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<document id="84D763B86C05560C858D397A7A427E09" ID-CLB-Dataset="25015" ID-DOI="10.11646/zootaxa.4691.1.7" ID-GBIF-Dataset="271f8735-1aa3-4f63-a742-aef356f505e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3527097" ID-ZooBank="76349E63-14F4-4E12-88B6-DCEC6F667694" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1572857365488" checkinUser="plazi" docAuthor="Yin, Zi-Wei &amp; Jantarit, Sopark" docDate="2019" docId="2827A302FF96C73E0385DBFE7679FDD7" docLanguage="en" docName="zootaxa.4691.1.7.pdf" docOrigin="Zootaxa 4691 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Araneibatrus phuphaphet Yin &amp; Jantarit 2019, sp. nov." docType="treatment" docVersion="6" lastPageNumber="85" masterDocId="D41EDB7AFF97C73C0312DB1F723EFFE8" masterDocTitle="Araneibatrus phuphaphet sp. nov., a new cavernicolous Pselaphinae from Satun Geopark, southern Thailand (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="87" masterPageNumber="83" pageNumber="84" updateTime="1698761628480" updateUser="ExternalLinkService">
<mods:mods id="A4AB3CD53E262672855EA2ABE45C524E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A8AAF38DD39A7DBE46847BE87E5B6694">
<mods:title id="8763573A10E5F0DB20461C90A0E2E4AD">Araneibatrus phuphaphet sp. nov., a new cavernicolous Pselaphinae from Satun Geopark, southern Thailand (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="EEE92F8CB1EC4D1950A7007E356A8DAD" type="personal">
<mods:role id="F287C008E3491D0EAA002942FFA92859">
<mods:roleTerm id="861410F15B0117008BC3AD528D74BFAA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="61C1D83A468D0690CA5B798DA79AE33D">Yin, Zi-Wei</mods:namePart>
</mods:name>
<mods:name id="00FD6A27535F608D887353C57196E4B1" type="personal">
<mods:role id="6854B965D25EEA0D510D914E9515AB31">
<mods:roleTerm id="3F55D6ADD9EE1E31772DA0A6ADA1DCBA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="94E0904155804831DD1779D2A7FCCACA">Jantarit, Sopark</mods:namePart>
</mods:name>
<mods:typeOfResource id="557BC9E024C39997CA4744F6EFE8C690">text</mods:typeOfResource>
<mods:relatedItem id="147AE5F2C7F1DED164BA0B3C167C3ECE" type="host">
<mods:titleInfo id="4A25210C99D5D668B71E122B123B4FF9">
<mods:title id="92231117437A9992EB0895E659823D51">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C6D891B4E8ACF6F8834AF2FFDF19A001">
<mods:date id="0F82479D87B71AD8EFFEF4D667E7F772">2019</mods:date>
<mods:detail id="18328D2E2A0561D7CAE7E9B7C81CAC6E" type="pubDate">
<mods:number id="B4000A54CE94BE34683B5223555486BC">2019-10-29</mods:number>
</mods:detail>
<mods:detail id="1B0D727B6A62A93F2CF67CFA428B0B4F" type="volume">
<mods:number id="CF32E34DF63B352049492673A9D2372E">4691</mods:number>
</mods:detail>
<mods:detail id="F373275C449775700ACA3C22BF2F91B7" type="issue">
<mods:number id="8B95AFF445751F2BF7743C6A07149EE2">1</mods:number>
</mods:detail>
<mods:extent id="AA6871BD7148F2EA4794441B7252509A" unit="page">
<mods:start id="0EA6EFF54F863E87223FC12BF0763460">83</mods:start>
<mods:end id="0F85AA527057858934F4EDBB60F7C0C9">87</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0345D1E4BC7429F9DC66AAD5CAA8548B">journal article</mods:classification>
<mods:identifier id="7B6BA184B14AC18B19B30EA568442F1D" type="CLB-Dataset">25015</mods:identifier>
<mods:identifier id="CB6FF6A6FACB9D2FFC75FE076817E939" type="DOI">10.11646/zootaxa.4691.1.7</mods:identifier>
<mods:identifier id="5005CF9ADFA8EB78FF7775B3341CB9B5" type="GBIF-Dataset">271f8735-1aa3-4f63-a742-aef356f505e1</mods:identifier>
<mods:identifier id="D650249C1FCBC87712F3BB69B7032170" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="53A4292D38437FE9B4A58BD959FB6F6D" type="Zenodo-Dep">3527097</mods:identifier>
<mods:identifier id="D7A5639E7B1140DF83257E34664BEC3F" type="ZooBank">76349E63-14F4-4E12-88B6-DCEC6F667694</mods:identifier>
</mods:mods>
<treatment id="2827A302FF96C73E0385DBFE7679FDD7" ID-GBIF-Taxon="160126644" LSID="urn:lsid:plazi:treatment:2827A302FF96C73E0385DBFE7679FDD7" httpUri="http://treatment.plazi.org/id/2827A302FF96C73E0385DBFE7679FDD7" lastPageId="2" lastPageNumber="85" pageId="1" pageNumber="84">
<subSubSection id="E894419FFF96C73D0385DBFE70D3FF14" box="[151,749,225,252]" pageId="1" pageNumber="84" type="nomenclature">
<paragraph id="A0311214FF96C73D0385DBFE70D3FF14" blockId="1.[151,749,225,287]" box="[151,749,225,252]" pageId="1" pageNumber="84">
<heading id="FB79A578FF96C73D0385DBFE70D3FF14" bold="true" box="[151,749,225,252]" fontSize="11" level="1" pageId="1" pageNumber="84" reason="1">
<emphasis id="92FACE06FF96C73D0385DBFE70D3FF14" bold="true" box="[151,749,225,252]" pageId="1" pageNumber="84">
<taxonomicName id="678E6997FF96C73D0385DBFE70BDFF13" ID-CoL="8XV29" authority="Yin &amp; Jantarit" authorityName="Yin &amp; Jantarit" authorityYear="2019" box="[151,643,225,252]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="phuphaphet" status="sp. nov.">
<emphasis id="92FACE06FF96C73D0385DBFE73FFFF14" bold="true" box="[151,449,225,252]" italics="true" pageId="1" pageNumber="84">Araneibatrus phuphaphet</emphasis>
Yin &amp; Jantarit
</taxonomicName>
,
