treatments-xml/data/23/52/32/235232213850FFDEFF2CF986FD3AF841.xml
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<document id="3649AE1C42E3A5A69C523B873F2D0D8D" ID-DOI="10.5281/zenodo.293981" ID-GBIF-Dataset="67c09726-7f1d-4e2a-b9f7-0b851b6f7cef" ID-ISSN="1175-5326" ID-Zenodo-Dep="293981" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460225559816" checkinUser="plazi" docAuthor="Grismer, Lee" docDate="2010" docId="235232213850FFDEFF2CF986FD3AF841" docLanguage="en" docName="zt02475p063.pdf" docOrigin="Zootaxa 2475" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cnemaspis laoensis Grismer, 2010, sp. nov." docType="treatment" docVersion="6" lastPageNumber="59" masterDocId="DF6B4A593852FFDAFFBBFFEBFF81FFF3" masterDocTitle="The first record of the genus Cnemaspis Strauch (Squamata: Gekkonidae) from Laos with the description of a new species" masterLastPageNumber="63" masterPageNumber="55" pageNumber="57" updateTime="1698238228737" updateUser="plazi">
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<mods:title id="FA67AA55826A9BE944199DE611AAFE42">The first record of the genus Cnemaspis Strauch (Squamata: Gekkonidae) from Laos with the description of a new species</mods:title>
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<mods:namePart id="9CDBAAC3F91E8628B3B2530664B95912">Grismer, Lee</mods:namePart>
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<mods:date id="FB4C65096122F4642A0FBE2AE0623651">2010</mods:date>
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<treatment id="235232213850FFDEFF2CF986FD3AF841" ID-DOI="http://doi.org/10.5281/zenodo.6196491" ID-GBIF-Taxon="119386186" ID-Zenodo-Dep="6196491" LSID="urn:lsid:plazi:treatment:235232213850FFDEFF2CF986FD3AF841" httpUri="http://treatment.plazi.org/id/235232213850FFDEFF2CF986FD3AF841" lastPageId="4" lastPageNumber="59" pageId="2" pageNumber="57">
<subSubSection id="E3E1D0BC3850FFD8FF2CF986FEDCF959" pageId="2" pageNumber="57" type="nomenclature">
<paragraph id="AB4483373850FFD8FF2CF986FE69F974" blockId="2.[151,488,1645,1743]" box="[151,488,1645,1671]" pageId="2" pageNumber="57">
<heading id="F00C345B3850FFD8FF2CF986FE69F974" bold="true" box="[151,488,1645,1671]" fontSize="11" level="1" pageId="2" pageNumber="57" reason="1">
<emphasis id="998F5F253850FFD8FF2CF986FE69F974" bold="true" box="[151,488,1645,1671]" pageId="2" pageNumber="57">
<taxonomicName id="6CFBF8B43850FFD8FF2CF986FE05F974" box="[151,388,1645,1671]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="57" phylum="Chordata" rank="species" species="laoensis" status="sp. nov.">
<emphasis id="998F5F253850FFD8FF2CF986FE05F974" bold="true" box="[151,388,1645,1671]" italics="true" pageId="2" pageNumber="57">Cnemaspis laoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82BCE25E3850FFD8FE30F986FE69F974" box="[395,488,1645,1671]" pageId="2" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="AB4483373850FFD8FF2CF97BFEDCF959" blockId="2.[151,488,1645,1743]" box="[151,349,1680,1706]" pageId="2" pageNumber="57">
<collectingCountry id="D3ECC3A73850FFD8FF2CF97BFF44F959" box="[151,197,1680,1706]" name="Laos" pageId="2" pageNumber="57">Lao</collectingCountry>
Rock Gecko
</paragraph>
</subSubSection>
<subSubSection id="E3E1D0BC3850FFD8FF2CF95EFF7BF93C" box="[151,250,1717,1743]" pageId="2" pageNumber="57" type="description">
<paragraph id="AB4483373850FFD8FF2CF95EFF7BF93C" blockId="2.[151,488,1645,1743]" box="[151,250,1717,1743]" pageId="2" pageNumber="57">
<figureCitation id="33C09FB23850FFD8FF2CF95EFF7BF93C" box="[151,250,1717,1743]" captionStart="FIGURE 2" captionStartId="3.[151,255,1287,1311]" captionTargetBox="[151,1434,421,1262]" captionTargetId="figure@3.[151,1436,421,1263]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cnemaspis laoensis (THNHM 12433)." httpUri="https://zenodo.org/record/293983/files/figure.png" pageId="2" pageNumber="57">Figure 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E3E1D0BC3850FFD8FF2CF914FCAAF8CC" pageId="2" pageNumber="57" type="materials_examined">
