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<document id="E042960E04B3744BDB856C846B115B08" ID-CLB-Dataset="5859" ID-DOI="10.11646/zootaxa.4985.2.9" ID-GBIF-Dataset="146beedf-2d3c-4ba4-944a-c6050ffde9cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="4943584" ID-ZooBank="F799B16C-4CAE-49B0-AA18-DB0C77B4264F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623661288461" checkinUser="plazi" docAuthor="Lee, Jae-Seok, Yoo, In-Seong &amp; Ahn, Kee-Jeong" docDate="2021" docId="231E878BFFD6FFF5D5DA607FEE35F8F9" docLanguage="en" docName="zootaxa.4985.2.9.pdf" docOrigin="Zootaxa 4985 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thinobius koreanus Lee &amp; Yoo &amp; Ahn 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="265" masterDocId="DF27FFF3FFD5FFF1D54D656BEF51FFF3" masterDocTitle="Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea" masterLastPageNumber="268" masterPageNumber="261" pageNumber="264" updateTime="1699014158557" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C4A98259D11D3875E53A92D029C04E9C">Taxonomy of the coastal Thinobius Kiesenwetter (Coleoptera: Staphylinidae Oxytelinae) in Korea</mods:title>
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<mods:namePart id="06D096340951203FA51935CCB93DA597">Lee, Jae-Seok</mods:namePart>
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<mods:affiliation id="75343B245BFF82D88376439E86921BCE">Department of Biology, Chungnam National University, Daejeon 34134, South Korea &amp; ljs 7615 @ naver. com; https: // orcid. org / 0000 - 0003 - 4502 - 7330</mods:affiliation>
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<mods:namePart id="FDE02F3F4A8BADFBC55F7BA761186CD3">Yoo, In-Seong</mods:namePart>
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<mods:affiliation id="6F30C501765D6F91444E7CB7675B7657">Division of Restoration Research, Research Center for Endangered Species, National Institute of Ecology, Yeongyang 36531, South Korea &amp; webila @ naver. com; https: // orcid. org / 0000 - 0002 - 6720 - 6025</mods:affiliation>
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<mods:namePart id="4A9BBA3EE63521397C0D837115D56040">Ahn, Kee-Jeong</mods:namePart>
<mods:affiliation id="F30C4F7E44E371C5694AB8DF109E9063">Department of Biology, Chungnam National University, Daejeon 34134, South Korea</mods:affiliation>
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<treatment id="231E878BFFD6FFF5D5DA607FEE35F8F9" ID-DOI="http://doi.org/10.5281/zenodo.5056445" ID-GBIF-Taxon="183232159" ID-Zenodo-Dep="5056445" LSID="urn:lsid:plazi:treatment:231E878BFFD6FFF5D5DA607FEE35F8F9" httpUri="http://treatment.plazi.org/id/231E878BFFD6FFF5D5DA607FEE35F8F9" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264">
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<paragraph id="AB08369DFFD6FFF2D5DA607FED0EFADC" blockId="3.[151,607,1300,1364]" box="[151,607,1300,1327]" pageId="3" pageNumber="264">
<heading id="F04081F1FFD6FFF2D5DA607FED0EFADC" bold="true" box="[151,607,1300,1327]" fontSize="11" level="1" pageId="3" pageNumber="264" reason="1">
<emphasis id="99C3EA8FFFD6FFF2D5DA607FED0EFADC" bold="true" box="[151,607,1300,1327]" pageId="3" pageNumber="264">
<taxonomicName id="6CB74D1EFFD6FFF2D5DA607FED44FADC" ID-CoL="8YBR7" authority="Lee &amp; Yoo" authorityName="Lee &amp; Yoo &amp; Ahn" authorityYear="2021" box="[151,533,1300,1327]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="99C3EA8FFFD6FFF2D5DA607FEEDBFADC" bold="true" box="[151,394,1300,1327]" italics="true" pageId="3" pageNumber="264">Thinobius koreanus</emphasis>
Lee &amp; Yoo
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<taxonomicNameLabel id="82F057F4FFD6FFF2D751607EED0EFADC" box="[540,607,1301,1327]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="E3AD6516FFD6FFF5D5DA6052EB89FE96" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264" type="description">
<paragraph id="AB08369DFFD6FFF2D5DA6052EE10FAA7" blockId="3.[151,607,1300,1364]" box="[151,321,1337,1364]" pageId="3" pageNumber="264">
(