<taxonomicNameLabel id="89C9737DFF96C73D0182DBFD70D3FF14" box="[656,749,226,252]" pageId="1" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E894419FFF96C73D0385DA1A7335FEF7" box="[151,267,261,287]" pageId="1" pageNumber="84" type="description">
<paragraph id="A0311214FF96C73D0385DA1A7335FEF7" blockId="1.[151,749,225,287]" box="[151,267,261,287]" pageId="1" pageNumber="84">
(
<figureCitation id="38B50E91FF96C73D038DDA1A733CFEF7" box="[159,258,261,287]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1967,1992]" captionStartId-1="3.[151,250,905,930]" captionStartId-2="3.[151,250,1979,2004]" captionTargetBox-0="[157,1429,634,1872]" captionTargetBox-1="[151,1433,174,885]" captionTargetBox-2="[161,1426,1017,1960]" captionTargetId-0="figure@2.[151,1436,626,1874]" captionTargetId-1="figure@3.[151,1436,174,886]" captionTargetId-2="figure@3.[161,1426,1017,1960]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURE 1. Habitus (A, C) and head (B, D) of Araneibatrus phuphaphet sp. nov., lateral. Note the female possesses distinctly smaller eyes than male. Scale bar: 0.5 mm for A, C; 0.2 mm for B, D." captionText-1="FIGURE 2. Diagnostic features of Araneibatrus phuphaphet sp. nov. male. A. Head dorsum and pronotum. B. Profemur. C, D. Aedeagus, dorsal (C) and lateral (D) (C’ shows apex of ventral lobe in ventral view). Abbreviations: dl = dorsal lobe; pm = paramere; vl = ventral lobe. Scale bar: 0.2 mm." captionText-2="FIGURE 3. Habitus in situ of Araneibatrus phuphaphet sp. nov. (A, B), and general cave environment (C, D) where the new species was collected." figureDoi-0="http://doi.org/10.5281/zenodo.3527099" figureDoi-1="http://doi.org/10.5281/zenodo.3527101" figureDoi-2="http://doi.org/10.5281/zenodo.3527103" httpUri-0="https://zenodo.org/record/3527099/files/figure.png" httpUri-1="https://zenodo.org/record/3527101/files/figure.png" httpUri-2="https://zenodo.org/record/3527103/files/figure.png" pageId="1" pageNumber="84">Figs 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E894419FFF96C73D0385DA517674FDF3" pageId="1" pageNumber="84" type="materials_examined">
<paragraph id="A0311214FF96C73D0385DA517674FDF3" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="84">
<emphasis id="92FACE06FF96C73D0385DA51737AFE80" bold="true" box="[151,324,333,360]" pageId="1" pageNumber="84">Type material.</emphasis>
<materialsCitation id="10E61849FF96C73D0258DA51767CFE47" ID-GBIF-Occurrence="2446790358" collectingDate="2019-02-08" collectionCode="NHM-PSU" collectorName="Jantarit, S &amp; Nilsai, A. &amp; Jantapaso, K." country="Thailand" county="Manang Dist." latitude="7.1264" location="Khao Banthat Wildlife Sanctuary" longLatPrecision="7" longitude="99.997" municipality="Phu Pha Phet Cave" pageId="1" pageNumber="84" specimenCount="1" specimenCount-male="1" stateProvince="Satun" typeStatus="holotype">
<emphasis id="92FACE06FF96C73D0258DA517058FE80" bold="true" box="[330,614,333,360]" pageId="1" pageNumber="84">
<typeStatus id="7F35ACB6FF96C73D0258DA517385FE80" box="[330,443,334,360]" pageId="1" pageNumber="84" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="D8995284FF96C73D02D4DA52705FFE80" box="[454,609,333,360]" name="Thailand" pageId="1" pageNumber="84">THAILAND</collectingCountry>
:
</emphasis>
<specimenCount id="B688D99DFF96C73D017FDA5170BBFE80" box="[621,645,334,360]" pageId="1" pageNumber="84" type="male">♂</specimenCount>
: ‘THAILAND:
<collectingRegion id="624ADCF6FF96C73D002ADA527185FE80" box="[824,955,333,360]" country="Thailand" name="Satun" pageId="1" pageNumber="84">Satun Prov.</collectingRegion>
,
<collectingCounty id="49506A98FF96C73D00D7DA517661FE8F" box="[965,1119,333,360]" pageId="1" pageNumber="84">Manang Dist.</collectingCounty>
,
<collectingMunicipality id="4055886EFF96C73D0778DA527700FE8F" box="[1130,1342,333,359]" pageId="1" pageNumber="84">Phu Pha Phet Cave</collectingMunicipality>
(sample # THA_SJ_STN10),
<location id="A55144CFFF96C73D0296DA6E70C4FE63" LSID="urn:lsid:plazi:treatment:2827A302FF96C73E0385DBFE7679FDD7:A55144CFFF96C73D0296DA6E70C4FE63" box="[388,762,369,395]" country="Thailand" county="Manang Dist." latitude="7.1264" longLatPrecision="7" longitude="99.997" municipality="Phu Pha Phet Cave" name="Khao Banthat Wildlife Sanctuary" pageId="1" pageNumber="84" stateProvince="Satun">Khao Banthat Wildlife Sanctuary</location>
,
<geoCoordinate id="C5BA74D3FF96C73D0014DA6E71B8FE64" box="[774,902,369,396]" degrees="99.9970" direction="east" orientation="longitude" pageId="1" pageNumber="84" precision="5" value="99.997">99.9970° E</geoCoordinate>
,
<geoCoordinate id="C5BA74D3FF96C73D0080DA6E7638FE64" box="[914,1030,369,396]" degrees="7.1264" direction="north" orientation="latitude" pageId="1" pageNumber="84" precision="5" value="7.1264">7.1264° N</geoCoordinate>
(entrance of the cave), ground floor of deep area,
<date id="D43034D4FF96C73D0224DA8A73A2FE47" box="[310,412,405,431]" pageId="1" pageNumber="84" value="2019-02-08">
<collectingDate id="C474CD3CFF96C73D0224DA8A73A2FE47" box="[310,412,405,431]" pageId="1" pageNumber="84" value="2019-02-08">8.ii.2019</collectingDate>
</date>
,
<collectorName id="0D7B77C2FF96C73D02B9DA8A701EFE47" box="[427,544,405,431]" pageId="1" pageNumber="84">Jantarit, S</collectorName>
.