<paragraph id="AB4483373850FFD8FF2CF914FCAAF8CC" blockId="2.[151,1437,1791,2004]" pageId="2" pageNumber="57">
<emphasis id="998F5F253850FFD8FF2CF914FE95F8EA" bold="true" box="[151,276,1791,1817]" pageId="2" pageNumber="57">
<typeStatus id="74403D953850FFD8FF2CF914FE95F8EA" box="[151,276,1791,1817]" pageId="2" pageNumber="57" type="holotype">Holotype</typeStatus>
</emphasis>
. Adult female (
<collectionCode id="CDEA1BF23850FFD8FE56F8EBFDE2F8E9" box="[493,611,1792,1818]" pageId="2" pageNumber="57">THNHM</collectionCode>
12433) collected at Dong Phu Vieng National Protected Area, Savannakhet,
<collectingCountry id="D3ECC3A73850FFD8FE80F8CEFEF5F8CC" box="[315,372,1829,1855]" name="Laos" pageId="2" pageNumber="57">Laos</collectingCountry>
by Tanya Chan-ard on
<date id="DF45A5F73850FFD8FD3CF8CEFCA6F8CC" box="[647,807,1829,1855]" pageId="2" pageNumber="57" value="1998-06-18">18 June 1998</date>
.
</paragraph>
</subSubSection>
<subSubSection id="E3E1D0BC3850FFD9FF7DF8A1FB22FE9F" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57" type="diagnosis">
<paragraph id="AB4483373850FFD9FF7DF8A1FB22FE9F" blockId="2.[151,1437,1791,2004]" lastBlockId="3.[151,1437,152,364]" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57">
<emphasis id="998F5F253850FFD8FF7DF8A1FEBEF897" bold="true" box="[198,319,1866,1892]" pageId="2" pageNumber="57">Diagnosis</emphasis>
. Adult female reaching 40.9 mm SVL, adult males unknown; relatively large, rugose scales on rostrum; nine supralabials; seven infralabials; subtibials, ventrals, and subcaudals smooth; forearm scales and dorsal tubercles keeled; body scales raised, smooth; 22 paravertebral tubercles; tubercles on flanks not linearly arranged; ventrolateral and lateral rows of caudal tubercles absent; caudal tubercles do not encircle tail; tubercles in lateral, caudal furrow; median row of slightly enlarged, smooth subcaudals; no pore-bearing scales; two or three postcloacal tubercles on each side of tail; shield-like subtibials and enlarged, submetatarsals absent; 29 subdigital lamellae on fourth toe; no dark, longitudinal gular markings or blotches; no dark patch on shoulder or neck enclosing a white to yellow ocellus; no yellow to white, prescapular crescent or transverse bars on flanks; and no orange tail, feet, and hands. These differences are summarized across all Southeast Asian species of
<taxonomicName id="6CFBF8B43851FFD9FDF7FEB9FD4CFE98" box="[588,717,338,363]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="998F5F253851FFD9FDF7FEB9FD4CFE98" box="[588,717,338,363]" italics="true" pageId="3" pageNumber="58">Cnemaspis</emphasis>
</taxonomicName>
in Grismer
<emphasis id="998F5F253851FFD9FCE1FEB9FC14FE98" box="[858,917,338,363]" italics="true" pageId="3" pageNumber="58">et al.</emphasis>
(2010:Tables 1 and 2).
</paragraph>
</subSubSection>
<caption id="FF84D3BF3851FFD9FF2CFAECFCCAFAED" box="[151,843,1287,1311]" httpUri="https://zenodo.org/record/293983/files/figure.png" pageId="3" pageNumber="58" targetBox="[151,1434,421,1262]" targetPageId="3">
<paragraph id="AB4483373851FFD9FF2CFAECFCCAFAED" blockId="3.[151,843,1287,1311]" box="[151,843,1287,1311]" pageId="3" pageNumber="58">
<emphasis id="998F5F253851FFD9FF2CFAECFE98FAEC" bold="true" box="[151,281,1287,1311]" pageId="3" pageNumber="58">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="6CFBF8B43851FFD9FE16FAECFD00FAED" box="[429,641,1287,1310]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="58" phylum="Chordata" rank="species" species="laoensis">
<emphasis id="998F5F253851FFD9FE16FAECFD00FAED" box="[429,641,1287,1310]" italics="true" pageId="3" pageNumber="58">Cnemaspis laoensis</emphasis>
</taxonomicName>
(THNHM 12433).