<figureCitation id="338C2A18FFD6FFF2D5D26052EE50FAA0" box="[159,257,1337,1363]" captionStart="FIGURE 1" captionStartId="1.[151,250,1452,1476]" captionTargetBox="[162,1422,319,1418]" captionTargetId="figure-75@1.[151,1436,308,1428]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Korean coastal Thinobius. A. Thinobius jejuensis sp. n., 1.04 mm; C. Thinobius koreanus sp. n., 1.60 mm; B. Thinobius kuroshio, 1.06 mm." figureDoi="http://doi.org/10.5281/zenodo.4943586" httpUri="https://zenodo.org/record/4943586/files/figure.png" pageId="3" pageNumber="264">Figs. 1B</figureCitation>
,
<figureCitation id="338C2A18FFD6FFF2D4406052EE68FAA7" box="[269,313,1337,1364]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="4.[151,250,1201,1225]" captionStartId-1="5.[151,247,931,955]" captionTargetBox-0="[183,1375,391,1154]" captionTargetBox-1="[156,1426,187,898]" captionTargetId-0="figure-133@4.[171,1416,381,1178]" captionTargetId-1="figure-19@5.[151,1436,181,907]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 4. Thinobius koreanus sp. n. A: antenna; B: head, dorsal aspect; C: pronotum, dorsal aspect; D: prosternum, ventral aspect; E: meso- and metaventrite, ventral aspect; F: ring structure with female genital appendage. Scales = 0.1 mm. Circles in B and E show magnification of sculptures." captionText-1="FIGURE 5. Thinobius koreanus sp. n. A: aedeagus, ventral aspect; B: aedeagus, lateral aspect; C: spermatheca. Scales = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4943598" figureDoi-1="http://doi.org/10.5281/zenodo.4943600" httpUri-0="https://zenodo.org/record/4943598/files/figure.png" httpUri-1="https://zenodo.org/record/4943600/files/figure.png" pageId="3" pageNumber="264">45</figureCitation>
)
</paragraph>
<paragraph id="AB08369DFFD6FFF5D5DA60EAEB89FE96" blockId="3.[151,1437,1409,2011]" lastBlockId="4.[151,1437,151,357]" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264">
<emphasis id="99C3EA8FFFD6FFF2D5DA60EAEE78FA68" bold="true" box="[151,297,1409,1435]" pageId="3" pageNumber="264">Description.</emphasis>
Length
<quantity id="6C4F9B78FFD6FFF2D4C460EAED60FA68" box="[393,561,1409,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="3" pageNumber="264" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.601.80 mm</quantity>
. Body flattened, surface pubescent; light brown to dark brown, head and abdomen dark brown, pronotum, elytra and antenna brown, mouthparts and legs light brown.
<emphasis id="99C3EA8FFFD6FFF2D1EB60CEEBB2FA4C" box="[1190,1251,1445,1471]" italics="true" pageId="3" pageNumber="264">Head</emphasis>
. Almost as long as wide and slightly narrower than pronotum; widest across eyes; clypeus trapezoidal; eye moderate in size and rounded in lateral aspect, with large eye facet and interfacetal setae; temple well developed, diameter of eye slightly longer than temple; gular suture fused and broadly divergent at posterior part; neck well developed, transversely reticulate sculptures distinct, seta absent (
<figureCitation id="338C2A18FFD6FFF2D700635EEDF5F9BC" box="[589,676,1589,1615]" captionStart="FIGURE 4" captionStartId="4.[151,250,1201,1225]" captionTargetBox="[183,1375,391,1154]" captionTargetId="figure-133@4.[171,1416,381,1178]" captionTargetPageId="4" captionText="FIGURE 4. Thinobius koreanus sp. n. A: antenna; B: head, dorsal aspect; C: pronotum, dorsal aspect; D: prosternum, ventral aspect; E: meso- and metaventrite, ventral aspect; F: ring structure with female genital appendage. Scales = 0.1 mm. Circles in B and E show magnification of sculptures." figureDoi="http://doi.org/10.5281/zenodo.4943598" httpUri="https://zenodo.org/record/4943598/files/figure.png" pageId="3" pageNumber="264">Fig. 4B</figureCitation>
); antenna slender, moderately elongate, and pubescent; antennomere 1 longest, 3 shortest, 111 elongate (
<figureCitation id="338C2A18FFD6FFF2D70E6332EDF0F980" box="[579,673,1625,1651]" captionStart="FIGURE 4" captionStartId="4.[151,250,1201,1225]" captionTargetBox="[183,1375,391,1154]" captionTargetId="figure-133@4.[171,1416,381,1178]" captionTargetPageId="4" captionText="FIGURE 4. Thinobius koreanus sp. n. A: antenna; B: head, dorsal aspect; C: pronotum, dorsal aspect; D: prosternum, ventral aspect; E: meso- and metaventrite, ventral aspect; F: ring structure with female genital appendage. Scales = 0.1 mm. Circles in B and E show magnification of sculptures." figureDoi="http://doi.org/10.5281/zenodo.4943598" httpUri="https://zenodo.org/record/4943598/files/figure.png" pageId="3" pageNumber="264">Fig. 4A</figureCitation>
).