<collectorName id="0D7B77C2FF96C73D013CDA8A70A2FE58" box="[558,668,405,432]" pageId="1" pageNumber="84">Nilsai, A.</collectorName>
and
<collectorName id="0D7B77C2FF96C73D01CADA89714FFE58" box="[728,881,406,432]" pageId="1" pageNumber="84">Jantapaso, K.</collectorName>
leg. (
<collectionCode id="C69F8AD1FF96C73D00A7DA8A7604FE47" box="[949,1082,405,431]" pageId="1" pageNumber="84">NHM-PSU</collectionCode>
)
</materialsCitation>
’
<materialsCitation id="10E61849FF96C73D075DDA89711AFE3B" ID-GBIF-Occurrence="2446790355" collectingDate="2019-02-08" collectionCode="THAI-, LAND" collectorName="Jantarit, S &amp; Nilsai, A. &amp; Jantapaso, K." country="Thailand" county="Manang Dist." latitude="7.1264" location="Khao Banthat Wildlife Sanctuary" longLatPrecision="7" longitude="99.997" municipality="Phu Pha Phet Cave" pageId="1" pageNumber="84" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Satun" typeStatus="paratype">
<emphasis id="92FACE06FF96C73D075DDA8976F9FE58" bold="true" box="[1103,1223,406,432]" pageId="1" pageNumber="84">
<typeStatus id="7F35ACB6FF96C73D075DDA8976F9FE58" box="[1103,1223,406,432]" pageId="1" pageNumber="84" type="paratype">Paratypes</typeStatus>
</emphasis>
(24 exx.):
<emphasis id="92FACE06FF96C73D0659DA8A72CEFE3C" bold="true" pageId="1" pageNumber="84">
<collectionCode id="C69F8AD1FF96C73D0659DA8A72D5FE3C" pageId="1" pageNumber="84">THAI- LAND</collectionCode>
:
</emphasis>
<specimenCount id="B688D99DFF96C73D03E4DAA67309FE3C" box="[246,311,441,468]" pageId="1" pageNumber="84" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="B688D99DFF96C73D0250DAA673BDFE3C" box="[322,387,441,468]" pageId="1" pageNumber="84" type="female">4 ♀♀</specimenCount>
: same collecting data as for holotype
</materialsCitation>
;
<materialsCitation id="10E61849FF96C73D003CDAA670A4FE1F" ID-GBIF-Occurrence="2446790357" collectingDate="2019-05-31" collectionCode="THAI-, LAND" collectorName="Jantarit, S &amp; Surakhamhaeng, K. &amp; Jantapaso, K." country="Thailand" county="Manang Dist." latitude="7.1264" location="Khao Banthat Wildlife Sanctuary" longLatPrecision="7" longitude="99.997" municipality="Phu Pha Phet Cave" pageId="1" pageNumber="84" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Satun" typeStatus="paratype">
<specimenCount id="B688D99DFF96C73D003CDAA67150FE3C" box="[814,878,441,468]" pageId="1" pageNumber="84" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="B688D99DFF96C73D0068DAA67184FE3C" box="[890,954,441,468]" pageId="1" pageNumber="84" type="female">4 ♀♀</specimenCount>
: same location, except ‘
<date id="D43034D4FF96C73D07D1DAA67709FE3B" box="[1219,1335,441,467]" pageId="1" pageNumber="84" value="2019-05-31">
<collectingDate id="C474CD3CFF96C73D07D1DAA67709FE3B" box="[1219,1335,441,467]" pageId="1" pageNumber="84" value="2019-05-31">31.v.2019</collectingDate>
</date>
,
<collectorName id="0D7B77C2FF96C73D0651DAA67297FE1F" pageId="1" pageNumber="84">Jantarit, S</collectorName>
.
<collectorName id="0D7B77C2FF96C73D03A4DAC273AAFE10" box="[182,404,477,504]" pageId="1" pageNumber="84">Surakhamhaeng, K.</collectorName>
and
<collectorName id="0D7B77C2FF96C73D02DEDAC1705DFE10" box="[460,611,478,504]" pageId="1" pageNumber="84">Jantapaso, K.</collectorName>
leg.