</paragraph>
</caption>
<subSubSection id="E3E1D0BC3851FFDEFF7EFAACFB22FC14" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58" type="description">
<paragraph id="AB4483373851FFD9FF7EFAACFB31F8D2" blockId="3.[151,1437,1351,2012]" pageId="3" pageNumber="58">
<emphasis id="998F5F253851FFD9FF7EFAACFE6FFA92" bold="true" box="[197,494,1351,1377]" pageId="3" pageNumber="58">
Description of
<typeStatus id="74403D953851FFD9FE3AFAACFE6FFA92" box="[385,494,1351,1377]" pageId="3" pageNumber="58" type="holotype">holotype</typeStatus>
</emphasis>
. Adult female; SVL 40.9 mm; head oblong in dorsal profile, moderate in size (HL/SVL 0.26), somewhat narrow (HW/SVL 0.39), flat (HD/HL 0.39), distinct from neck; snout short (ES/ HL 0.49), concave in lateral profile; postnasal region constricted medially, raised; scales of rostrum flat, juxtaposed, rugose, much larger than similarly shaped scales on occiput; moderate, supraorbital ridges; shallow frontonasal sulcus; canthus rostralis smoothly rounded; eye large (ED/HL 0.23); extra-brillar fringe scales small in general but largest anteriorly; pupil round; ear opening oval, taller than wide; rostral concave dorsally, raised posteriorly, dorsal 80% divided by longitudinal groove; rostral bordered posteriorly by two supranasals and one much smaller azygous scale, laterally by first supralabials and nostrils; supranaslals followed by three prominently raised scales; 9R,L raised supralabials of similar size; 8R,L infralabials, decreasing gradually in size posteriorly; nostrils small, oblong, posteriorly directed, bordered posteriorly by small, granular, postnasal scales; mental large, subtriangular, deeply concave medially, extending to level of second infralabials, bordered posteriorly by two large, raised postmentals; gular scales raised, smooth, somewhat pointed, anterior gulars largest; throat scales smooth, raised, larger than gulars.
</paragraph>
<paragraph id="AB4483373851FFDEFF7EF8C6FBBDFD4F" blockId="3.[151,1437,1351,2012]" lastBlockId="4.[151,1437,152,1970]" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">Body slender, elongate (AG/SVL 0.50); small, smooth, slightly raised, dorsal scales equal in size throughout body, intermixed with numerous, large, multi-keeled, longitudinally arranged tubercles; tubercles extend from top of head to base of tail and are smallest anteriorly; 22 paravertebral tubercles; pectoral and abdominal scales flat, smooth, subimbricate; ventrals slightly larger than pectorals, much larger than dorsals; no pore-bearing, precloacal scales; forelimbs moderately long, slender, dorsal scales keeled; ventral scales of forearm smooth, juxtaposed to subimbricate; palmar scales smooth, raised, subimbricate; digits long with an inflected joint; claws recurved; subdigital lamellae unnotched; subdigital lamellae wide throughout length of digits, bearing a larger scale at digital inflections; interdigital webbing weak; fingers increase in length from first to fourth with fifth nearly same length as fourth; hind limbs longer and thicker than forelimbs; dorsal scales keeled, raised, juxtaposed; ventral scales of thigh, raised, keeled; subtibials keeled, larger than dorsal tibials; plantar scales rugose, slightly raised, subimbricate; no enlarged submetatarsal scales beneath first metatarsal; digits elongate with an inflected joint; claws recurved; subdigital lamellae unnotched; lamellae wide throughout length of digits except at base where scales are more granular; enlarged, scale at the digital inflections; interdigital webbing absent; toes increase in length from first to fourth with fourth being longest; 27R,L subdigital lamellae on fourth toe; caudal scales arranged in segmented whorls; dorsal caudal scales raised, keeled, juxtaposed anteriorly; moderate, middorsal furrow; deep, single, lateral furrow; subcaudals smooth; midventral subcaudals slightly enlarged; four or five scales per caudal segment; paravertebral and dorsolateral rows of large, keeled, equally sized, caudal tubercles, lateral tubercular row and ventrolateral caudal tubercles absent; tubercles in lateral, caudal furrow; caudal tubercles do not encircle tail; 2R,3L postcloacal tubercles; tail approximately 1.29% of SVL; tail complete, original.</paragraph>
<paragraph id="AB4483373856FFDEFF7EFD2CFC81FCF4" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="59">
<emphasis id="998F5F253856FFDEFF7EFD2CFE75FD12" bold="true" box="[197,500,711,737]" pageId="4" pageNumber="59">Measurements (in mm).</emphasis>
TL 52.7, TW 4.7, FL 6.5, TBL 7.1, AG 20.5, HL 10.6, HW 6.6, HD 4.1, ED 2.4, EE 2.9, ES 5.2, EN 2.3, IO 2.3, EL 1.1, IN 0.08.