<emphasis id="99C3EA8FFFD6FFF2D7FA6332EC6BF980" box="[695,826,1625,1651]" italics="true" pageId="3" pageNumber="264">Mouthparts</emphasis>
. Labrum transverse, lateral margin rounded, anterior margin slightly emarginate; mandible triangular, apex bifid with subapical tooth, prostheca and molar tooth well developed; maxillary palpus with four palpomeres, palpomere 1 transverse and small, 2 slightly dilated to apex, 3 largest, 23 with many setae, 4 short and slender; apical margin of 3 more than 2.0 times as wide as basal margin of 4; labial palpus with three elongate palpi, palpomere 1 with one long, one short seta and two pores, 2 with one long seta and one pore, 3 with few short sensilla at apex; mentum quadrate with two pairs of long macrosetae and many setae.
<emphasis id="99C3EA8FFFD6FFF2D46E625AEE25F8B8" box="[291,372,1841,1867]" italics="true" pageId="3" pageNumber="264">Thorax</emphasis>
. Pronotum transverse, width
<quantity id="6C4F9B78FFD6FFF2D7F3625AEC32F8B8" box="[702,867,1841,1867]" metricMagnitude="-4" metricUnit="m" metricValue="2.9499999999999997" metricValueMax="3.0" metricValueMin="2.9" pageId="3" pageNumber="264" unit="mm" value="0.295" valueMax="0.3" valueMin="0.29">0.290.30 mm</quantity>
, widest near half of length, surface pubescent with seven pairs of macrosetae and many setae; prosternum with one pair of long setae and a few short setae, apex of prosternal process sharp and elevated (
<figureCitation id="338C2A18FFD6FFF2D7196212ED8DF860" box="[596,732,1913,1939]" captionStart="FIGURE 4" captionStartId="4.[151,250,1201,1225]" captionTargetBox="[183,1375,391,1154]" captionTargetId="figure-133@4.[171,1416,381,1178]" captionTargetPageId="4" captionText="FIGURE 4. Thinobius koreanus sp. n. A: antenna; B: head, dorsal aspect; C: pronotum, dorsal aspect; D: prosternum, ventral aspect; E: meso- and metaventrite, ventral aspect; F: ring structure with female genital appendage. Scales = 0.1 mm. Circles in B and E show magnification of sculptures." figureDoi="http://doi.org/10.5281/zenodo.4943598" httpUri="https://zenodo.org/record/4943598/files/figure.png" pageId="3" pageNumber="264">Figs. 4CD</figureCitation>
); scutellum an inverted triangle, anterior margin of scutellum emarginate, posterior part reticulate with a few setae and inverted triangle impression, impression about 1.6 times as wide as long, elytron elongate and pubescent; posterior margin with membranous lobe; hind wing present; mesoventral process slightly extended and pointed at apex; metaventral carina absent; metaventrite distinctly reticulate with two pairs of long macrosetae and many setae (
<figureCitation id="338C2A18FFD1FFF5D78365D0EC74FF26" box="[718,805,187,213]" captionStart="FIGURE 4" captionStartId="4.[151,250,1201,1225]" captionTargetBox="[183,1375,391,1154]" captionTargetId="figure-133@4.[171,1416,381,1178]" captionTargetPageId="4" captionText="FIGURE 4. Thinobius koreanus sp. n. A: antenna; B: head, dorsal aspect; C: pronotum, dorsal aspect; D: prosternum, ventral aspect; E: meso- and metaventrite, ventral aspect; F: ring structure with female genital appendage. Scales = 0.1 mm. Circles in B and E show magnification of sculptures." figureDoi="http://doi.org/10.5281/zenodo.4943598" httpUri="https://zenodo.org/record/4943598/files/figure.png" pageId="4" pageNumber="265">Fig. 4E</figureCitation>
); all tibiae with one long macroseta in middle.
<emphasis id="99C3EA8FFFD1FFF5D06665D0EAC4FF26" box="[1323,1429,187,213]" italics="true" pageId="4" pageNumber="265">Abdomen</emphasis>
. Posterior margin of tergite VII with fringe of setae; posterior margin of tergite VIII emarginate; posterior margin of sternite VIII convex.