</materialsCitation>
’;
<materialsCitation id="10E61849FF96C73D01BFDAC27679FDF3" ID-GBIF-Occurrence="2446790356" collectingDate="2019-06-23" collectionCode="NHM-PSU, SNUC" collectorName="Jantarit, S. &amp; Jantapaso, K." country="Thailand" county="Manang Dist." latitude="7.1264" location="Khao Banthat Wildlife Sanctuary" longLatPrecision="7" longitude="99.997" municipality="Phu Pha Phet Cave" pageId="1" pageNumber="84" specimenCount="32" specimenCount-female="18" specimenCount-male="14" stateProvince="Satun" typeStatus="paratype">
<specimenCount id="B688D99DFF96C73D01BFDAC270D1FE10" box="[685,751,477,504]" pageId="1" pageNumber="84" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="B688D99DFF96C73D01EEDAC27100FE10" box="[764,830,477,504]" pageId="1" pageNumber="84" type="female">5 ♀♀</specimenCount>
: same location, except ‘
<date id="D43034D4FF96C73D0759DAC276F4FE1F" box="[1099,1226,477,503]" pageId="1" pageNumber="84" value="2019-06-23">
<collectingDate id="C474CD3CFF96C73D0759DAC276F4FE1F" box="[1099,1226,477,503]" pageId="1" pageNumber="84" value="2019-06-23">23.vi.2019</collectingDate>
</date>
,
<collectorName id="0D7B77C2FF96C73D07C5DAC27770FE1F" box="[1239,1358,477,503]" pageId="1" pageNumber="84">Jantarit, S.</collectorName>
Prom- dam, R. and
<collectorName id="0D7B77C2FF96C73D0234D91D7382FDF4" box="[294,444,514,540]" pageId="1" pageNumber="84">Jantapaso, K.</collectorName>
leg.’ (
<specimenCount id="B688D99DFF96C73D0111D91E707AFDF4" box="[515,580,513,540]" pageId="1" pageNumber="84" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="B688D99DFF96C73D0142D91E70B1FDF4" box="[592,655,513,540]" pageId="1" pageNumber="84" type="female">9 ♀♀</specimenCount>
in
<collectionCode id="C69F8AD1FF96C73D01A1D91E710AFDF3" box="[691,820,513,539]" pageId="1" pageNumber="84">NHM-PSU</collectionCode>
,
<specimenCount id="B688D99DFF96C73D002DD91E71BEFDF4" box="[831,896,513,540]" pageId="1" pageNumber="84" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="B688D99DFF96C73D009ED91E71F5FDF4" box="[908,971,513,540]" pageId="1" pageNumber="84" type="female">4 ♀♀</specimenCount>
in
<collectionCode id="C69F8AD1FF96C73D00FDD91E7601FDF3" box="[1007,1087,513,539]" pageId="1" pageNumber="84">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E894419FFF96C73D03D5D93A700EFD1B" pageId="1" pageNumber="84" type="diagnosis">
<paragraph id="A0311214FF96C73D03D5D93A700EFD1B" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="84">
<emphasis id="92FACE06FF96C73D03D5D93A737CFDA8" bold="true" box="[199,322,549,576]" pageId="1" pageNumber="84">Diagnosis.</emphasis>
Body length
<quantity id="6776BFF1FF96C73D02CDD93A70B9FDA8" box="[479,647,549,576]" metricMagnitude="-3" metricUnit="m" metricValue="2.19" metricValueMax="2.26" metricValueMin="2.12" pageId="1" pageNumber="84" unit="mm" value="2.19" valueMax="2.26" valueMin="2.12">2.12–2.26 mm</quantity>
. Vertex with elongate median carina, lateral postantennal carinae absent; clypeus short, less than 1/5 total head length. Pronotum lacking sulci or carinae, with large antebasal spines, sides dentate. Elytra lacking discal striae; anterolateral margins non-emarginate. Metaventrite with narrow notch at middle of posterior margin. Tergite IV with two and sternite IV with four basolateral foveae, each lacking mediobasal fovea. Male antennal club unmodified; profemur ventrally protuberant at basal 2/5; aedeagus asymmetric, with elongate globose basal capsule.