</paragraph>
<paragraph id="AB4483373856FFDEFF7EFCF9FB22FC14" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="59">
<emphasis id="998F5F253856FFDEFF7EFCF9FDAAFCDF" bold="true" box="[197,555,786,812]" pageId="4" pageNumber="59">Coloration in alcohol (fig. 2).</emphasis>
Dorsal ground color of head, body, limbs and tail pale brown; rostrum and top of head bearing diffuse, faint, brown markings; postorbital stripes absent; light colored, subelliptical, paired, paravertebral markings extending from nape to base of tail, extending onto tail as wide, diffuse, light colored, caudal bands separated by thinner, dark brown bands; dark brown markings within interspaces between light markings; large, light colored markings on flanks; forelimbs and hind limbs mottled; digits bearing dark bands; all ventral surfaces beige bearing small, black stipples in each scale.
</paragraph>
</subSubSection>
<subSubSection id="E3E1D0BC3856FFDEFF7DFC19FB25FB34" pageId="4" pageNumber="59" type="distribution">
<paragraph id="AB4483373856FFDEFF7DFC19FEAEFBC1" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="59">
<emphasis id="998F5F253856FFDEFF7DFC19FEDEFBFF" bold="true" box="[198,351,1010,1036]" pageId="4" pageNumber="59">Distribution</emphasis>
.
<taxonomicName id="6CFBF8B43856FFDEFECFFC19FDDDFBF8" box="[372,604,1010,1035]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="laoensis">
<emphasis id="998F5F253856FFDEFECFFC19FDDDFBF8" box="[372,604,1010,1035]" italics="true" pageId="4" pageNumber="59">Cnemaspis laoensis</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="74403D953856FFDEFCC1FC19FC2DFBFF" box="[890,940,1010,1036]" pageId="4" pageNumber="59">type</typeStatus>
locality at Dong Phu Vieng, Savannakhet,
<collectingCountry id="D3ECC3A73856FFDEFF2CFBF3FF51FBC1" box="[151,208,1048,1074]" name="Laos" pageId="4" pageNumber="59">Laos</collectingCountry>
(fig. 1).
</paragraph>
<paragraph id="AB4483373856FFDEFF7DFBD6FB25FB34" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="59">
<emphasis id="998F5F253856FFDEFF7DFBD6FE0AFBA4" bold="true" box="[198,395,1085,1111]" pageId="4" pageNumber="59">Natural History</emphasis>
. Nothing is known of the natural history of
<emphasis id="998F5F253856FFDEFC22FBD6FB08FBA5" box="[921,1161,1085,1110]" italics="true" pageId="4" pageNumber="59">
<taxonomicName id="6CFBF8B43856FFDEFC22FBD6FB07FBA5" box="[921,1158,1085,1110]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="laoensis">Cnemaspis laoensis</taxonomicName>
.
</emphasis>
However, on the basis of it having smooth ventrals and subcaudals, it is hypothesized that this species is a karst dweller. All other karst dwelling, microhabitat specialists share this combination of characters. Furthermore, there are numerous outcroppings of karst tower formations in the general area of the
<typeStatus id="74403D953856FFDEFBB7FB46FBBFFB34" box="[1036,1086,1197,1223]" pageId="4" pageNumber="59">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="E3E1D0BC3856FFDEFF7DFB39FD3AF841" pageId="4" pageNumber="59" type="etymology">
<paragraph id="AB4483373856FFDEFF7DFB39FEBFFAE1" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="59">
<emphasis id="998F5F253856FFDEFF7DFB39FECAFB1F" bold="true" box="[198,331,1234,1260]" pageId="4" pageNumber="59">Etymology</emphasis>
. The specific epithet is in reference to this being the first species of
<taxonomicName id="6CFBF8B43856FFDEFB1BFB38FAA0FB1F" box="[1184,1313,1235,1260]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="998F5F253856FFDEFB1BFB38FAA0FB1F" box="[1184,1313,1235,1260]" italics="true" pageId="4" pageNumber="59">Cnemaspis</emphasis>
</taxonomicName>
known to occur in
<collectingCountry id="D3ECC3A73856FFDEFF46FB13FEB8FAE1" box="[253,313,1272,1298]" name="Laos" pageId="4" pageNumber="59">Laos</collectingCountry>
.