<emphasis id="99C3EA8FFFD1FFF5D4C96468EDA5FEEE" box="[388,756,259,285]" italics="true" pageId="4" pageNumber="265">Secondary sexual characteristics</emphasis>
. Male tergite IX with ventral structs, struts shorter than other part, female gonocoxites present; female with one ring structure (
<figureCitation id="338C2A18FFD1FFF5D63C644CEC97FEB2" box="[881,966,295,321]" captionStart="FIGURE 4" captionStartId="4.[151,250,1201,1225]" captionTargetBox="[183,1375,391,1154]" captionTargetId="figure-133@4.[171,1416,381,1178]" captionTargetPageId="4" captionText="FIGURE 4. Thinobius koreanus sp. n. A: antenna; B: head, dorsal aspect; C: pronotum, dorsal aspect; D: prosternum, ventral aspect; E: meso- and metaventrite, ventral aspect; F: ring structure with female genital appendage. Scales = 0.1 mm. Circles in B and E show magnification of sculptures." figureDoi="http://doi.org/10.5281/zenodo.4943598" httpUri="https://zenodo.org/record/4943598/files/figure.png" pageId="4" pageNumber="265">Fig. 4F</figureCitation>
).
<emphasis id="99C3EA8FFFD1FFF5D694644CEB12FEB2" box="[985,1091,295,321]" italics="true" pageId="4" pageNumber="265">Genitalia</emphasis>
. Aedeagus oval (
<figureCitation id="338C2A18FFD1FFF5D049644CEA0AFEB2" box="[1284,1371,294,321]" captionStart="FIGURE 5" captionStartId="5.[151,247,931,955]" captionTargetBox="[156,1426,187,898]" captionTargetId="figure-19@5.[151,1436,181,907]" captionTargetPageId="5" captionText="FIGURE 5. Thinobius koreanus sp. n. A: aedeagus, ventral aspect; B: aedeagus, lateral aspect; C: spermatheca. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4943600" httpUri="https://zenodo.org/record/4943600/files/figure.png" pageId="4" pageNumber="265">Fig. 5A</figureCitation>
) with serrated pattern in lateral aspect (
<figureCitation id="338C2A18FFD1FFF5D7466420ED32FE96" box="[523,611,330,357]" captionStart="FIGURE 5" captionStartId="5.[151,247,931,955]" captionTargetBox="[156,1426,187,898]" captionTargetId="figure-19@5.[151,1436,181,907]" captionTargetPageId="5" captionText="FIGURE 5. Thinobius koreanus sp. n. A: aedeagus, ventral aspect; B: aedeagus, lateral aspect; C: spermatheca. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4943600" httpUri="https://zenodo.org/record/4943600/files/figure.png" pageId="4" pageNumber="265">Fig. 5B</figureCitation>
); paramere long and slender; spermatheca as in
<figureCitation id="338C2A18FFD1FFF5D1306420EB85FE96" box="[1149,1236,330,357]" captionStart="FIGURE 5" captionStartId="5.[151,247,931,955]" captionTargetBox="[156,1426,187,898]" captionTargetId="figure-19@5.[151,1436,181,907]" captionTargetPageId="5" captionText="FIGURE 5. Thinobius koreanus sp. n. A: aedeagus, ventral aspect; B: aedeagus, lateral aspect; C: spermatheca. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4943600" httpUri="https://zenodo.org/record/4943600/files/figure.png" pageId="4" pageNumber="265">Fig. 5C</figureCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="AB08369DFFD1FFF5D5DA61DAED19FAE2" blockId="4.[151,1437,1201,1297]" pageId="4" pageNumber="265">
<emphasis id="99C3EA8FFFD1FFF5D5DA61DAEE45FB3A" bold="true" box="[151,276,1201,1225]" pageId="4" pageNumber="265">FIGURE 4.</emphasis>
<taxonomicName id="6CB74D1EFFD1FFF5D45661DAEEB2FB3A" authority="Lee &amp; Yoo &amp; Ahn, 2021" authorityName="Lee &amp; Yoo &amp; Ahn" authorityYear="2021" box="[283,483,1201,1225]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="99C3EA8FFFD1FFF5D45661DAEEB2FB3A" box="[283,483,1201,1225]" italics="true" pageId="4" pageNumber="265">Thinobius koreanus</emphasis>
</taxonomicName>
<emphasis id="99C3EA8FFFD1FFF5D4A761D9ED72FB39" bold="true" box="[490,547,1202,1226]" pageId="4" pageNumber="265">
<taxonomicNameLabel id="82F057F4FFD1FFF5D4A761D9ED72FB39" box="[490,547,1202,1226]" pageId="4" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A: antenna; B: head, dorsal aspect; C: pronotum, dorsal aspect; D: prosternum, ventral aspect; E: meso- and metaventrite, ventral aspect; F: ring structure with female genital appendage. Scales = 0.1 mm. Circles in B and E show magnification of sculptures.