</paragraph>
</subSubSection>
<subSubSection id="E894419FFF96C73D03D5D9E276B1F8EF" pageId="1" pageNumber="84" type="description">
<paragraph id="A0311214FF96C73D03D5D9E2716EFA94" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="84">
<emphasis id="92FACE06FF96C73D03D5D9E27367FCF0" bold="true" box="[199,345,765,792]" pageId="1" pageNumber="84">Description.</emphasis>
Male (
<figureCitation id="38B50E91FF96C73D02A2D9E27033FCFF" box="[432,525,765,791]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1992]" captionTargetBox="[157,1429,634,1872]" captionTargetId="figure@2.[151,1436,626,1874]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A, C) and head (B, D) of Araneibatrus phuphaphet sp. nov., lateral. Note the female possesses distinctly smaller eyes than male. Scale bar: 0.5 mm for A, C; 0.2 mm for B, D." figureDoi="http://doi.org/10.5281/zenodo.3527099" httpUri="https://zenodo.org/record/3527099/files/figure.png" pageId="1" pageNumber="84">Fig. 1A</figureCitation>
). Length
<quantity id="6776BFF1FF96C73D0192D9E27114FCF0" box="[640,810,765,792]" metricMagnitude="-3" metricUnit="m" metricValue="2.245" metricValueMax="2.26" metricValueMin="2.23" pageId="1" pageNumber="84" unit="mm" value="2.245" valueMax="2.26" valueMin="2.23">2.23–2.26 mm</quantity>
. Body uniformly reddish-brown, mouthparts and tarsi lighter in color. Head (
<figureCitation id="38B50E91FF96C73D0286D83E73D2FCD3" box="[404,492,801,827]" captionStart="FIGURE 2" captionStartId="3.[151,250,905,930]" captionTargetBox="[151,1433,174,885]" captionTargetId="figure@3.[151,1436,174,886]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Araneibatrus phuphaphet sp. nov. male. A. Head dorsum and pronotum. B. Profemur. C, D. Aedeagus, dorsal (C) and lateral (D) (C’ shows apex of ventral lobe in ventral view). Abbreviations: dl = dorsal lobe; pm = paramere; vl = ventral lobe. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3527101" httpUri="https://zenodo.org/record/3527101/files/figure.png" pageId="1" pageNumber="84">Fig. 2A</figureCitation>
) slightly longer than wide, length from clypeal anterior margin to basal construction
<quantity id="6776BFF1FF96C73D0385D85A72C0FC88" box="[151,254,837,864]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="84" unit="mm" value="0.46">0.46 mm</quantity>
, width across eyes
<quantity id="6776BFF1FF96C73D02CBD85A7041FC88" box="[473,639,837,864]" metricMagnitude="-4" metricUnit="m" metricValue="4.45" metricValueMax="4.6" metricValueMin="4.3" pageId="1" pageNumber="84" unit="mm" value="0.445" valueMax="0.46" valueMin="0.43">0.43–0.46 mm</quantity>
; dorsum sparsely and finely punctate; vertex with elongate median carina extending from level of vertexal foveae to antennal tubercles, lacking short mediobasal carina; lacking lateral postantennal carinae; clypeus relatively short, slightly less than one-fifth of total head length; each eye composed of about 12 facets (
<figureCitation id="38B50E91FF96C73D0243D8AE7399FC23" box="[337,423,945,971]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1992]" captionTargetBox="[157,1429,634,1872]" captionTargetId="figure@2.[151,1436,626,1874]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A, C) and head (B, D) of Araneibatrus phuphaphet sp. nov., lateral. Note the female possesses distinctly smaller eyes than male. Scale bar: 0.5 mm for A, C; 0.2 mm for B, D." figureDoi="http://doi.org/10.5281/zenodo.3527099" httpUri="https://zenodo.org/record/3527099/files/figure.png" pageId="1" pageNumber="84">Fig. 1B</figureCitation>
); antennal club formed by three apical antennomeres, which are slightly broader than proceeding antennomeres; antennomere 11 asymmetrically narrowed at apical 1/3. Pronotum (
<figureCitation id="38B50E91FF96C73D0781D8CA76D2FC07" box="[1171,1260,981,1007]" captionStart="FIGURE 2" captionStartId="3.[151,250,905,930]" captionTargetBox="[151,1433,174,885]" captionTargetId="figure@3.[151,1436,174,886]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Araneibatrus phuphaphet sp. nov. male. A. Head dorsum and pronotum. B. Profemur. C, D. Aedeagus, dorsal (C) and lateral (D) (C’ shows apex of ventral lobe in ventral view). Abbreviations: dl = dorsal lobe; pm = paramere; vl = ventral lobe. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3527101" httpUri="https://zenodo.org/record/3527101/files/figure.png" pageId="1" pageNumber="84">Fig. 2A</figureCitation>
) approximately hexagonal, wider than head and slightly wider than long, length along midline
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, maximum width
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; disc sparsely and finely punctate, lacking sulcus or carina, with two large, elongate antebasal spines; sides widest at apical 2/5, distinctly dentate. Elytra slightly longer than wide, length along suture
<quantity id="6776BFF1FF96C73D076BDF5E7720FBB4" box="[1145,1310,1089,1116]" metricMagnitude="-4" metricUnit="m" metricValue="6.799999999999999" metricValueMax="6.8999999999999995" metricValueMin="6.7" pageId="1" pageNumber="84" unit="mm" value="0.6799999999999999" valueMax="0.69" valueMin="0.67">0.67–0.69 mm</quantity>
, maximum width
<quantity id="6776BFF1FF96C73D03CFDF7A73BCFB68" box="[221,386,1125,1152]" metricMagnitude="-4" metricUnit="m" metricValue="6.45" metricValueMax="6.5" metricValueMin="6.4" pageId="1" pageNumber="84" unit="mm" value="0.645" valueMax="0.65" valueMin="0.64">0.64–0.65 mm</quantity>
, lacking discal striae, anterolateral margins evenly curved, non-emarginate. Mesoventrite with lateral foveae separated from median foveae. Metaventrite lacking protuberances; posterior margin with narrow notch at middle. Profemur (
<figureCitation id="38B50E91FF96C73D02DCDFB2701BFB2F" box="[462,549,1197,1223]" captionStart="FIGURE 2" captionStartId="3.[151,250,905,930]" captionTargetBox="[151,1433,174,885]" captionTargetId="figure@3.[151,1436,174,886]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Araneibatrus phuphaphet sp. nov. male. A. Head dorsum and pronotum. B. Profemur. C, D. Aedeagus, dorsal (C) and lateral (D) (C’ shows apex of ventral lobe in ventral view). Abbreviations: dl = dorsal lobe; pm = paramere; vl = ventral lobe. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3527101" httpUri="https://zenodo.org/record/3527101/files/figure.png" pageId="1" pageNumber="84">Fig. 2B</figureCitation>
) ventrally protuberant at basal 2/5. Tergite IV with complete and oblique inner marginal carinae, with complete basal impression ended by two basolateral foveae. Sternite IV with two pairs of basolateral foveae covered under dense setae. Length of aedeagus (
<figureCitation id="38B50E91FF96C73D0098DFEA762FFAE7" box="[906,1041,1269,1295]" captionStart="FIGURE 2" captionStartId="3.[151,250,905,930]" captionTargetBox="[151,1433,174,885]" captionTargetId="figure@3.[151,1436,174,886]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Araneibatrus phuphaphet sp. nov. male. A. Head dorsum and pronotum. B. Profemur. C, D. Aedeagus, dorsal (C) and lateral (D) (C’ shows apex of ventral lobe in ventral view). Abbreviations: dl = dorsal lobe; pm = paramere; vl = ventral lobe. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3527101" httpUri="https://zenodo.org/record/3527101/files/figure.png" pageId="1" pageNumber="84">Fig. 2C, 2D</figureCitation>
)
<quantity id="6776BFF1FF96C73D0733DFEA76B6FAF8" box="[1057,1160,1269,1296]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="84" unit="mm" value="0.44">0.44 mm</quantity>
; median lobe with stout, elongate basal capsule and large foramen; ventral lobe expanded along left side dorsal-ventrally (natural position) and curved ventrad; dorsal lobe deeply split at base and comprised of two elongate sclerites; parameres attached to apex of basal capsule, reduced to semi-membranous structure.