</paragraph>
<paragraph id="AB4483373856FFDEFF7DFAF6FD3AF841" blockId="4.[151,1437,152,1970]" pageId="4" pageNumber="59">
<emphasis id="998F5F253856FFDEFF7DFAF6FEEDFAC4" bold="true" box="[198,364,1309,1335]" pageId="4" pageNumber="59">Comparisons</emphasis>
.
<taxonomicName id="6CFBF8B43856FFDEFE3BFAF6FDECFAC5" box="[384,621,1309,1334]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="laoensis">
<emphasis id="998F5F253856FFDEFE3BFAF6FDECFAC5" box="[384,621,1309,1334]" italics="true" pageId="4" pageNumber="59">Cnemaspis laoensis</emphasis>
</taxonomicName>
can be separated from all species of
<taxonomicName id="6CFBF8B43856FFDEFBFDFAF6FB46FAC5" box="[1094,1223,1309,1334]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="998F5F253856FFDEFBFDFAF6FB46FAC5" box="[1094,1223,1309,1334]" italics="true" pageId="4" pageNumber="59">Cnemaspis</emphasis>
</taxonomicName>
except
<taxonomicName id="6CFBF8B43856FFDEFA90FAF6FE9EFAAF" authority="Stoliczka" authorityName="Stoliczka" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="affinis">
<emphasis id="998F5F253856FFDEFA90FAF6FA1DFAC5" box="[1323,1436,1309,1334]" italics="true" pageId="4" pageNumber="59">C. affinis</emphasis>
(Stoliczka)
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFE96FAA8FD49FAAF" authority="Grismer &amp; Ngo" authorityName="Grismer &amp; Ngo" box="[301,712,1346,1372]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="aurantiacopes">
<emphasis id="998F5F253856FFDEFE96FAA8FE7EFAAF" box="[301,511,1347,1372]" italics="true" pageId="4" pageNumber="59">C. aurantiacopes</emphasis>
Grismer &amp; Ngo
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFD62FAA8FC67FAAF" authority="Strauch" authorityName="Strauch" box="[729,998,1346,1372]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="998F5F253856FFDEFD62FAA8FC00FAAF" box="[729,897,1347,1372]" italics="true" pageId="4" pageNumber="59">C. boulengeri</emphasis>
Strauch
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFC4CFAA8FAF2FAAF" authority="Grismer &amp; Ngo" authorityName="Grismer &amp; Ngo" box="[1015,1395,1346,1372]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="caudanivea">
<emphasis id="998F5F253856FFDEFC4CFAA8FB26FAAF" box="[1015,1191,1347,1372]" italics="true" pageId="4" pageNumber="59">C. caudanivea</emphasis>
Grismer &amp; Ngo
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFA3BFAA8FD86FA71" authority="Bauer &amp; Das" authorityName="Bauer &amp; Das" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="chanthaburiensis">
<emphasis id="998F5F253856FFDEFA3BFAA8FEE3FA72" italics="true" pageId="4" pageNumber="59">C. chanthaburiensis</emphasis>
Bauer &amp; Das
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFDAFFA83FC02FA71" authority="Chan &amp; Grismer" authorityName="Chan &amp; Grismer" box="[532,899,1384,1410]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="flavigaster">
<emphasis id="998F5F253856FFDEFDAFFA83FD35FA72" box="[532,692,1384,1409]" italics="true" pageId="4" pageNumber="59">C. flavigaster</emphasis>
Chan &amp; Grismer
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFC2BFA83FBE2FA71" authority="Smith" authorityName="Smith" box="[912,1123,1384,1410]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="kumpoli">
<emphasis id="998F5F253856FFDEFC2BFA83FB93FA72" box="[912,1042,1384,1409]" italics="true" pageId="4" pageNumber="59">C. kumpoli</emphasis>
Smith
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFBD4FA83FD6BFA54" authority="Grismer, Norhayati, Chan, Belabut, Muin, Wood &amp; Grismer" authorityName="Grismer, Norhayati, Chan, Belabut, Muin, Wood &amp; Grismer" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="monachorum">
<emphasis id="998F5F253856FFDEFBD4FA83FAAFFA72" box="[1135,1326,1384,1409]" italics="true" pageId="4" pageNumber="59">C. monachorum</emphasis>
Grismer, Norhayati, Chan, Belabut, Muin, Wood &amp; Grismer
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFD4EFA66FBE8FA54" authority="Grismer &amp; Ngo" authorityName="Grismer &amp; Ngo" box="[757,1129,1421,1447]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="nuicamensis">