</paragraph>
</caption>
<subSubSection id="E3AD6516FFD1FFF5D58A602BECAFF9FD" pageId="4" pageNumber="265" type="materials_examined">
<paragraph id="AB08369DFFD1FFF5D58A602BECAFF9FD" blockId="4.[151,1437,1343,1802]" pageId="4" pageNumber="265">
<emphasis id="99C3EA8FFFD1FFF5D58A602BEE26FAAA" bold="true" box="[199,375,1343,1370]" pageId="4" pageNumber="265">Type material.</emphasis>
<materialsCitation id="1BDF3CC0FFD1FFF5D4CD602BEB0BFA35" ID-GBIF-Occurrence="3312842303" collectingDate="2018-07-12" collectorName="IS Yoo &amp; JS Lee &amp; JG Jung" country="South Korea" county="Pohang-si" elevation="17" latitude="36.177425" location="Cheongjin-ri" longLatPrecision="1" longitude="129.39365" municipality="Buk-gu" pageId="4" pageNumber="265" specimenCount="1" specimenCount-male="1" stateProvince="Gyeongbuk" typeStatus="holotype">
<typeStatus id="740C883FFFD1FFF5D4CD602BEEBAFAA9" box="[384,491,1344,1370]" pageId="4" pageNumber="265" type="holotype">Holotype</typeStatus>
,
<specimenCount id="BDB1FD14FFD1FFF5D4B4602BED76FAA9" box="[505,551,1344,1370]" pageId="4" pageNumber="265" type="male">1 ♂</specimenCount>
, labeled as follows:
<collectingCountry id="D3A0760DFFD1FFF5D654602BECD2FAA9" box="[793,899,1344,1370]" name="South Korea" pageId="4" pageNumber="265">KOREA</collectingCountry>
:
<collectingRegion id="6973F87FFFD1FFF5D6C2602BEB09FAA9" box="[911,1112,1344,1370]" country="South Korea" name="Gyeongbuk" pageId="4" pageNumber="265">Gyeongbuk Prov.</collectingRegion>
,
<collectingCounty id="42694E11FFD1FFF5D128602BEB89FAA9" box="[1125,1240,1344,1370]" pageId="4" pageNumber="265">Pohang-si</collectingCounty>
,
<collectingMunicipality id="4B6CACE7FFD1FFF5D1AB602BEA6CFAA9" box="[1254,1341,1344,1370]" pageId="4" pageNumber="265">Buk-gu</collectingMunicipality>
, Cheongha-myeon,
<location id="AE686046FFD1FFF5D468600FEEE6FA8D" LSID="urn:lsid:plazi:treatment:231E878BFFD6FFF5D5DA607FEE35F8F9:AE686046FFD1FFF5D468600FEEE6FA8D" box="[293,439,1380,1406]" country="South Korea" county="Pohang-si" latitude="36.177425" longLatPrecision="1" longitude="129.39365" municipality="Buk-gu" name="Cheongjin-ri" pageId="4" pageNumber="265" stateProvince="Gyeongbuk">Cheongjin-ri</location>
,
<geoCoordinate id="CE83505AFFD1FFF5D48E600FED3BFA8D" box="[451,618,1380,1406]" degrees="36" direction="north" minutes="10" orientation="latitude" pageId="4" pageNumber="265" precision="1" seconds="38.73" value="36.177425">N36°10'38.73&quot;</geoCoordinate>
<geoCoordinate id="CE83505AFFD1FFF5D73F600FEC75FA8C" box="[626,804,1380,1407]" degrees="129" direction="east" minutes="23" orientation="longitude" pageId="4" pageNumber="265" precision="1" seconds="37.15" value="129.39365">E129°23'37.15&quot;</geoCoordinate>
<quantity id="6C4F9B78FFD1FFF5D660600FEC38FA8D" box="[813,873,1380,1406]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="265" unit="m" value="17.0">
<elevation id="209AD1AEFFD1FFF5D660600FEC38FA8D" box="[813,873,1380,1406]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="265" unit="m" value="17.0">17 m</elevation>
</quantity>
,
<date id="DF09105DFFD1FFF5D63B600FEB54FA8D" box="[886,1029,1380,1406]" pageId="4" pageNumber="265" value="2018-07-12">
<collectingDate id="CF4DE9B5FFD1FFF5D63B600FEB54FA8D" box="[886,1029,1380,1406]" pageId="4" pageNumber="265" value="2018-07-12">12 VII 2018</collectingDate>
</date>
,
<collectorName id="0642534BFFD1FFF5D15C600FEB30FA8D" box="[1041,1121,1380,1406]" pageId="4" pageNumber="265">IS Yoo</collectorName>
,