</paragraph>
<paragraph id="A0311214FF96C73D03D5DE9A70A4F9E4" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="84">
Female. Similar to male in external morphology, except each eye composed of about 6 facets (
<figureCitation id="38B50E91FF96C73D061BDE9A775FFA77" box="[1289,1377,1413,1439]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1992]" captionTargetBox="[157,1429,634,1872]" captionTargetId="figure@2.[151,1436,626,1874]" captionTargetPageId="2" captionText="FIGURE 1. Habitus (A, C) and head (B, D) of Araneibatrus phuphaphet sp. nov., lateral. Note the female possesses distinctly smaller eyes than male. Scale bar: 0.5 mm for A, C; 0.2 mm for B, D." figureDoi="http://doi.org/10.5281/zenodo.3527099" httpUri="https://zenodo.org/record/3527099/files/figure.png" pageId="1" pageNumber="84">Fig. 1D</figureCitation>
) and profemur simple. Measurements (as for male): body
<quantity id="6776BFF1FF96C73D01E0DEB671A7FA2C" box="[754,921,1449,1476]" metricMagnitude="-3" metricUnit="m" metricValue="2.17" metricValueMax="2.22" metricValueMin="2.12" pageId="1" pageNumber="84" unit="mm" value="2.17" valueMax="2.22" valueMin="2.12">2.12–2.22 mm</quantity>
, length /width of head 0.43–0.48 /
<quantity id="6776BFF1FF96C73D063EDEB672FBFA00" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.4" metricValueMin="3.9" pageId="1" pageNumber="84" unit="mm" value="0.415" valueMax="0.44" valueMin="0.39">0.39–0.44 mm</quantity>
, length / width of pronotum 0.47 /
<quantity id="6776BFF1FF96C73D0143DED270C9FA00" box="[593,759,1485,1512]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="1" pageNumber="84" unit="mm" value="0.5" valueMax="0.52" valueMin="0.48">0.48–0.52 mm</quantity>
, length / width of elytra 0.62–0.65 /
<quantity id="6776BFF1FF96C73D0787DED27705FA00" box="[1173,1339,1485,1512]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="6.1" metricValueMin="5.8999999999999995" pageId="1" pageNumber="84" unit="mm" value="0.6" valueMax="0.61" valueMin="0.59">0.59–0.61 mm</quantity>
, length / width of abdomen 0.60–0.62 /
<quantity id="6776BFF1FF96C73D02E2DEEE70A8F9E4" box="[496,662,1521,1548]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" metricValueMax="6.0" metricValueMin="5.6" pageId="1" pageNumber="84" unit="mm" value="0.58" valueMax="0.6" valueMin="0.56">0.56–0.60 mm</quantity>
.
</paragraph>
<paragraph id="A0311214FF96C73D03D5DD0A76B1F8EF" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="84">
<emphasis id="92FACE06FF96C73D03D5DD0A738EF9D8" bold="true" box="[199,432,1557,1584]" pageId="1" pageNumber="84">Comparative notes.</emphasis>
The genus
<taxonomicName id="678E6997FF96C73D0120DD0A70F8F9C7" authorityName="Yin &amp; Li" authorityYear="2010" box="[562,710,1557,1583]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="92FACE06FF96C73D0120DD0A70F8F9C7" box="[562,710,1557,1583]" italics="true" pageId="1" pageNumber="84">Araneibatrus</emphasis>
</taxonomicName>
now includes seven species from southern
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(5 spp.), northern
<collectingCountry id="D8995284FF96C73D03D1DD2572C7F9BC" box="[195,249,1594,1620]" name="Laos" pageId="1" pageNumber="84">Laos</collectingCountry>
(
<specimenCount id="B688D99DFF96C73D021EDD26737FF9BC" box="[268,321,1593,1620]" pageId="1" pageNumber="84" type="generic">1 sp.</specimenCount>
), and southern
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(
<specimenCount id="B688D99DFF96C73D0177DD2670A5F9BC" box="[613,667,1593,1620]" pageId="1" pageNumber="84" type="generic">1 sp.</specimenCount>
). Using the key provided by
<bibRefCitation id="C41F6FE5FF96C73D00F1DD2676A0F9BC" author="Yin, Z. - W. &amp; Jiang, R. - X. &amp; Steiner, H." box="[995,1182,1593,1620]" pageId="1" pageNumber="84" pagination="475 - 494" refId="ref2326" refString="Yin, Z. - W., Jiang, R. - X. &amp; Steiner, H. (2016) Revision of the genus Araneibatrus (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4097 (4), 475 - 494. https: // doi. org / 10.11646 / zootaxa. 4097.4.2" type="journal article" year="2016">
Yin
<emphasis id="92FACE06FF96C73D0701DD25767BF9BB" box="[1043,1093,1593,1619]" italics="true" pageId="1" pageNumber="84">et al</emphasis>
. (2016)
</bibRefCitation>
the new species could be keyed out at couplet 3, near
<taxonomicName id="678E6997FF96C73D02E4DD417142F99F" authority="Yin, Li &amp; Zhao 2010" authorityName="Yin, Li &amp; Zhao" authorityYear="2010" box="[502,892,1629,1656]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="92FACE06FF96C73D02E4DD417039F990" box="[502,519,1630,1656]" italics="true" pageId="1" pageNumber="84">A</emphasis>
.
