFAC3651FD54FA68B84DFA19" box="[738,890,1413,1437]" pageId="42" pageNumber="41" value="1929-06-01">
<collectingDate id="C433E9ACFFAC3651FD54FA68B84DFA19" box="[738,890,1413,1437]" pageId="42" pageNumber="41" value="1929-06-01">1 June 1929</collectingDate>
</date>
,
<collectingMunicipality id="4012ACFEFFAC3651FC33FA68B8FFFA19" box="[901,968,1413,1437]" pageId="42" pageNumber="41">Ralfs</collectingMunicipality>
, determined
<location id="A516605FFFAC3651FBD1FA68BF99FA19" LSID="urn:lsid:plazi:treatment:28608792FFAC3651FF76FB70BAD1F9F7:A516605FFFAC3651FBD1FA68BF99FA19" box="[1127,1198,1413,1437]" county="Story Co." municipality="Ralfs" name="Davis" pageId="42" pageNumber="41" stateProvince="Iowa">Davis</location>
(
<collectionCode id="C6D8AE41FFAC3651FB0BFA68BFCDFA19" box="[1213,1274,1413,1437]" pageId="42" pageNumber="41">ISIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="10A13CD9FFAC3651FAB8FA68B887FA3E" collectingDate="2005-05-24" collectionCode="ELFC" collectorName="Cedar Bend Park" county="Bremer Co." location="Bremer Co." pageId="42" pageNumber="41" specimenCount="1" stateProvince="Iowa">
<collectorName id="0D3C5352FFAC3651FAB8FA68BBC9FA3E" pageId="42" pageNumber="41">Cedar Bend Park</collectorName>
,
<collectingCounty id="49174E08FFAC3651FEBCFA4EBAAEFA3E" box="[266,409,1442,1466]" pageId="42" pageNumber="41">Bremer Co.</collectingCounty>
,
<collectingRegion id="620DF866FFAC3651FE13FA4EBAD3FA3E" box="[421,484,1443,1466]" country="United States of America" name="Iowa" pageId="42" pageNumber="41">Iowa</collectingRegion>
,
<date id="D4771044FFAC3651FE59FA4FB9B8FA3E" box="[495,655,1442,1467]" pageId="42" pageNumber="41" value="2005-05-24">
<collectingDate id="C433E9ACFFAC3651FE59FA4FB9B8FA3E" box="[495,655,1442,1467]" pageId="42" pageNumber="41" value="2005-05-24">24 May 2005</collectingDate>
</date>
, UV light, ELF (
<collectionCode id="C6D8AE41FFAC3651FCECFA4FB89FFA3E" box="[858,936,1442,1466]" pageId="42" pageNumber="41">ELFC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="10A13CD9FFAC3651FC0AFA4FB995FA53" collectingDate="2007-04-20" collectionCode="ELFC" county="Pammel State Park Madison Co." location="Pammel State Park Madison Co." pageId="42" pageNumber="41" specimenCount="1" stateProvince="Iowa">
<collectingCounty id="49174E08FFAC3651FC0AFA4FBE67FA3E" box="[956,1360,1442,1466]" pageId="42" pageNumber="41">Pammel State Park Madison Co.</collectingCounty>
,
<collectingRegion id="620DF866FFAC3651FAEAFA4EBEACFA3E" box="[1372,1435,1443,1466]" country="United States of America" name="Iowa" pageId="42" pageNumber="41">Iowa</collectingRegion>
,
<date id="D4771044FFAC3651FF76FA52BA5FFA53" box="[192,360,1471,1495]" pageId="42" pageNumber="41" value="2007-04-20">
<collectingDate id="C433E9ACFFAC3651FF76FA52BA5FFA53" box="[192,360,1471,1495]" pageId="42" pageNumber="41" value="2007-04-20">20 April 2007</collectingDate>
</date>
, under rock, ELF (
<collectionCode id="C6D8AE41FFAC3651FDFDFA52B9AEFA53" box="[587,665,1471,1495]" pageId="42" pageNumber="41">ELFC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="10A13CD9FFAC3651FD1BFA52BA92FA70" collectingDate="2018-07-12" collectionCode="ELFC" county="Appanoose Co." location="Sharon Bluffs State Park" pageId="42" pageNumber="41" specimenCount="1" stateProvince="Iowa">
<location id="A516605FFFAC3651FD1BFA52B8D2FA53" LSID="urn:lsid:plazi:treatment:28608792FFAC3651FF76FB70BAD1F9F7:A516605FFFAC3651FD1BFA52B8D2FA53" box="[685,997,1471,1495]" county="Appanoose Co." name="Sharon Bluffs State Park" pageId="42" pageNumber="41" stateProvince="Iowa">Sharon Bluffs State Park</location>
,
<collectingCounty id="49174E08FFAC3651FC46FA52BF90FA53" box="[1008,1191,1471,1495]" pageId="42" pageNumber="41">Appanoose Co.</collectingCounty>
,
<collectingRegion id="620DF866FFAC3651FB04FA52BFC6FA52" box="[1202,1265,1471,1494]" country="United States of America" name="Iowa" pageId="42" pageNumber="41">Iowa</collectingRegion>
,
<date id="D4771044FFAC3651FB4AFA52BEACFA53" box="[1276,1435,1471,1495]" pageId="42" pageNumber="41" value="2018-07-12">
<collectingDate id="C433E9ACFFAC3651FB4AFA52BEACFA53" box="[1276,1435,1471,1495]" pageId="42" pageNumber="41" value="2018-07-12">12 July 2018</collectingDate>
</date>
, light, ELF (
<collectionCode id="C6D8AE41FFAC3651FEF8FA31BAAAFA70" box="[334,413,1500,1524]" pageId="42" pageNumber="41">ELFC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A0763684FFAC3651FF76F9EEBE66F99F" blockId="42.[192,1361,1539,1564]" box="[192,1361,1539,1564]" pageId="42" pageNumber="41">
County records (
<figureCitation id="38F22A01FFAC3651FE3DF9EEB923F99F" box="[395,532,1539,1563]" captionStart="Figures 146153" captionStartId="79.[192,286,1847,1869]" captionTargetBox="[123,1439,217,1801]" captionTargetPageId="79" captionText="Figures 146153. Distribution of Scarabaeoidea in Iowa.146) Phyllophaga quercus (Knoch, 1801).147) Phyllophaga rugosa (Melsheimer, 1846). 148) Phyllophaga spreta (Horn, 1887). 149) Phyllophaga tristis (Fabricius, 1781). 150) Phyllophaga vehemens (Horn, 1887). 151) Phyllophaga vilifrons (LeConte, 1856). 152) Polyphylla hammondi" figureDoi="http://doi.org/10.5281/zenodo.5353834" httpUri="https://zenodo.org/record/5353834/files/figure.png" pageId="42" pageNumber="41">Figure 151</figureCitation>
): Allamakee, Appanoose, Bremer, Henry, Johnson, Madison, Story.
</paragraph>
<paragraph id="A0763684FFAC3651FF76F9B7BAD1F9F7" blockId="42.[192,486,1626,1651]" box="[192,486,1626,1651]" pageId="42" pageNumber="41">
<taxonomicName id="67C94D07FFAC3651FF76F9B7BAD1F9F7" authority="Harris, 1841" authorityName="Harris" authorityYear="1841" box="[192,486,1626,1651]" class="Insecta" family="Melolonthidae" genus="Polyphylla" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="92BDEA96FFAC3651FF76F9B7BA74F9F6" box="[192,323,1626,1650]" italics="true" pageId="42" pageNumber="41">Polyphylla</emphasis>
Harris, 1841
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>