<collectorName id="0642534BFFD1FFF5D123600FEBEFFA8D" box="[1134,1214,1380,1406]" pageId="4" pageNumber="265">JS Lee</collectorName>
,
<collectorName id="0642534BFFD1FFF5D186600FEA7BFA8D" box="[1227,1322,1380,1406]" pageId="4" pageNumber="265">JG Jung</collectorName>
, in coarse sand/gravels on seashore, flotation
<typeStatus id="740C883FFFD1FFF5D76460E3EDC8FA51" box="[553,665,1416,1442]" pageId="4" pageNumber="265" type="holotype">Holotype</typeStatus>
,
<taxonomicName id="6CB74D1EFFD1FFF5D7E960E3EE5CFA35" authority="Lee and Yoo, Desig. Jae-Seok Lee and In-Seong Yoo, 2021" authorityName="Lee and Yoo, Desig. Jae-Seok Lee and In-Seong Yoo" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="koreanus">
<emphasis id="99C3EA8FFFD1FFF5D7E960E3ECD1FA51" box="[676,896,1416,1442]" italics="true" pageId="4" pageNumber="265">Thinobius koreanus</emphasis>
Lee and Yoo, Desig. Jae-Seok Lee and In-Seong Yoo, 2021
</taxonomicName>
<collectingCounty id="42694E11FFD1FFF5D45660C7EEC3FA35" box="[283,402,1452,1478]" pageId="4" pageNumber="265">Deposited</collectingCounty>
in the
<collectingRegion id="6973F87FFFD1FFF5D49660C7ED07FA35" box="[475,598,1452,1478]" country="South Korea" name="Chungnam" pageId="4" pageNumber="265">Chungnam</collectingRegion>
<collectingMunicipality id="4B6CACE7FFD1FFF5D71160C7EB51FA35" box="[604,1024,1452,1478]" pageId="4" pageNumber="265">National University Insect Collection</collectingMunicipality>
,
<location id="AE686046FFD1FFF5D14460C7EB0BFA35" LSID="urn:lsid:plazi:treatment:231E878BFFD6FFF5D5DA607FEE35F8F9:AE686046FFD1FFF5D14460C7EB0BFA35" box="[1033,1114,1452,1478]" country="South Korea" county="Deposited" municipality="National University Insect Collection" name="Korea'" pageId="4" pageNumber="265" stateProvince="Chungnam">Korea</location>
</materialsCitation>
.
<materialsCitation id="1BDF3CC0FFD1FFF5D12960C7ECA6F9FD" ID-GBIF-Occurrence="3312842305" collectingDate="2010-08-27" collectorName="KJ Ahn &amp; TK Kim &amp; YH Kim &amp; IS Yoo &amp; JH Song &amp; SG Lee &amp; JH Jeon" country="South Korea" county="Deposited" elevation="9" latitude="36.17717" location="Korea'" longLatPrecision="1" longitude="129.39389" municipality="National University Insect Collection" pageId="4" pageNumber="265" specimenCount="2" stateProvince="Chungnam" typeStatus="paratype">
<typeStatus id="740C883FFFD1FFF5D12960C7EB85FA35" box="[1124,1236,1452,1478]" pageId="4" pageNumber="265" type="paratype">Paratypes</typeStatus>
, 7 exx., same data as holotype, 5 exx., same data as
<typeStatus id="740C883FFFD1FFF5D75D60BBED23FA19" box="[528,626,1488,1514]" pageId="4" pageNumber="265" type="holotype">holotype</typeStatus>
except for
<geoCoordinate id="CE83505AFFD1FFF5D7BA60BAECD7FA1F" box="[759,902,1489,1516]" degrees="36" direction="north" minutes="10" orientation="latitude" pageId="4" pageNumber="265" precision="1" seconds="37.8" value="36.17717">N36°10ʹ37.8</geoCoordinate>
<geoCoordinate id="CE83505AFFD1FFF5D6CB60BAEB69FA1F" box="[902,1080,1489,1516]" degrees="129" direction="east" minutes="23" orientation="longitude" pageId="4" pageNumber="265" precision="1" seconds="38.0" value="129.39389">ʺE129°23ʹ38.0ʺ</geoCoordinate>
<quantity id="6C4F9B78FFD1FFF5D10D60BBEB37FA19" box="[1088,1126,1488,1514]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="265" unit="m" value="9.0">
<elevation id="209AD1AEFFD1FFF5D10D60BBEB37FA19" box="[1088,1126,1488,1514]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="265" unit="m" value="9.0">9m</elevation>
</quantity>
,