<emphasis id="92FACE06FF96C73D0107DD4270BBF99F" box="[533,645,1629,1655]" italics="true" pageId="1" pageNumber="84">gracilipes</emphasis>
Yin, Li &amp; Zhao 2010
</taxonomicName>
. These two species share a short clypeus and the lack of lateral postantennal carinae of the head, but can be readily distinguished by the presence of large antebasal spines and distinct spinose lateral margins of the pronotum of
<taxonomicName id="678E6997FF96C73D0047DDB971C7F957" authorityName="Yin &amp; Jantarit" authorityYear="2019" box="[853,1017,1701,1728]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="phuphaphet">
<emphasis id="92FACE06FF96C73D0047DDB97158F928" box="[853,870,1702,1728]" italics="true" pageId="1" pageNumber="84">A</emphasis>
.
<emphasis id="92FACE06FF96C73D0067DDBA71C7F957" box="[885,1017,1701,1727]" italics="true" pageId="1" pageNumber="84">phuphaphet</emphasis>
</taxonomicName>
. Beyond that,
<taxonomicName id="678E6997FF96C73D078DDDB9777CF957" authorityName="Yin &amp; Jantarit" authorityYear="2019" box="[1183,1346,1701,1728]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="phuphaphet">
<emphasis id="92FACE06FF96C73D078DDDB9768EF928" box="[1183,1200,1702,1728]" italics="true" pageId="1" pageNumber="84">A</emphasis>
.
<emphasis id="92FACE06FF96C73D07ACDDBA777CF957" box="[1214,1346,1701,1727]" italics="true" pageId="1" pageNumber="84">phuphaphet</emphasis>
</taxonomicName>
is quite unique among the members of the genus in possessing conspicuously long discal spines of the pronotum, sexually protuberant profemora of the male, as well as a unique, complex structure of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="E894419FFF96C73E03D5DC0E72ECFEF7" lastPageId="2" lastPageNumber="85" pageId="1" pageNumber="84" type="biology_ecology">
<paragraph id="A0311214FF96C73E03D5DC0E72ECFEF7" blockId="1.[151,1437,333,2016]" lastBlockId="2.[151,1437,152,575]" lastPageId="2" lastPageNumber="85" pageId="1" pageNumber="84">
<emphasis id="92FACE06FF96C73D03D5DC0E7319F8C4" bold="true" box="[199,295,1809,1836]" pageId="1" pageNumber="84">Biology.</emphasis>
<taxonomicName id="678E6997FF96C73D023DDC0E7071F8C3" authorityName="Yin &amp; Jantarit" authorityYear="2019" box="[303,591,1809,1835]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="phuphaphet">
<emphasis id="92FACE06FF96C73D023DDC0E7071F8C3" box="[303,591,1809,1835]" italics="true" pageId="1" pageNumber="84">Araneibatrus phuphaphet</emphasis>
</taxonomicName>
is a cavernicolous species endemic to Phu Pha Phet Cave. The second author and his team have extensively surveyed and collected materials from almost 100 caves in southern
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, and in total almost 200 caves throughout the country. However, the new species has been only found in this cave. The populations of
<taxonomicName id="678E6997FF96C73D0263DC61702AF87F" authorityName="Yin &amp; Jantarit" authorityYear="2019" box="[369,532,1917,1944]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="phuphaphet">
<emphasis id="92FACE06FF96C73D0263DC6173BCF870" box="[369,386,1918,1944]" italics="true" pageId="1" pageNumber="84">A</emphasis>
.
<emphasis id="92FACE06FF96C73D0282DC62702AF87F" box="[400,532,1917,1943]" italics="true" pageId="1" pageNumber="84">phuphaphet</emphasis>
</taxonomicName>
are rather limited and restricted to the dark zone of certain chambers, with a few individuals being found wandering at the tourist spots. The beetles moved quite fast when disturbed and were usually found roaming around the humid ground floor (
<figureCitation id="38B50E91FF96C73D01CEDCDA715DF837" box="[732,867,1989,2015]" captionStart="FIGURE 3" captionStartId="3.[151,250,1979,2004]" captionTargetBox="[161,1426,1017,1960]" captionTargetId="figure@3.[161,1426,1017,1960]" captionTargetPageId="3" captionText="FIGURE 3. Habitus in situ of Araneibatrus phuphaphet sp. nov. (A, B), and general cave environment (C, D) where the new species was collected." figureDoi="http://doi.org/10.5281/zenodo.3527103" httpUri="https://zenodo.org/record/3527103/files/figure.png" pageId="1" pageNumber="84">Fig. 3A, 3B</figureCitation>
).