<emphasis id="998F5F253856FFDEFD4EFA66FC28FA55" box="[757,937,1421,1446]" italics="true" pageId="4" pageNumber="59">C. nuicamensis</emphasis>
Grismer &amp; Ngo
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFBCEFA66FED5FA3F" authority="Grismer, Ngo &amp; Grismer" authorityName="Grismer, Ngo &amp; Grismer" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="psychedelica">
<emphasis id="998F5F253856FFDEFBCEFA66FAAEFA55" box="[1141,1327,1421,1446]" italics="true" pageId="4" pageNumber="59">C. psychedelica</emphasis>
Grismer, Ngo &amp; Grismer
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFEDAFA58FDDCFA3F" authority="Smith" authorityName="Smith" box="[353,605,1458,1484]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="998F5F253856FFDEFEDAFA58FE79FA3F" box="[353,504,1459,1484]" italics="true" pageId="4" pageNumber="59">C. siamensis</emphasis>
(Smith)
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFDD0FA58FC5FFA3F" authority="Grismer &amp; Ngo" authorityName="Grismer &amp; Ngo" box="[619,990,1458,1484]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="tucdupensis">
<emphasis id="998F5F253856FFDEFDD0FA58FC9BFA3F" box="[619,794,1459,1484]" italics="true" pageId="4" pageNumber="59">C. tucdupensis</emphasis>
Grismer &amp; Ngo
</taxonomicName>
and in having smooth, as opposed to keeled, ventral scales. Lacking, as opposed to having, a lateral row of caudal tubercles separates
<taxonomicName id="6CFBF8B43856FFDEFAA2FA33FA1DFA02" box="[1305,1436,1496,1521]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="laoensis">
<emphasis id="998F5F253856FFDEFAA2FA33FA1DFA02" box="[1305,1436,1496,1521]" italics="true" pageId="4" pageNumber="59">C. laoensis</emphasis>
</taxonomicName>
from all species of
<taxonomicName id="6CFBF8B43856FFDEFE3AFA16FD84F9E5" box="[385,517,1533,1558]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="998F5F253856FFDEFE3AFA16FD84F9E5" box="[385,517,1533,1558]" italics="true" pageId="4" pageNumber="59">Cnemaspis</emphasis>
</taxonomicName>
except
<emphasis id="998F5F253856FFDEFDDFFA16FA90F9E5" box="[612,1297,1533,1558]" italics="true" pageId="4" pageNumber="59">
<taxonomicName id="6CFBF8B43856FFDEFDDFFA16FC8CF9E5" box="[612,781,1533,1558]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="boulengeri">C. boulengeri</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFCA0FA16FC22F9E5" box="[795,931,1533,1558]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="kumpoli">C. kumpoli</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFC0BFA16FBF2F9E5" box="[944,1139,1533,1558]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="psychedelica">C. psychedelica</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFB3BFA16FA90F9E5" authority="Grismer &amp; Chan" authorityName="Grismer &amp; Chan" box="[1152,1297,1533,1558]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="roticanai">C. roticanai</taxonomicName>
</emphasis>
Grismer &amp; Chan,
<taxonomicName id="6CFBF8B43856FFDEFF5FF9C8FEFAF9CF" box="[228,379,1571,1596]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="998F5F253856FFDEFF5FF9C8FEFAF9CF" box="[228,379,1571,1596]" italics="true" pageId="4" pageNumber="59">C. siamensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="6CFBF8B43856FFDEFE07F9C8FCD6F9CF" authority="Grismer &amp; Ngo. It" authorityName="Grismer &amp; Ngo. It" box="[444,855,1570,1596]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="tucdupensis">
<emphasis id="998F5F253856FFDEFE07F9C8FDEAF9CF" box="[444,619,1571,1596]" italics="true" pageId="4" pageNumber="59">C. tucdupensis</emphasis>
Grismer &amp; Ngo. It
</taxonomicName>
can be separated from
<taxonomicName id="6CFBF8B43856FFDEFBD6F9C8FAF0F9CF" authority="Stoliczka" authorityName="Stoliczka" box="[1133,1393,1570,1596]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="affinins">
<emphasis id="998F5F253856FFDEFBD6F9C8FB69F9CF" box="[1133,1256,1571,1596]" italics="true" pageId="4" pageNumber="59">C. affinins</emphasis>