<date id="DF09105DFFD1FFF5D13F60BBEA59FA19" box="[1138,1288,1488,1514]" pageId="4" pageNumber="265" value="2010-08-27">
<collectingDate id="CF4DE9B5FFD1FFF5D13F60BBEA59FA19" box="[1138,1288,1488,1514]" pageId="4" pageNumber="265" value="2010-08-27">27 VIII 2010</collectingDate>
</date>
,
<collectorName id="0642534BFFD1FFF5D05E60BBEA3DFA19" box="[1299,1388,1488,1514]" pageId="4" pageNumber="265">KJ Ahn</collectorName>
,
<collectorName id="0642534BFFD1FFF5D03A60BBEF9DF9FD" pageId="4" pageNumber="265">TK Kim</collectorName>
,
<collectorName id="0642534BFFD1FFF5D59B609FEE6BF9FD" box="[214,314,1524,1550]" pageId="4" pageNumber="265">YH Kim</collectorName>
,
<collectorName id="0642534BFFD1FFF5D408609FEEC5F9FD" box="[325,404,1524,1550]" pageId="4" pageNumber="265">IS Yoo</collectorName>
,
<collectorName id="0642534BFFD1FFF5D4D2609FED53F9FD" box="[415,514,1524,1550]" pageId="4" pageNumber="265">JH Song</collectorName>
,
<collectorName id="0642534BFFD1FFF5D740609FED34F9FD" box="[525,613,1524,1550]" pageId="4" pageNumber="265">SG Lee</collectorName>
,
<collectorName id="0642534BFFD1FFF5D73D609FED9DF9FD" box="[624,716,1524,1550]" pageId="4" pageNumber="265">JH Jeon</collectorName>
, under stones on seashore
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E3AD6516FFD1FFF5D58A637CEEE5F9C1" box="[199,436,1559,1586]" pageId="4" pageNumber="265" type="distribution">
<paragraph id="AB08369DFFD1FFF5D58A637CEEE5F9C1" blockId="4.[151,1437,1343,1802]" box="[199,436,1559,1586]" pageId="4" pageNumber="265">
<emphasis id="99C3EA8FFFD1FFF5D58A637CEE30F9C2" bold="true" box="[199,353,1559,1585]" pageId="4" pageNumber="265">Distribution.</emphasis>
<collectingCountry id="D3A0760DFFD1FFF5D42A6373EEFEF9C1" box="[359,431,1560,1586]" name="South Korea" pageId="4" pageNumber="265">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E3AD6516FFD1FFF5D58A6357EE35F8F9" pageId="4" pageNumber="265" type="discussion">
<paragraph id="AB08369DFFD1FFF5D58A6357EE35F8F9" blockId="4.[151,1437,1343,1802]" pageId="4" pageNumber="265">
<emphasis id="99C3EA8FFFD1FFF5D58A6357EE6AF9A5" bold="true" box="[199,315,1596,1622]" pageId="4" pageNumber="265">Remarks.</emphasis>
This species is similar to
<taxonomicName id="6CB74D1EFFD1FFF5D7256356EDB0F9A5" baseAuthorityName="Cameron" baseAuthorityYear="1917" box="[616,737,1597,1622]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="marinus">
<emphasis id="99C3EA8FFFD1FFF5D7256356EDB0F9A5" box="[616,737,1597,1622]" italics="true" pageId="4" pageNumber="265">T. marinus</emphasis>
</taxonomicName>
, but can be distinguished by the shape of antennomeres. Antennomeres 111 are elongate in
<taxonomicName id="6CB74D1EFFD1FFF5D75C630BEDBEF989" authority="Lee &amp; Yoo &amp; Ahn, 2021" authorityName="Lee &amp; Yoo &amp; Ahn" authorityYear="2021" box="[529,751,1632,1658]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="koreanus" status="sp. nov.">
<emphasis id="99C3EA8FFFD1FFF5D75C630BEDBEF989" box="[529,751,1632,1658]" italics="true" pageId="4" pageNumber="265">Thinobius koreanus</emphasis>
</taxonomicName>
<emphasis id="99C3EA8FFFD1FFF5D7B5630BEC68F989" bold="true" box="[760,825,1632,1658]" pageId="4" pageNumber="265">