<taxonomicName id="678E6997FF96C73D006ADCDA76A9F837" authorityName="Yin &amp; Jantarit" authorityYear="2019" box="[888,1175,1989,2015]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="phuphaphet">
<emphasis id="92FACE06FF96C73D006ADCDA76A9F837" box="[888,1175,1989,2015]" italics="true" pageId="1" pageNumber="84">Araneibatrus phuphaphet</emphasis>
</taxonomicName>
prefers an oligotrophic habitat (without bat guano,
<figureCitation id="38B50E91FF95C73E02DDDB86706BFF5B" box="[463,597,153,179]" captionStart="FIGURE 3" captionStartId="3.[151,250,1979,2004]" captionTargetBox="[161,1426,1017,1960]" captionTargetId="figure@3.[161,1426,1017,1960]" captionTargetPageId="3" captionText="FIGURE 3. Habitus in situ of Araneibatrus phuphaphet sp. nov. (A, B), and general cave environment (C, D) where the new species was collected." figureDoi="http://doi.org/10.5281/zenodo.3527103" httpUri="https://zenodo.org/record/3527103/files/figure.png" pageId="2" pageNumber="85">Fig. 3C, 3D</figureCitation>
) where the air temperature is relatively low (24.4–25.2 °C; at altitude
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), and the cave soil temperature ranges from 24.4–25.0 °C. The humidity is very high at 93%, with the air at 21% oxygen. During our three surveys we rarely found other micro/macro-invertebrates (except for crickets) in this cave.
</paragraph>
</subSubSection>
<subSubSection id="E894419FFF95C73E03D5DA377743FE1F" pageId="2" pageNumber="85" type="discussion">
<paragraph id="A0311214FF95C73E03D5DA377743FE1F" blockId="2.[151,1437,152,575]" pageId="2" pageNumber="85">
<emphasis id="92FACE06FF95C73E03D5DA377371FEAB" bold="true" box="[199,335,296,323]" pageId="2" pageNumber="85">Comments.</emphasis>
The Phu Pha Phet Cave is the largest cave in
<collectingCountry id="D8995284FF95C73E0070DA3671F8FEAB" box="[866,966,297,323]" name="Thailand" pageId="2" pageNumber="85">Thailand</collectingCountry>
(ca. 20 acres with more than 20 separate rooms) and is ranked the fourth largest in the world. The cave developed in limestone of the Ordovician Thung Song Group, one of the oldest sedimentary rock in
<collectingCountry id="D8995284FF95C73E01B4DA6E7133FE63" box="[678,781,369,395]" name="Thailand" pageId="2" pageNumber="85">Thailand</collectingCountry>
. Unfortunately, little attention has been paid to its fauna. Hence, the discovery of a new cavernicolous pselaphine not only emphasizes the biological importance of the cave and
<collectingRegion id="624ADCF6FF95C73E03DADAA67336FE3B" box="[200,264,441,467]" country="Thailand" name="Satun" pageId="2" pageNumber="85">Satun</collectingRegion>
Geopark, but also generates major implications in the development of the conservation policy and ecotourism plan for the area, as well as raising awareness about the unique subterranean environment of the region.
</paragraph>
</subSubSection>
<subSubSection id="E894419FFF95C73E03D5D91F76A4FDF3" box="[199,1178,512,539]" pageId="2" pageNumber="85" type="distribution">
<paragraph id="A0311214FF95C73E03D5D91F76A4FDF3" blockId="2.[151,1437,152,575]" box="[199,1178,512,539]" pageId="2" pageNumber="85">
<emphasis id="92FACE06FF95C73E03D5D91F735FFDF3" bold="true" box="[199,353,512,539]" pageId="2" pageNumber="85">Distribution.</emphasis>
Only known from the
<typeStatus id="7F35ACB6FF95C73E0172D91E70AEFDF3" box="[608,656,513,539]" pageId="2" pageNumber="85">type</typeStatus>
locality in
<collectingRegion id="624ADCF6FF95C73E001DD91E7182FDF3" box="[783,956,513,539]" country="Thailand" name="Satun" pageId="2" pageNumber="85">Satun Province</collectingRegion>
, southern
<collectingCountry id="D8995284FF95C73E073CD91E76ABFDF3" box="[1070,1173,513,539]" name="Thailand" pageId="2" pageNumber="85">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E894419FFF95C73E03D5D93A7679FDD7" box="[199,1095,549,575]" pageId="2" pageNumber="85" type="etymology">
<paragraph id="A0311214FF95C73E03D5D93A7679FDD7" blockId="2.[151,1437,152,575]" box="[199,1095,549,575]" pageId="2" pageNumber="85">
<emphasis id="92FACE06FF95C73E03D5D93A7370FDD7" bold="true" box="[199,334,549,575]" pageId="2" pageNumber="85">Etymology.</emphasis>
The new specific epithet ‘
<taxonomicName id="678E6997FF95C73E0165D93A70C5FDD7" authorityName="Yin &amp; Jantarit" authorityYear="2019" box="[631,763,549,575]" class="Insecta" family="Staphylinidae" genus="Araneibatrus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="phuphaphet">
<emphasis id="92FACE06FF95C73E0165D93A70C5FDD7" box="[631,763,549,575]" italics="true" pageId="2" pageNumber="85">phuphaphet</emphasis>
</taxonomicName>
’ is based on the
<typeStatus id="7F35ACB6FF95C73E00A4D93A71D8FDD7" box="[950,998,549,575]" pageId="2" pageNumber="85">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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