(Stoliczka)
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFA3AF9C8FDA3F991" authority="Grismer &amp; Chan" authorityName="Grismer &amp; Chan" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="perhentianensis">
<emphasis id="998F5F253856FFDEFA3AF9C8FED0F992" italics="true" pageId="4" pageNumber="59">C. perhentianensis</emphasis>
Grismer &amp; Chan
</taxonomicName>
, and
<taxonomicName id="6CFBF8B43856FFDEFDDAF9A3FD76F992" box="[609,759,1608,1633]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="siamensis">
<emphasis id="998F5F253856FFDEFDDAF9A3FD76F992" box="[609,759,1608,1633]" italics="true" pageId="4" pageNumber="59">C. siamensis</emphasis>
</taxonomicName>
in having sooth as opposed to keeled subcaudals. Being less than
<quantity id="6C032ED23856FFDEFEB7F986FEE3F974" box="[268,354,1645,1671]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="59" unit="mm" value="56.0">56 mm</quantity>
SVL separates it from
<taxonomicName id="6CFBF8B43856FFDEFD3DF986FCC5F974" authority="Dring" authorityName="Dring" box="[646,836,1645,1671]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="argus">
<emphasis id="998F5F253856FFDEFD3DF986FD71F975" box="[646,752,1645,1670]" italics="true" pageId="4" pageNumber="59">C. argus</emphasis>
Dring
</taxonomicName>
,
<emphasis id="998F5F253856FFDEFCEAF986FB2CF975" box="[849,1197,1645,1670]" italics="true" pageId="4" pageNumber="59">
<taxonomicName id="6CFBF8B43856FFDEFCEAF986FBAAF975" box="[849,1067,1645,1670]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="aurantiacopes">C. aurantiacopes</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFB8CF986FB2CF975" authority="Das &amp; Grismer" authorityName="Das &amp; Grismer" box="[1079,1197,1645,1670]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="baueri">C. baueri</taxonomicName>
</emphasis>
Das &amp; Grismer,
<emphasis id="998F5F253856FFDEFA3BF986FE9EF95F" italics="true" pageId="4" pageNumber="59">
<taxonomicName id="6CFBF8B43856FFDEFA3BF986FE9BF95F" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="boulengeri">C. boulengeri</taxonomicName>
,
</emphasis>
<taxonomicName id="6CFBF8B43856FFDEFE9DF978FD89F95F" authority="(Gray)" authorityName="Gray" baseAuthorityName="Gray" box="[294,520,1682,1708]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="kendallii">
<emphasis id="998F5F253856FFDEFE9DF978FE33F95F" box="[294,434,1683,1708]" italics="true" pageId="4" pageNumber="59">C. kendallii</emphasis>
(Gray)
</taxonomicName>
,
<emphasis id="998F5F253856FFDEFDA3F978FD7AF95F" box="[536,763,1683,1708]" italics="true" pageId="4" pageNumber="59">
<taxonomicName id="6CFBF8B43856FFDEFDA3F978FD1CF95F" box="[536,669,1683,1708]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="kumpoli">C. kumpoli</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFD11F978FD7AF95F" authority="Das &amp; Grismer" authorityName="Das &amp; Grismer" box="[682,763,1683,1708]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="limi">C. limi</taxonomicName>
</emphasis>
Das &amp; Grismer,
<taxonomicName id="6CFBF8B43856FFDEFC7DF978FF59F921" authority="Grismer, Grismer, Wood &amp; Chan" authorityName="Grismer, Grismer, Wood &amp; Chan" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="mcguirei">
<emphasis id="998F5F253856FFDEFC7DF978FBD0F95F" box="[966,1105,1683,1708]" italics="true" pageId="4" pageNumber="59">C. mcguirei</emphasis>
Grismer, Grismer, Wood &amp; Chan
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFF58F953FE42F921" authority="Smith" authorityName="Smith" box="[227,451,1720,1746]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="nigridia">
<emphasis id="998F5F253856FFDEFF58F953FEE3F922" box="[227,354,1720,1745]" italics="true" pageId="4" pageNumber="59">C. nigridia</emphasis>
(Smith)
</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFE6AF953FCE3F921" authority="Grismer &amp; Das" authorityName="Grismer &amp; Das" box="[465,866,1720,1746]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="pemanggilensis">