<taxonomicNameLabel id="82F057F4FFD1FFF5D7B5630BEC68F989" box="[760,825,1632,1658]" pageId="4" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="338C2A18FFD1FFF5D604630BECF4F989" box="[841,933,1632,1658]" captionStart="FIGURE 4" captionStartId="4.[151,250,1201,1225]" captionTargetBox="[183,1375,391,1154]" captionTargetId="figure-133@4.[171,1416,381,1178]" captionTargetPageId="4" captionText="FIGURE 4. Thinobius koreanus sp. n. A: antenna; B: head, dorsal aspect; C: pronotum, dorsal aspect; D: prosternum, ventral aspect; E: meso- and metaventrite, ventral aspect; F: ring structure with female genital appendage. Scales = 0.1 mm. Circles in B and E show magnification of sculptures." figureDoi="http://doi.org/10.5281/zenodo.4943598" httpUri="https://zenodo.org/record/4943598/files/figure.png" pageId="4" pageNumber="265">Fig. 4A</figureCitation>
), but 48 transverse in
<taxonomicName id="6CB74D1EFFD1FFF5D1FF630AEA7AF989" baseAuthorityName="Cameron" baseAuthorityYear="1917" box="[1202,1323,1633,1658]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="marinus">
<emphasis id="99C3EA8FFFD1FFF5D1FF630AEA7AF989" box="[1202,1323,1633,1658]" italics="true" pageId="4" pageNumber="265">T. marinus</emphasis>
</taxonomicName>
. The new species is also similar to
<taxonomicName id="6CB74D1EFFD1FFF5D4FD63EEED71F96D" authorityName="Hatch" authorityYear="1957" box="[432,544,1668,1694]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="frizzelli">
<emphasis id="99C3EA8FFFD1FFF5D4FD63EEED71F96D" box="[432,544,1668,1694]" italics="true" pageId="4" pageNumber="265">T. frizzelli</emphasis>
</taxonomicName>
from
<collectingCountry id="D3A0760DFFD1FFF5D72B63EFEDEBF96D" box="[614,698,1668,1694]" name="Canada" pageId="4" pageNumber="265">Canada</collectingCountry>
and
<collectingCountry id="D3A0760DFFD1FFF5D7BC63EFEC76F96D" box="[753,807,1668,1694]" name="United States of America" pageId="4" pageNumber="265">USA</collectingCountry>
but can be distinguished by the shape of antennomeres. Antennomeres 410 are quadrate and 46 are almost same length in the new species (
<figureCitation id="338C2A18FFD1FFF5D12863C3EB91F931" box="[1125,1216,1704,1730]" captionStart="FIGURE 4" captionStartId="4.[151,250,1201,1225]" captionTargetBox="[183,1375,391,1154]" captionTargetId="figure-133@4.[171,1416,381,1178]" captionTargetPageId="4" captionText="FIGURE 4. Thinobius koreanus sp. n. A: antenna; B: head, dorsal aspect; C: pronotum, dorsal aspect; D: prosternum, ventral aspect; E: meso- and metaventrite, ventral aspect; F: ring structure with female genital appendage. Scales = 0.1 mm. Circles in B and E show magnification of sculptures." figureDoi="http://doi.org/10.5281/zenodo.4943598" httpUri="https://zenodo.org/record/4943598/files/figure.png" pageId="4" pageNumber="265">Fig. 4A</figureCitation>
), but 410 rounded form and 5 larger than 4 or
<quantity id="6C4F9B78FFD1FFF5D48763A7EEA5F915" box="[458,500,1740,1766]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="265" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="6CB74D1EFFD1FFF5D4B663A6EC77F914" authority="(Kincaid, 1961)" baseAuthorityName="Kincaid" baseAuthorityYear="1961" box="[507,806,1740,1767]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="frizzelli">
<emphasis id="99C3EA8FFFD1FFF5D4B663A6ED3BF915" box="[507,618,1740,1766]" italics="true" pageId="4" pageNumber="265">T. frizzelli</emphasis>
(
<bibRefCitation id="CF264B6CFFD1FFF5D73463A7EC4CF914" author="Kincaid, T." box="[633,797,1740,1767]" pageId="4" pageNumber="265" refId="ref3912" refString="Kincaid, T. (1961) The Ecology and Morphology of Thinobius frizzelli Hatch an Intertidal beetle. The Calliostoma Co., Seattle, Washington, 12 pp., 6 pls." type="book" year="1961">Kincaid, 1961</bibRefCitation>
)
</taxonomicName>
. All specimens were collected in the southern part of the Korean peninsula.
</paragraph>
</subSubSection>
</treatment>
</document>