<emphasis id="998F5F253856FFDEFE6AF953FD28F922" box="[465,681,1720,1745]" italics="true" pageId="4" pageNumber="59">C. pemanggilensis</emphasis>
Grismer &amp; Das
</taxonomicName>
, and
<taxonomicName id="6CFBF8B43856FFDEFC1AF953FBDBF922" box="[929,1114,1720,1745]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="psychedelica">
<emphasis id="998F5F253856FFDEFC1AF953FBDBF922" box="[929,1114,1720,1745]" italics="true" pageId="4" pageNumber="59">C. psychedelica</emphasis>
</taxonomicName>
which all have a maximum SVL much greater than
<quantity id="6C032ED23856FFDEFE14F936FD84F904" box="[431,517,1757,1783]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="59" unit="mm" value="56.0">56 mm</quantity>
. It is separated from
<taxonomicName id="6CFBF8B43856FFDEFD45F936FC2BF905" box="[766,938,1757,1782]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="tucdupensis">
<emphasis id="998F5F253856FFDEFD45F936FC2BF905" box="[766,938,1757,1782]" italics="true" pageId="4" pageNumber="59">C. tucdupensis</emphasis>
</taxonomicName>
in lacking, as opposed to having, enlarged scales beneath the first metatarsal. Lacking, as opposed to having, dark shoulder and/or neck patches enclosing white to yellow ocelli separate
<taxonomicName id="6CFBF8B43856FFDEFD33F8C3FC8FF8B2" box="[648,782,1832,1857]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="laoensis">
<emphasis id="998F5F253856FFDEFD33F8C3FC8FF8B2" box="[648,782,1832,1857]" italics="true" pageId="4" pageNumber="59">C. laoensis</emphasis>
</taxonomicName>
from
<emphasis id="998F5F253856FFDEFCECF8C3FBEEF8B2" box="[855,1135,1832,1857]" italics="true" pageId="4" pageNumber="59">
<taxonomicName id="6CFBF8B43856FFDEFCECF8C3FC49F8B2" box="[855,968,1832,1857]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="affinis">C. affinis</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFC6EF8C3FBEEF8B2" authority="Grismer, Chan, Nasir" authorityName="Grismer, Chan, Nasir" box="[981,1135,1832,1857]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="biocellata">C. biocellata</taxonomicName>
</emphasis>
Grismer, Chan, Nasir, &amp; Sumontha,
<emphasis id="998F5F253856FFDEFE9FF8A6FDCFF895" box="[292,590,1869,1894]" italics="true" pageId="4" pageNumber="59">
<taxonomicName id="6CFBF8B43856FFDEFE9FF8A6FE2EF895" box="[292,431,1869,1894]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="kumpoli">C. kumpoli</taxonomicName>
,
<taxonomicName id="6CFBF8B43856FFDEFE06F8A6FDCFF895" authority="Grismer, Grismer, Wood" authorityName="Grismer, Grismer, Wood" box="[445,590,1869,1894]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="mcguirei">C. mcguirei</taxonomicName>
</emphasis>
Grismer, Grismer, Wood, &amp; Chan, and
<taxonomicName id="6CFBF8B43856FFDEFB84F8A6FDEFF87F" authority="Grismer, Norhayati, Chan, Belabut, Muin, Wood" authorityName="Grismer, Norhayati, Chan, Belabut, Muin, Wood" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="pseudomcguirei">
<emphasis id="998F5F253856FFDEFB84F8A6FAA9F895" box="[1087,1320,1869,1894]" italics="true" pageId="4" pageNumber="59">C. pseudomcguirei</emphasis>
Grismer, Norhayati, Chan, Belabut, Muin, Wood
</taxonomicName>
, &amp; Grismer. These characters as well as others are scored across all species of
<taxonomicName id="6CFBF8B43856FFDEFEA8F873FE15F842" box="[275,404,1944,1969]" class="Reptilia" family="Gekkonidae" genus="Cnemaspis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="998F5F253856FFDEFEA8F873FE15F842" box="[275,404,1944,1969]" italics="true" pageId="4" pageNumber="59">Cnemaspis</emphasis>
</taxonomicName>
in
<bibRefCitation id="CF6AFEC63856FFDEFE02F873FD3AF841" box="[441,699,1944,1970]" pageId="4" pageNumber="60" refString="Grismer, L. L., Chan, K. O., Nurolhuda, N. &amp; Sumontha, M. (2008 a) A new species of karst dwelling gecko (genus Cnemaspis Strauch 1887) from the border region of Thailand and Peninsular Malaysia. Zootaxa, 1875, 51 - 68." type="journal article">Grismer et al. (2008a)</bibRefCitation>
</paragraph>
</subSubSection>
</treatment>
</document>